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Ville Formation

{{Infobox rockunit

| name = Ville Formation

| image = File:20070812 Lackabzug Tagebau Garzweiler Miocene Frimmersdorf Sand Germany.jpg

| caption = Frimmersdorf sand and lignite

| type = Geological formation

| age = Neogene

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| country = Germany

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The Ville Formation is a geologic formation in Germany. It preserves fossils dating back to the Neogene period. Lignite of the Ville Formation is excavated in North Rhine-Westphalia.

Description

The Middle Miocene Hambach 6C site located within the Ville Formation is believed to have been deposited in an estuarine setting in a large fluviatile system with extended coal swamps surrounding it, as supported by sedimentological and palaeobotanical evidence. Many tetrapod remains found in the Ville Formation are associated with tropical elements such as primates, chameleons and carettochelyine turtles, suggesting the age of the fauna to be at approximately 15.5 Ma during the Middle Miocene Climate Transition. The paleoflora of the formation also supports a tropical-like climate at the time of deposition.{{Cite journal |last1=Čerňanský |first1=A. |last2=Szyndlar |first2=Z. |last3=Mörs |first3=T. |date=2016 |title=Fossil squamate faunas from the Neogene of Hambach (northwestern Germany) |journal=Palaeobiodiversity and Palaeoenvironments |language=en |volume=97 |issue=2 |pages=329–354 |doi=10.1007/s12549-016-0252-1|s2cid=132267739 }}

The younger late Pliocene (approximately 2.5 Ma) sites of Hambach 11 & 13 are believed to be deposited in oxygenated water and currents in a river channel setting in close vicinity to lakes or oxbows, as supported by sedimentological and palaeobotanical evidence

Fossil content

=Mammals=

==Rodents==

class="wikitable" align="center"
colspan="7" align="center" | Rodents reported from the Ville Formation
GenusSpeciesPresence

!Age

!Material

NotesImages
Myoglis

|M. meini

|Hambach 6C.{{Cite journal |last1=Nemetschek |first1=Antje |last2=Mörs |first2=Thomas |date=January 2003 |title=Myoglis meini (de Bruijn, 1965 [1966]) (Mammalia: Gliridae) aus dem Miozän von Hambach 6C (NW-Deutschland) |url=https://www.researchgate.net/publication/226876010 |journal=Paläontologische Zeitschrift |language=de |volume=77 |issue=2 |pages=401–416 |doi=10.1007/BF03006950 |s2cid=131075063 |issn=0031-0220}}

|Middle Miocene.

|Numerous teeth.

|A dormouse.

|

Steneofiber

|S. depereti

|Hambach 6C.{{Cite journal |last1=Mörs |first1=Thomas |last2=Stefen |first2=Clara |date=11 January 2010 |title=The Castorid Steneofiber from NW Germany and Its Implications for the Taxonomy of Miocene Beavers |journal=Acta Palaeontologica Polonica |language=en |volume=55 |issue=2 |pages=189–198 |doi=10.4202/app.2009.0013 |issn=0567-7920|doi-access=free }}

|Middle Miocene.

|6 mandibles and 56 isolated teeth.

|A castorid.

|File:Steneofiber.jpg

==Ungulates==

class="wikitable" align="center"
colspan="7" align="center" | Ungulates reported from the Ville Formation
GenusSpeciesPresence

!Age

!Material

NotesImages
Orygotherium

|O. escheri

| Hambach 6C.{{Cite journal |last1=Rössner |first1=Gertrud E. |last2=Mörs |first2=Thomas |date=2001-08-22 |title=A new record of the enigmatic Eurasian Miocene ruminant artiodactyl Orygotherium |url=http://www.tandfonline.com/doi/abs/10.1671/0272-4634%282001%29021%5B0591%3AANROTE%5D2.0.CO%3B2 |journal=Journal of Vertebrate Paleontology |language=en |volume=21 |issue=3 |pages=591–595 |doi=10.1671/0272-4634(2001)021[0591:ANROTE]2.0.CO;2 |s2cid=85657626 |issn=0272-4634|url-access=subscription }}

|Middle Miocene.

|An isolated molar.

| A ruminant.

|

=Reptiles=

==Squamates==

class="wikitable" align="center"
colspan="7" align="center" | Squamates reported from the Ville Formation
GenusSpeciesPresence

!Age

!Material

NotesImages
rowspan=2|Anguidae

|Anguidae indet. 1

|Hambach 6C.

|Middle Miocene.

|20 osteoderms (IPB-HaH 3700), caudal vertebra (IPB-HaH 3709).

|An anguid lizard.

|

Anguidae indet. 2

|Hambach 11.

|Upper Pliocene.

|2 caudal vertebrae (IPB-HaR 3009).

|An anguid lizard.

|

Bavarioboa

|B. sp.

|Hambach 6C.

|Middle Miocene.

|5 trunk vertebrae.

|A boid snake.

|

Chamaeleo

|C. aff. andrusovi

| Hambach 6C.

|Middle Miocene.

|Right squamosal (IPB-HaH 3900).

| A chameleon.

|

rowspan=2|?Chamaeleonidae

|Tooth morphotype 1.

|Hambach 6C.

|Middle Miocene.

|Right maxilla (IPB-HaH 3901) & left maxilla (IPB-HaH 3902).

|A chameleon.

|

Tooth morphotype 2.

|Hambach 6C.

|Middle Miocene.

|Right maxilla (IPB-HaH 3903).

|A chameleon.

|

"Coluber"

|"C." sp.

|Hambach 6C.

|Middle Miocene.

|42 trunk vertebrae.

|A colubrid snake.

|

rowspan=2|"Colubrinae"

|"Colubrinae" indet. 1

|Hambach 6C.

|Middle Miocene.

|57 trunk vertebrae.

|A colubrine snake.

|

"Colubrinae" indet. 2

|Hambach 11.

|Upper Pliocene.

|A trunk vertebra (IPB-HaR 3001).

|A colubrine snake.

|

Colubroidea

|Colubroidea indet.

|Hambach 6C.

|Middle Miocene.

|36 caudal vertebrae.

|A colubroid snake.

|

Eoanilius

|E. sp.

|Hambach 6C.

|Middle Miocene.

|5 trunk vertebrae.

|An aniliid snake.

|

cf. Falseryx

|cf. F. sp.

|Hambach 6C.

|Middle Miocene.

|A trunk vertebra (HaH-3830).

|A tropidophiid snake.

|

rowspan="2" |Lacertidae

|Lacertidae indet. 1

|Hambach 6C.

|Middle Miocene.

|Left maxilla (IPB-HaH 3701), two left dentaries (IPB-HaH 3702–3703).

|A lacertid lizard.

|

Lacertidae indet. 2

|Hambach 6C.

|Middle Miocene.

|Premaxilla (IPB-HaH3704), left dentary (IPB-HaH 3705).

|A lacertid lizard.

|

cf. Naja

|cf. N. sp.

|Hambach 6C.

|Middle Miocene.

|6 trunk vertebrae.

|A cobra.

|

"Natricinae"

|Indeterminate

|Hambach 6C.

|Middle Miocene.

|9 trunk vertebrae.

|A natricine snake.

|

rowspan=2|Natrix

|N. sp. 1

|Hambach 6C.

|Middle Miocene.

|2 trunk vertebrae (IPB-HaH 3842; 3843).

|A natricine snake.

|

N. sp. 2

|Hambach.

|Upper Pliocene.

|4 trunk vertebrae (IPB-HaR 3010; 3401).

|A natricine snake.

|

rowspan="4" |Pseudopus

|P. cf. ahnikoviensis

|Hambach 6C.

|Middle Miocene.

|Right dentary (IPB-HaH3706), left dentary (IPB-HaH 3707).

|A glass lizard, material likely represents a juvenile.

|

P. cf. pannonicus

|Hambach 11 & 13.

|Upper Pliocene.

|Jaw elements.

|A glass lizard.

|

P. sp. 1

|Hambach 6C.

|Middle Miocene.

|Dorsal vertebra (IPB-HaH 3708).

|A glass lizard.

|

P. sp. 2

|Hambach 11 & 13.

|Upper Pliocene.

|Right nasal, vertebrae & osteoderms.

|A glass lizard.

|

Telescopus

|T. sp.

|Hambach 6C.

|Middle Miocene.

|A trunk vertebra (IPB-HaH 3853).

|A colubrid snake.

|

Texasophis

|T. sp.

|Hambach 6C.

|Middle Miocene.

|2 trunk vertebrae (IPB-HaH 3832; 3841).

|A colubrid snake.

|

rowspan=2|Vipera

|V. sp. ("Oriental viper")

|Hambach 6C.

|Middle Miocene.

|8 trunk vertebrae.

|A viper.

|

V. sp. ("aspis complex")

|Hambach 6C.

|Middle Miocene.

|3 trunk vertebrae.

|A viper.

|

==Testudines==

class="wikitable" align="center"
colspan="7" align="center" | Testudines reported from the Ville Formation
GenusSpeciesPresence

!Age

!Material

NotesImages
Carettochelyidae

|

| Hambach 6C.{{Cite journal |last1=Joyce |first1=Walter G. |last2=Klein |first2=Nicole |last3=Mörs |first3=Thomas |date=2004 |title=Carettochelyine Turtle from the Neogene of Europe |url=https://www.academia.edu/12924044 |journal=Copeia |volume=2004 |issue=2 |pages=406–411 |doi=10.1643/CH-03-172R |s2cid=86333324 |issn=0045-8511}}

|Middle Miocene.

|A peripheral bone (IPB HaH 3028).

| A carettochelyid turtle.

|

=Amphibians=

class="wikitable" align="center"
colspan="7" align="center" | Amphibians reported from the Ville Formation
GenusSpeciesPresence

!Age

!Material

NotesImages
Euronecturus

|E. grogu

| Hambach 6C.{{Cite journal |last1=Macaluso |first1=Loredana |last2=Villa |first2=Andrea |last3=Mörs |first3=Thomas |date=January 2022 |editor-last=Mannion |editor-first=Philip |title=A new proteid salamander (Urodela, Proteidae) from the middle Miocene of Hambach (Germany) and implications for the evolution of the family |url=https://onlinelibrary.wiley.com/doi/10.1111/pala.12585 |journal=Palaeontology |language=en |volume=65 |issue=1 |doi=10.1111/pala.12585 |s2cid=245471400 |issn=0031-0239|hdl=2318/1825718 |hdl-access=free }}

|Late Orleanian (early Middle Miocene).

|Multiple specimens.

| A proteid salamander.

|

See also

{{Portal|Earth sciences|Germany|Paleontology}}

References

{{Reflist}}

  • {{cite web|title= Fossilworks: Gateway to the Paleobiology Database|author= ((Various Contributors to the Paleobiology Database))|url= https://www.fossilworks.org|access-date= 17 December 2021|df= dmy-all}}

Category:Neogene Germany

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