Viverravidae
{{Short description|Extinct family of carnivorous mammals}}
{{About|an extinct family of carnivorous mammals|the extant family of carnivorans|Viverridae}}
{{Automatic taxobox
|fossil_range = {{fossil range|66.043|33.9}} early Paleocene - late Eocene
|image = Viverravus minutus 2.jpg
|image_caption = skull of Viverravus minutus
|image2 = Didymictis protenus.jpg
|image2_caption = skull of Didymictis protenus
|display_parents = 3
|taxon = Viverravidae
|parent_authority = Wortman & Matthew, 1899
|authority = Wortman & Matthew, 1899{{Cite journal |last1=Wortman |first1=J. L. |last2=Matthew |first2=W. D. |author2-link=William Diller Matthew |title=The ancestry of certain members of the Canidae, Viverridae, and Procyonidae |year=1899 |journal=Bulletin of the American Museum of Natural History |volume=12 |pages=109–138 |oclc=46687698 |hdl=2246/1535}}
|type_genus = †Viverravus
|type_genus_authority = Marsh, 1872
|subdivision_ranks = Genera
|subdivision = {{center|[see classification]}}
|synonyms =
{{collapsible list |bullets=true |title=synonyms of family:
|Didymictida {{small|(Kretzoi, 1945)}}Miklos Kretzoi (1945) "Bemerkungen über das Raubtiersystem." Annales Historico-Naturales Musei Nationalis Hungarici, Budapest, vol. 38, pp. 59–83.
|Didymictidae {{small|(Flynn & Galiano, 1982)}}
|Viverravidea
|Viverravinae {{small|(Matthew, 1909)}}W. D. Matthew (1909) [https://digitallibrary.amnh.org/handle/2246/5744 "The Carnivora and Insectivora of the Bridger Basin, middle Eocene."] Memoirs of the American Museum of Natural History 9:289-567
}}
}}
Viverravidae ("ancestors of viverrids") is an extinct monophyletic family of mammals from extinct superfamily Viverravoidea within the clade Carnivoramorpha, that lived from the early Palaeocene to the late Eocene in North America, Europe and Asia.{{BioRef |fossilworks |title=†family Viverravidae (Wortman & Matthew, 1899) (placental) |id=40992 |access-date=29 June 2019}}Malcolm C. McKenna, Susan K. Bell: Classification of Mammals: Above the Species Level in Columbia University Press, New York (1997), 631 Seiten. {{page|year=|isbn=0-231-11013-8|pages=}}{{cite journal |last1=Zack |first1=Shawn P. |title=Deciduous dentition of Didymictis (Carnivoramorpha: Viverravidae): implications for the first appearance of "Creodonta" |journal=Journal of Mammalogy |volume=93 |issue=3 |year=2012 |pages=808–817 |issn=0022-2372 |doi=10.1644/11-MAMM-A-245.1 |doi-access=free}} They were once thought to be the earliest carnivorans and ancestral to extant ones, but now are placed outside the order Carnivora based on cranial morphology as relatives (a plesion-group) to extant carnivorans.{{Cite journal |last1=Wesley-Hunt |first1=G. D. |last2=Flynn |first2=J. J. |title=Phylogeny of the Carnivora: basal relationships among the carnivoramorphans, and assessment of the position of 'Miacoidea' relative to Carnivora |year=2005 |journal=Journal of Systematic Palaeontology |volume=3 |pages=1–28 |doi=10.1017/S1477201904001518 |s2cid=86755875}}{{cite journal |author=Polly, David, Gina D. Wesley-Hunt, Ronald E. Heinrich, Graham Davis and Peter Houde |year=2006 |title=Earliest known carnivoran auditory bulla and support for a recent origin of crown-clade carnivora (Eutheria, Mammalia) |journal=Palaeontology |volume=49 |issue=5 |pages=1019–1027 |url=http://mypage.iu.edu/~pdpolly/Papers/Polly%20et%20al,%202006,%20Viverravus.pdf |doi=10.1111/j.1475-4983.2006.00586.x |doi-access=free}}
General characteristics
Wang and Tedford propose that they arose in North America 66–60 million years ago, spread to Asia then later to Europe, and were the first carnivoramorphans and possessed the first true pair of carnassial teeth.Wang, Xiaoming; Tedford, Richard H. (2008.) "Dogs: Their Fossil Relatives and Evolutionary History." New York: Columbia University Press In viverravids, the skull is elongated and the number of molars is reduced to two (M1/m1 and M2/m2 are present and M3/m3 are absent).
Classification and phylogeny
=Classification=
Taxonomy retrieved from the Paleobiology DatabasePaleobiology Database. Retrieved with [https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=40992 Fossilworks] (March 3, 2017)
class="wikitable" |
style="vertical-align:top;"
|style="text-align: center;"|Superfamily: †Viverravoidea {{small|(Wortman & Matthew, 1899)}} ---- :{| |
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=Phylogeny=
The phylogenetic relationships of family Viverravidae are shown in the following cladogram:J. J. Flynn and H. Galiano. (1982.) "Phylogeny of Early Tertiary Carnivora, With a Description of a New Species of Protictis From the Middle Eocene of Northwestern Wyoming" American Museum Novitates 2725:1-64P. D. Gingerich and D. A. Winkler. (1985.) [https://deepblue.lib.umich.edu/bitstream/handle/2027.42/48525/ID378.pdf?sequence=2&isAllowed=y "Systematics of Paleocene Viverravidae (Mammalia, Carnivora) in the Bighorn Basin and Clark's Fork Basin, Wyoming."] Contributions from the Museum of Paleontology, University of Michigan 27(4):87-128P. D. Polly. (1997.) [https://deepblue.lib.umich.edu/bitstream/handle/2027.42/48638/ID504.pdf?sequence=2 "Ancestry and Species Definition in Paleontology: A Stratocladistic Analysis of Paleocene-Eocene Viverravidae (Mammalia, Carnivora) from Wyoming."] Contributions from the Museum of Paleontology, University of Michigan 30(1):1-53{{cite journal |last1=Solé |first1=Floréal |last2=Smith |first2=Thierry |last3=De Bast |first3=Eric |last4=Codrea |first4=Vlad |last5=Gheerbrant |first5=Emmanuel |title=New carnivoraforms from the latest Paleocene of Europe and their bearing on the origin and radiation of Carnivoraformes (Carnivoramorpha, Mammalia) |journal=Journal of Vertebrate Paleontology |volume=36 |issue=2 |year=2016 |pages=e1082480 |issn=0272-4634 |doi=10.1080/02724634.2016.1082480|s2cid=87537565 }}S. Faurby, L. Werdelin, A. Antonelli (2019.) [https://www.biorxiv.org/content/biorxiv/early/2019/09/05/755207.full.pdf "Dispersal ability predicts evolutionary success among mammalian carnivores"] Department of Biological and Environmental Sciences, University of Gothenburg, Box 461, SE
{{clade |style=font-size:85%;line-height:100%;
|label1=Carnivoramorpha
|1={{clade
|thickness2=2 |thickness3=2 |thickness4=2
|label2=?|state2=dotted
|2=†Carnivoramorpha sp. (UALVP 31176)
|label3=?|state3=dotted
|3=†Carnivoramorpha sp. (USNM 538395)
|label4=?|state4=dotted
|label5=†Viverravidae|sublabel5=sensu lato
|5={{clade
|thickness1=2 |thickness2=2 |thickness3=2 |label1=?|state1=dotted
|1=†Ravenictis
|label2=?|state2=dotted
|2=†Carnivoramorpha sp. (UALVP 50993 & UALVP 50994)
|label3=†Viverravoidea
|3={{clade
|thickness=2 |label1=†Viverravidae|sublabel1=sensu stricto
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1={{clade
|thickness=2 |label1=†Pappictidops
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1=†Pappictidops acies
|2=†Pappictidops obtusus
}}
|2=†Pappictidops orientalis
}}
|label2=†Orientictis
|2=†Orientictis spanios
|label3=†Preonictis
|3=†Preonictis youngi
|label4=†Variviverra
|4=†Variviverra vegetatus
}}
|2={{clade
|thickness=2 |label1=†Ictidopappinae
|1={{clade
|thickness=2 |label1=†Ictidopappus
|1=†Ictidopappus mustelinus
}} }} }}
|2={{clade
|thickness=2 |label1=†Viverravinae
|1={{clade
|thickness=2 |label1=†Viverravus
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1={{clade
|thickness=2
|1=†Viverravus minutus
|2=†Viverravus sicarius
|3=†Viverravus sp. (Locality Group 2, Washakie Basin, Wyoming)
}}
|2=†Viverravus gracilis
|3=†Viverravus lutosus
}}
|2=†Viverravus acutus
|3=†Viverravus rosei
}}
|2=†Viverravus laytoni
}}
|label2=?|state2=dotted
|2=†Viverravus sp. (V11141)
|3=†Viverravus lawsoni
|4=†Viverravus politus
}}
|2={{clade
|thickness=2 |label1=†Viverriscus
|1=†Viverriscus omnivorus
|label2=†Simpsonictis
|2={{clade
|thickness=2
|1=†Simpsonictis jaynanneae
|2=†Simpsonictis pegus
|3=†Simpsonictis tenuis
}} }} }}
|label2=?|state2=dotted
|2=†Viverravidae sp. (CM 71188 & CM 71189)
|label3=†Didymictinae
|3={{clade
|thickness=2
|1={{clade
|thickness=2 |label1=†Bryanictis
|1={{clade
|thickness=2
|1=†Bryanictis microlestes
|2=†Bryanictis paulus
|3=†Bryanictis terlinguae
}}
|label2=†Intyrictis
|2=†Intyrictis vanvaleni
}}
|2={{clade
|thickness=2 |caption={{Uline|c=green|†Protictis}} |captionstyle=text-align:center;font-weight:bold;padding:2px 10px 1px 1px; |color=green
|1={{Uline|c=green|†Protictis haydenianus}}
|2={{Uline|c=green|†Protictis minor}}
|3={{Uline|c=green|†Protictis simpsoni}}
|4={{clade
|thickness=2 |color=green
|1={{clade
|thickness=2 |label1=†Raphictis|color=green |color1=black
|1={{clade
|thickness=2
|1=†Raphictis gausion
|2=†Raphictis iota
|3=†Raphictis machaera
|4=†Raphictis nanoptexis
}}
|2={{Uline|c=green|†Protictis paralus}}
}}
|2={{clade
|thickness=2 |color=green |color3=black |color4=black |color5=black
|1={{Uline|c=green|†Protictis agastor}}
|label2=†(Protictoides)
|2={{Uline|c=green|†Protictis aprophatos}}
|label3=†Pristinictis
|3=†Pristinictis connata
|label4=?|state4=dotted
|label5=†Didymictis
|5={{clade
|thickness=2
|1=†Didymictis dellensis
|2=†Didymictis proteus
|label3=?|state3=dotted
|3=†Didymictis sp. (Erquelinnes, Hainaut, Belgium)
|4={{clade
|thickness=2
|1=†Didymictis altidens
|2=†Didymictis leptomylus
|3=†Didymictis protenus
|4=†Didymictis vancleveae
}} }} }} }} }} }} }} }} }} }} }} }}
See also
References
{{Reflist|2}}
{{Pan-Carnivora|C.|state=collapsed}}
{{Taxonbar|from=Q4889586}}
Category:Paleocene first appearances
Category:Prehistoric mammal families
Category:Long stubs with short prose
{{Paleo-carnivora-stub}}