Wryneck#Subspecies

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{{Automatic taxobox

| name = Wrynecks

| image = Wryneck, Village, Behlolpur, district Mohali, Punjab, India.JPG|

| image_caption = Eurasian wryneck (Jynx torquilla) Punjab, India

| taxon = Jynx

| authority = Linnaeus, 1758

| type_species = Jynx torquilla (Eurasian wryneck)

| type_species_authority = Linnaeus, 1758

| subdivision_ranks = Species

| subdivision =

Jynx torquilla

Jynx ruficollis

| range_map = Jynx torquilla distr .png

| range_map_caption = Range map for Eurasian wryneck

| range_map2 = Jynx ruficollis map.jpg

| range_map2_caption = Range map for red-necked wryneck
{{legend0|#F7A229|Summer|border=solid 1px black}} {{legend0|#53A27B|Resident|border=solid 1px black}} {{legend0|#5379F6|Winter|border=solid 1px black}}

}}

The wrynecks (genus Jynx) are a small but distinctive group of small Old World woodpeckers. Jynx is from the Ancient Greek iunx, the Eurasian wryneck.

These birds get their English name from their ability to turn their heads almost 180°. When disturbed at the nest, they use this snake-like head twisting and hissing as a threat display. It has occasionally been called "snake-bird" for that reason.{{Cite EB1911|wstitle=Wryneck|volume=28|page=854|first=Alfred|last=Newton|author-link=Alfred Newton}}

Like the true woodpeckers, wrynecks have large heads, long tongues, which they use to extract their insect prey, and zygodactyl feet, with two toes pointing forward and two backwards, but they lack the stiff tail feathers that the true woodpeckers use when climbing trees, so they are more likely than their relatives to perch on a branch rather than an upright trunk. The sexes have a similar appearance.{{Cite web|url=https://www.encyclopedia.com/environment/encyclopedias-almanacs-transcripts-and-maps/woodpeckers-wrynecks-and-piculets-picidae|title=Woodpeckers, Wrynecks, and Piculets (Picidae) | Encyclopedia.com|website=www.encyclopedia.com|access-date=28 March 2023}}

Their bills are shorter and less dagger-like than in the true woodpeckers, but their chief prey is ants and other insects, which they find in decaying wood or almost bare soil. They reuse woodpecker holes for nesting, rather than making their own holes. The eggs are white, as with many hole nesters.

The two species have cryptic plumage, with intricate patterning of greys and browns. The adult moults rapidly between July and September, although some moult continues in its winter quarters.RSPB Handbook of British Birds (2014). UK. {{ISBN|978-1-4729-0647-2}}.

Taxonomy and etymology

The woodpeckers are an ancient bird family consisting of three subfamilies, the wrynecks, the piculets and the true woodpeckers, Picinae. DNA sequencing and phylogenetic analysis show that the wrynecks are a sister clade to other woodpeckers including the Picinae and probably diverged early from the rest of the family.Gorman (2022) p. 3.

The wryneck subfamily Jynginae has one genus, Jynx, introduced in 1758 by Swedish naturalist Carl Linnaeus in the 10th edition of his Systema Naturae.{{cite book |last=Linnaeus |first=Carl |author-link=Carl Linnaeus |year=1758 |title= Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis |volume=1 |edition=10th |page=112 |publisher=Laurentii Salvii |location=Holmiae (Stockholm) |language=Latin |url=https://www.biodiversitylibrary.org/page/727017 }} Linnaeus placed a single species in the genus, the Eurasian wryneck (Jynx torquilla), which is therefore the type species.{{ cite book | editor-last=Peters | editor-first=James Lee | editor-link=James L. Peters | year=1948 | title=Check-List of Birds of the World | volume=6 | publisher=Harvard University Press | place=Cambridge, Massachusetts | page=86 | url=https://www.biodiversitylibrary.org/page/14477519 }} The genus name Jynx is from the Ancient Greek name for the Eurasian wryneck, ιυγξ, iunx, and ruficollis is from the Latin {{Lang|la|rufus}}, "rufous" and {{Lang|la|collum}} "neck".{{cite book |last=Jobling |first=James A |year=2010 |title=The Helm Dictionary of Scientific Bird Names |url=https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling |publisher=Christopher Helm |location=London |isbn=978-1-4081-2501-4 |pages=[https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling/page/n212 212], [https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling/page/n341/mode/2up 341] }} The English "wryneck" refers to the habit of birds in this genus of twisting and writhing their necks when agitated. It was first recorded in 1585.{{OED|Wryneck}}

The red-throated wryneck was first identified by German ornithologist Johann Georg Wagler in 1830.{{cite book |last1=Wagler |first1=Johann Georg |title=Naturliches System der Amphibien |publisher=J. G. Cotta'schen |series= |volume= |edition= |year=1830 |location=Munich |page=118 |language=German |url=https://www.biodiversitylibrary.org/page/48675921#page/126/mode/1up |isbn= }}Gorman (2022) pp. 35–36. It is also known as the rufous-necked wryneck or red-breasted wryneck.Gorman (2014) pp. 38–39.

The two wrynecks form a superspecies that probably separated early in their evolution from the piculets, although there has subsequently been only limited divergence between the Jynx species.Gorman (2022) pp. 39–40.{{cite journal |last1=Tarboton |first1=Warwick |title=Aspects of the Biology of Jynx ruficollis |year=1976 |volume=47 |issue=2–3 |journal=The Ostrich: Journal of African Ornithology |pages=99–112 |doi=10.1080/00306525.1976.9639545 |bibcode=1976Ostri..47...99T }}

=Fossil record=

The woodpecker family appears to have diverged from other Piciformes about fifty million years ago, and a 2017 study considered that the split between Jynx and other woodpeckers occurred about 22.5{{nbs}}million years ago.{{cite journal |last1=Shakya |first1=S B |last2=Fuchs |first2=J |last3=Pons |first3=J-M |last4=Sheldon |first4=F H |year=2017 |title=Tapping the woodpecker tree for evolutionary insight |journal=Molecular Phylogenetics and Evolution |volume=116 |issue= |pages=182–191 |doi=10.1016/j.ympev.2017.09.005 |pmid=28890006 |doi-access=free }} A fossil dating from the early Miocene, more than twenty million years ago, consisting of the distal end of a tarsometatarsus had some ‘’Jynx’’-like features, but was classed as an early piculet.{{cite journal |last1=De Pietri |first1=Vanesa L |last2=Manegold |first2=Albrecht |last3=Costeur |first3=Loïc |last4=Mayr |first4=Gerald |year=2011 |title=A new species of woodpecker (Aves; Picidae) from the early Miocene of Saulcet (Allier, France) |journal=Swiss Journal of Palaeontology |volume=130 |issue=2 |pages=307–314 |doi=10.1007/s13358-011-0021-8 |s2cid=129249316 |doi-access=free |bibcode=2011SwJP..130..307D }} By the Pliocene (five million years ago) woodpeckers were similar to those now extant. Fossil wrynecks are known from Europe in the Pleistocene, between 2.6{{nbs}}million and 11,700 years ago.

=Species=

The two species in Jynx are restricted to the Palearctic biogeographic realm and Africa. The Eurasian wryneck breeds across temperate Europe and Asia, and one of only two Old World woodpeckers to undertake long-distance migration mainly wintering in sub-Saharan Africa and tropical Asia.Gorman (2022) pp. 104–106. The rufous-necked wryneck has a disjunct distribution confined to sub-Saharan Africa. It is resident, although there may be local movements and post-breeding dispersal.Gorman (2022) p. 42. Both wrynecks have several geographical subspecies.Gorman (2022) pp. 4–6.

{{Species table |genus=Jynx |authority-name=Linnaeus |authority-year=1758 |species-count=two|no-note=y|narrow-percent=75}}

{{Species table/row

|name= Eurasian wryneck|binomial=Jynx torquilla

|image=File:Wryneck by Pepe Reigada.jpg|image-size=180px |image-alt=

|authority-name=Linnaeus |authority-year=1758 |authority-not-original=

|range= Palearctic from the Arctic Circle south to Spain, Algeria, Morocco, southern Siberia, Central Asia, Japan and southern China.{{cite web |url=http://www.birdlife.org/datazone/speciesfactsheet.php?id=538 |title=Eurasian Wryneck Jynx torquilla |author1=Butchart, S. |author2=Ekstrom, J. |publisher=BirdLife International |access-date=2013-08-06 |archive-date=2016-10-21 |archive-url=https://web.archive.org/web/20161021202343/http://www.birdlife.org/datazone/speciesfactsheet.php?id=538 |url-status=dead }} Winters south of the Sahara from Senegal in the west to Ethiopia in the east, and in tropical South and Southeast Asia.

|range-image=File:Jynx torquilla distr.png

|range-image-size=180px

|size={{convert|17|cm|in|abbr=on}}.{{cite book |title=Handbook of British Birds, Volume 2: Warblers to Owls |editor-last=Witherby |editor-first=H. F. |year=1943 |publisher=H. F. and G. Witherby Ltd. |pages=292–296 }}
{{convert|26|to|50|g|oz|abbr=on}}.CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), {{ISBN|978-0-8493-4258-5}}.

|habitat=Open countryside and gardens, especially with some old trees.

|hunting=Mainly ants

|iucn-status= LC

|population=

|direction=

|subspecies={{Collapsible list |expand=yes |title=Six subspecies |bullets=on

|J. t. torquilla Linnaeus, 1758

|J. t. chinensis Hesse, 1911

|J. t. himalayana Vaurie, 1959

|J. t. mauretanica Rothschild, 1909

|J. t. sarudnyi Loudon, 1912

|J. t. tschusii Kleinschmidt, 1907

}}

{{Species table/row

|name=Red-throated wryneck |binomial=Jynx ruficollis

|image=File:Red-throated Wryneck, Jynx ruficollis at Rietvlei Nature Reserve, Gauteng, South Africa (14564203969).jpg|image-size=180px |image-alt=

|authority-name=Wagler |authority-year=1830 |authority-not-original=yes

|range= Resident in sub-Saharan Africa from Nigeria, Cameroon, Central African Republic and Ethiopia south to South Africa and Eswatini.{{cite iucn |author=BirdLife International |date=2016 |title=Jynx ruficollis |volume=2016 |page=e.T22680689A92872725 |doi=10.2305/IUCN.UK.2016-3.RLTS.T22680689A92872725.en |access-date= }}

|range-image=File:Jynx ruficollis map.jpg

|range-image-size=180px

|size={{convert|19|cm|in|abbr=on}} in length. Male{{convert|52|–|59|g|oz|abbr=on}} Female {{convert|46|–|52|g|oz|abbr=on}}

|habitat=

|hunting=

|iucn-status= LC

|population=

|direction=

|subspecies={{Collapsible list |expand=yes |title=Three subspecies |bullets=on

|J. r. ruficollis Wagler, 1830

|J. r. aequatorialis Ruppell, 1842

|J. r. pulchricollis Hartlaub, 1884

}}

References

Cited texts

{{cite book |last1=Gorman |first1=Gerard |title=Woodpeckers of the World |publisher=Christopher Helm |series=Helm Photographic Guides |year=2014 |location=London |isbn=978-1-4081-4715-3 |ref=none }}
{{cite book|last1=Gorman|first1=Gerard|title=The Wryneck |year=2022 |publisher=Pelagic |location=Exeter |isbn=978-1-78427-288-3 |ref=none }}

External links

[http://aulaenred.ibercaja.es/wp-content/uploads/287_WryneckJtorquilla.pdf Ageing and sexing of Eurasian wryneck (PDF; 4.6 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze]

Category:Picidae