arctostaphylos
{{Short description|Genus of flowering plants in the heath family}}
{{See also|Manzanita}}
{{Automatic taxobox
|image = Arctostaphylos patula 08154.JPG
|image_caption = Whiteleaf manzanita (Arctostaphylos viscida)
|taxon = Arctostaphylos
|authority = Adanson
|subdivision_ranks = Species
|subdivision = About 60, see text.
|type_species = Arctostaphylos uva-ursi
|type_species_authority = (Linnaeus) Sprengel
}}
Arctostaphylos ({{IPAc-en|ˌ|ɑːr|k|t|oʊ|ˈ|s|t|æ|f|ᵻ|l|ə|s|,_|-|l|ɒ|s}};{{refn|{{MerriamWebsterDictionary|access-date=2016-01-21|Arctostaphylos}}}}Sunset Western Garden Book, 1995:606–607 from {{wikt-lang|grc|ἄρκτος}} {{grc-tr|ἄρκτος}} "bear" and {{wikt-lang|grc|σταφυλή}} {{grc-tr|σταφυλή}} "bunch of grapes") is a genus of plants comprising the manzanitas ({{IPAc-en|ˌ|m|æ|n|z|ə|ˈ|n|iː|t|ə|z}}{{refn|{{MerriamWebsterDictionary|access-date=2016-01-21|manzanita}}}}{{refn|{{Cite encyclopedia |url=http://www.lexico.com/definition/manzanita |archive-url=https://web.archive.org/web/20200322182747/https://www.lexico.com/definition/manzanita |url-status=dead |archive-date=2020-03-22 |title=manzanita |dictionary=Lexico UK English Dictionary |publisher=Oxford University Press}} }}) and bearberries. There are about 60 species of Arctostaphylos, ranging from ground-hugging arctic, coastal, and mountain shrub to small trees up to {{Convert|6|m|abbr=on}} tall. Most are evergreen (one species deciduous), with small oval leaves {{Convert|1-7|cm|abbr=on|sigfig=1}} long, arranged spirally on the stems. The flowers are bell-shaped, white or pale pink, and borne in small clusters of 2–20 together; flowering is in the spring. The fruit are small berries, ripening in the summer or autumn. The berries of some species are edible.
Arctostaphylos species are used as food plants by the larvae of some Lepidoptera species including Coleophora arctostaphyli (which feeds exclusively on A. uva-ursi) and Coleophora glaucella.
Distribution
File:Arctostaphylos nevadensis 7744.JPG to California.]]
File:Arctostaphylos uva-ursi 3 RF.jpg
Manzanitas, the bulk of Arctostaphylos species, are present in the chaparral biome of western North America, where they occur from southern British Columbia in Canada, Washington to California and New Mexico in the United States, and throughout much of northern and central Mexico.
One species, however, A. uva-ursi (common bearberry), is adapted to arctic and subarctic climates and has a circumpolar distribution in northern North America, Asia and Europe.
An unusual association of manzanita occurs on Hood Mountain, in Sonoma County, California, where stands of pygmy forest dominated by Hesperocyparis sargentii are found.
Fossil record
One fossil fruit of †Arctostaphylos globula and several fossil fruits of †Arctostaphylos menzelii have been described from middle Miocene strata of the Fasterholt area near Silkeborg in Central Jutland, Denmark.{{cite journal| title=Angiosperm fruits and seeds from the Middle Miocene of Jutland (Denmark) |first=Else Marie |last=Friis |journal=Det Kongelige Danske Videnskaberne Selskab, Biologiske Skrifter |volume=24 |issue=3 |pages=1–165 |year=1985 |url=https://www.royalacademy.dk/Publications/Low/369_Friis,%20Else%20Marie.pdf }}
Cultivation
Taxonomy
The following species are recognised in the genus Arctostaphylos:{{Cite web |title=Arctostaphylos Adans. {{!}} Plants of the World Online {{!}} Kew Science |url=http://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:331363-2 |access-date=2022-04-29 |website=Plants of the World Online |language=en}}
{{div col|colwidth=22em}}
- Arctostaphylos acutifolia {{small|Eastw.}}
- Arctostaphylos andersonii {{small|A.Gray}}
- Arctostaphylos auriculata {{small|Eastw.}}
- Arctostaphylos australis {{small|Eastw.}}
- Arctostaphylos bakeri {{small|Eastw.}}
- Arctostaphylos bolensis {{small|P.V.Wells}}
- Arctostaphylos canescens {{small|Eastw.}}
- Arctostaphylos catalinae {{small|P.V.Wells}}
- Arctostaphylos caucasica {{small|Lipsch.}}
- Arctostaphylos columbiana {{small|Piper}}
- Arctostaphylos confertiflora {{small|Eastw.}}
- Arctostaphylos cratericola {{small|(Donn.Sm.) Donn.Sm.}}
- Arctostaphylos crustacea {{small|Eastw.}}
- Arctostaphylos cruzensis {{small|Roof}}
- Arctostaphylos densiflora {{small|M.S.Baker}}
- Arctostaphylos edmundsii {{small|J.T.Howell}}
- Arctostaphylos franciscana {{small|Eastw.}}
- Arctostaphylos gabilanensis {{small|V.T.Parker & M.C.Vasey}}
- Arctostaphylos glandulosa {{small|Eastw.}}
- Arctostaphylos glauca {{small|Lindl.}}
- Arctostaphylos glutinosa {{small|B.Schreib.}}
- Arctostaphylos × helleri {{small|Eastw.}}
- Arctostaphylos hispidula {{small|Howell}}
- Arctostaphylos hookeri {{small|G.Don}}
- Arctostaphylos hooveri {{small|P.V.Wells}}
- Arctostaphylos imbricata {{small|Eastw.}}
- Arctostaphylos incognita {{small|J.E.Keeley, Massihi, J.Delgad. & Hirales}}
- Arctostaphylos insularis {{small|Greene & Parry}}
- Arctostaphylos × jepsonii {{small|Eastw.}}
- Arctostaphylos klamathensis {{small|S.W.Edwards, Keeler-Wolf & W.Knight}}
- Arctostaphylos × laxiflora {{small|A.Heller ex Eastw.}}
- Arctostaphylos luciana {{small|P.V.Wells}}
- Arctostaphylos malloryi {{small|(W.Knight & Gankin) P.V.Wells}}
- Arctostaphylos manzanita {{small|Parry}}
- Arctostaphylos × media {{small|Greene}}
- Arctostaphylos mewukka {{small|Merriam}}
- Arctostaphylos montana {{small|Eastw.}}
- Arctostaphylos montaraensis {{small|Roof}}
- Arctostaphylos montereyensis {{small|Hoover}}
- Arctostaphylos moranii {{small|P.V.Wells}}
- Arctostaphylos morroensis {{small|Wiesl. & B.Schreib.}}
- Arctostaphylos myrtifolia {{small|Parry}}
- Arctostaphylos nevadensis {{small|A.Gray}}
- Arctostaphylos nissenana {{small|Merriam}}
- Arctostaphylos nortensis {{small|(P.V.Wells) P.V.Wells}}
- Arctostaphylos nummularia {{small|A.Gray}}
- Arctostaphylos obispoensis {{small|Eastw.}}
- Arctostaphylos ohloneana {{small|M.C.Vasey & V.T.Parker}}
- Arctostaphylos osoensis {{small|P.V.Wells}}
- Arctostaphylos otayensis {{small|Wiesl. & B.Schreib.}}
- Arctostaphylos pacifica {{small|Roof}}
- Arctostaphylos pajaroensis {{small|J.E.Adams}}
- Arctostaphylos pallida {{small|Eastw.}}
- Arctostaphylos parryana {{small|Lemmon}}
- Arctostaphylos × parvifolia {{small|Howell}}
- Arctostaphylos patula {{small|Greene}}
- Arctostaphylos pechoensis {{small|(Abrams) Dudley ex Abrams}}
- Arctostaphylos peninsularis {{small|P.V.Wells}}
- Arctostaphylos pilosula {{small|Jeps. & Wiesl.}}
- Arctostaphylos pringlei {{small|Parry}}
- Arctostaphylos pumila {{small|Nutt.}}
- Arctostaphylos pungens {{small|Kunth}}
- Arctostaphylos purissima {{small|P.V.Wells}}
- Arctostaphylos rainbowensis {{small|J.E.Keeley & Massihi}}
- Arctostaphylos refugioensis {{small|Gankin}}
- Arctostaphylos regismontana {{small|Eastw.}}
- Arctostaphylos × repens {{small|(Howell) P.V.Wells}}
- Arctostaphylos rudis {{small|Jeps. & Wiesl.}}
- Arctostaphylos sensitiva {{small|Jeps.}}
- Arctostaphylos silvicola {{small|Jeps. & Wiesl.}}
- Arctostaphylos stanfordiana {{small|Parry}}
- Arctostaphylos × strigosa {{small|Howell}}
- Arctostaphylos tomentosa {{small|(Pursh) Lindl.}}
- Arctostaphylos uva-ursi {{small|(L.) Spreng.}}
- Arctostaphylos virgata {{small|Eastw.}}
- Arctostaphylos viridissima {{small|(Eastw.) McMinn}}
- Arctostaphylos viscida {{small|Parry}}
- Arctostaphylos wellsii {{small|W.Knight}}
{{div col end}}
= Ranges =
- Subgenus Micrococcus
- Sect. Micrococcus
class="wikitable"
! Species ! Common name ! Range |
mendocinoensis
| Pygmy manzanita | |
myrtifolia
| Ione manzanita | California (Amador, Calaveras Counties) |
nissenana
| Nissenan manzanita | California (coastal and inland ranges north of San Francisco Bay) |
nummularia
| Glossyleaf manzanita | California (Mendocino County) |
- Subgenus Arctostaphylos, which has three sections:
- Sect. Arctostaphylos
class="wikitable"
! Species ! Common name ! Range |
alpina
| Alpine bearberry | |
bakeri
| Baker's manzanita | California (Sonoma County) |
densiflora
| Sonoma manzanita | California (Sonoma County) |
edmundsii
| Little Sur manzanita | California (Monterey County) |
franciscana
| Franciscan manzanita | California (San Francisco County) |
gabrielensis
| San Gabriel manzanita | California (Los Angeles County) |
glauca
| Bigberry manzanita | California and Baja California |
hispidula
| Gasquet manzanita | Coastal mountain ranges of southern Oregon and northern California |
hookeri
| Hooker's manzanita | California |
insularis
| Island manzanita | California (Santa Cruz Island) |
klamathensis
| Klamath manzanita | California (Klamath Mountains) |
manzanita
| Common manzanita, whiteleaf manzanita | California (Coast Ranges and Sierra Nevada foothills) |
mewukka
| Indian manzanita | California (Sierra Nevada) |
nevadensis
| Pinemat manzanita | California |
parryana
| Parry manzanita | California (southern) |
patula
| Greenleaf manzanita | Western North America |
pumila
| Sandmat manzanita | California (Monterey County) |
pungens
| Pointleaf manzanita | Southwestern United States and to northern and central Mexico |
rudis
| Shagbark manzanita | California (southern central coast) |
stanfordiana
| Stanford's manzanita | California (Outer North Coast Ranges north of the San Francisco Bay Area) |
uva-ursi
| Bearberry | Europe, Asia, North America |
viscida
| Sticky manzanita, whiteleaf manzanita | California and Oregon |
- Sect. Foliobracteata
class="wikitable"
! Species ! Common name ! Range |
andersonii
| Santa Cruz manzanita | Santa Cruz mountains (CA) |
auriculata
| Mount Diablo manzanita | Mount Diablo (CA) |
canescens
| Hoary manzanita | Coastal ranges of SW OR and N CA |
catalinae
| Santa Catalina Island manzanita | Santa Catalina Island (CA) |
columbiana
| Hairy manzanita | West coast from N. CA to S. BC |
confertiflora
| Santa Rosa Island manzanita | Santa Rosa Island (CA) |
cruzensis
| La Cruz manzanita | Monterey and San Luis Obispo counties (CA) |
glandulosa
| Eastwood manzanita | Coastal slops from OR, CA, Baja California |
glutinosa
| Schreiber's manzanita | Santa Cruz County (CA) |
hooveri
| Hoover's manzanita | Santa Lucia Mountains (CA) |
imbricata
| San Bruno Mountain manzanita | San Bruno Mountain (CA) |
luciana
| Santa Lucia manzanita | southern Santa Lucia Mountains (CA) |
malloryi
| Mallory's manzanita | Inner North Coast Ranges west and northwest of the Sacramento Valley (CA) |
montaraensis
| Montara manzanita | San Bruno Mountain and Montara Mountain (CA) |
montereyensis
| Monterey manzanita | Monterey County (CA) |
morroensis
| Morro manzanita | Morro Bay (CA) |
nortensis
| Del Norte manzanita | Del Norte County (CA), Curry County (OR), Josephine County (OR) |
obispoensis
| Serpentine manzanita | Southern Santa Lucia Mountains (CA) |
osoensis
| Oso manzanita | Los Osos Valley (San Luis Obispo, CA) |
otayensis
| Otay manzanita | San Diego County (CA) |
pajaroensis
| Pajaro manzanita | Monterey County, Santa Cruz County, San Benito County (CA) |
pallida
| Pallid manzanita | |
pechoensis
| Pecho manzanita | |
pilosula
| La Panzo manzanita | |
purissima
| La Purissima manzanita | |
refugioensis
| Refugio manzanita | |
regismontana
| Kings Mountain manzanita | |
silvicola
| Bonny Doon manzanita | |
tomentosa
| Woolyleaf manzanita | |
virgata
| Bolinas manzanita | |
viridissima
| Whitehair manzanita | |
wellsii
| Wells' manzanita | |
- Sect. Pictobracteata
class="wikitable"
! Species ! Common name ! Range |
pringlei
| Pringle manzanita | |
- Unassigned
class="wikitable"
! Species ! Common name ! Range |
rainbowensis
| Rainbow manzanita | |
gabilanensis
| Gabilan manzanita | |
ohloneana
| Ohlone manzanita | |
See also the closely related genus Comarostaphylis, previously often included in Arctostaphylos.
References
{{Reflist}}
Further reading
- {{citation |author=Hickman, James C. |year=1993 |title=The Jepson Manual: higher plants of California |publisher=University of California Press |location=Berkeley and Los Angeles, California |isbn=0-520-082559 |url-access=registration |url=https://archive.org/details/jepsonmanualhigh00hick }}
- {{citation |author=Hogan, C. Michael |year=2012 |url=http://www.eoearth.org/article/Arctostaphylos |title=Arctostaphylos |editor1=McGinley, M. |editor2=C. J. Cleveland |encyclopedia=Encyclopedia of Earth |publisher=National Council for Science and the Environment}}
- {{Citation |last=Kauffmann |first=Michael |title=Field Guide to Manzanitas: California, North America, and Mexico |date=2021-07-29 |edition=2nd |publisher=Backcountry Press |isbn=978-1941624159 |last2=Parker |first2=Tom |last3=Vasey |first3=Michael |last4=Bisbee |first4=Jeff}}
- [http://ucjeps.berkeley.edu/cgi-bin/get_JM_treatment.pl?Arctostaphylos Treatment from the Jepson Manual]
- {{citation |author=Wells, Philip V. |year=2000 |title=Manzanitas of California |publisher=University of Kansas, Department of Ecology & Evolutionary Biology |location=Lawrence, Kansas |isbn=0-933994-22-2 |url=https://books.google.com/books?id=jC5FAQAAIAAJ }}
- {{cite journal |author=Wells, Philip V. |year=1992 |title=Subgenera and sections of Arctostaphylos |journal=The Four Seasons |volume=9 |pages=64–69}}
External links
- [https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=9173 Jepson Manual Treatment]
{{Arbutoideae}}
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Category:Plants used in Native American cuisine
Category:Plants used in traditional Native American medicine
Category:Natural history of the California chaparral and woodlands
Category:Garden plants of North America