snail#Anatomy

Snails that respire using a lung belong to the group Pulmonata. As traditionally defined, the Pulmonata were found to be polyphyletic in a molecular study per Jörger et al., dating from 2010.{{cite journal|last=Jörger|first=Katharina M|author2=Stöger, Isabella |author3=Kano, Yasunori |author4=Fukuda, Hiroshi |author5=Knebelsberger, Thomas |author6= Schrödl, Michael |title=On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia|journal=BMC Evolutionary Biology|year=2010|volume=10|issue=1|pages=323|doi=10.1186/1471-2148-10-323|pmid=20973994|pmc=3087543 |doi-access=free |bibcode=2010BMCEE..10..323J }} But snails with gills also form a polyphyletic group; in other words, snails with lungs and snails with gills form a number of taxonomic groups that are not necessarily more closely related to each other than they are related to some other groups.

Both snails that have lungs and snails that have gills have diversified so widely over geological time that a few species with gills can be found on land and numerous species with lungs can be found in freshwater. Even a few marine species have lungs.

Snails can be found in a very wide range of environments, including ditches, deserts, and the abyssal depths of the sea. Although land snails may be more familiar to laymen, marine snails constitute the majority of snail species, and have much greater diversity and a greater biomass. Numerous kinds of snail can also be found in fresh water.

Most snails have thousands of microscopic tooth-like structures located on a banded ribbon-like tongue called a radula. The radula works like a file, ripping food into small pieces. Many snails are herbivorous, eating plants or rasping algae from surfaces with their radulae, though a few land species and many marine species are omnivores or predatory carnivores. Snails cannot absorb colored pigments when eating paper or cardboard so their feces are also colored.{{cite web|url=https://www.dezeen.com/2013/11/26/tiles-made-from-snail-poo-by-lieske-schreuder/|title=Floor tiles made of coloured snail poo by Lieske Schreuder - design|date=26 November 2013|access-date=29 March 2018}}

Several species of the genus Achatina and related genera are known as giant African land snails; some grow to {{Convert|38|cm|in|abbr=on}} from snout to tail, and weigh {{convert|1|kg|0|abbr=on}}.{{cite book |last=Fredericks |first=Anthony D. |year=2010 |title=How Long Things Live & How They Live As Long As They Do |publisher=Stackpole Books |isbn=9780811736220 |page=[https://archive.org/details/howlongthingsliv0000fred/page/73 73] |url=https://archive.org/details/howlongthingsliv0000fred |url-access=registration |access-date=19 June 2012}} The largest living species of sea snail is Syrinx aruanus; its shell can measure up to {{convert|90|cm|abbr=on}} in length, and the whole animal with the shell can weigh up to {{convert|18|kg|abbr=on}}. The smallest land snail, Angustopila psammion, was discovered in 2022 and measures 0.6 mm in diameter.{{Cite web |date=2 Feb 2022 |title=New snail species are world's smallest, tinier than grains of sand |url=https://www.nationalgeographic.com/animals/article/smallest-snails-on-earth-discovered?linkId=150748492 |url-status=live |archive-url=https://web.archive.org/web/20220830205208/https://www.nationalgeographic.com/animals/article/smallest-snails-on-earth-discovered?linkId=150748492 |archive-date=30 Aug 2022 |access-date=16 Oct 2024 |website=National Geographic}}

The largest known land gastropod is the African giant snail Achatina achatina, the largest recorded specimen of which measured {{Convert|39.3|cm|in}} from snout to tail when fully extended, with a shell length of {{Convert|27.3|cm|in|abbr=on}} in December 1978. It weighed exactly 900 g (about 2 lb). Named Gee Geronimo, this snail was owned by Christopher Hudson (1955–79) of Hove, East Sussex, UK, and was collected in Sierra Leone in June 1976.{{Cite web|website=Guinness World Records|title=Largest Snail|url=http://www.guinnessworldrecords.com/explore-records/animals?page=2|access-date=17 December 2016}}

Snails are protostomes. That means during development, in the gastrulation phase, the blastopore forms the mouth first. Cleavage in snails is spiral holoblastic patterning. In spiral holoblastic cleavage, the cleavage plane rotates each division and the cell divisions are complete. Snails do not undergo metamorphosis after hatching. Snails hatch in the form of small adults. The only additional development they will undergo is to consume calcium to strengthen their shell. Snails can be male, female, hermaphroditic, or parthenogenetic so there are many different systems of sexual determination.

Snails have complex organ systems and anatomies that differ greatly from most animals. Snails and most other Mollusca share three anatomical features; the foot, the mantle, and the radula.

Foot: The foot is a muscular organ used by Gastropods for locomotion. Gastropods' stomachs are located within their foot. Both land and sea snails travel by contracting foot muscles to deform the mucus layer beneath it into different wave-like patterns.{{Cite web |last=Pegler |first=Sam |date=2018 |title=Snail locomotion |url=https://gfd.whoi.edu/wp-content/uploads/sites/18/2018/03/SamPegler_136807.pdf |access-date=October 7, 2024 |website=Woods Hole Oceanographic Institution}}

Mantle: The mantle is the organ that produces shells for most species of mollusca. In snails, the mantle secretes the shell along the snail shell opening, continuously growing and producing the shell for the entirety of the snail’s life.{{Cite web |date=2024-09-03 |title=Gastropod - Shell Structure, Anatomy, Diversity {{!}} Britannica |url=https://www.britannica.com/animal/gastropod/The-shell |access-date=2024-10-07 |website=www.britannica.com |language=en}} The mantle creates a compartment known as the mantle cavity and is used by many mollusca as the surface where gas exchange occurs. Snails that use the mantle cavity as a lung are known as Pulmonate snails. Other snails may only have a gill. Snails in the Caenogastropoda families like Ampullariidae, have both a gill and a lung.{{Cite journal |last1=Rodriguez |first1=Cristian |last2=Prieto |first2=Guido I. |last3=Vega |first3=Israel A. |last4=Castro-Vazquez |first4=Alfredo |date=2021-04-14 |title=Morphological grounds for the obligate aerial respiration of an aquatic snail: functional and evolutionary perspectives |journal=PeerJ |volume=9 |pages=e10763 |doi=10.7717/peerj.10763 |doi-access=free |issn=2167-8359 |pmc=8052964 |pmid=33954023}}

Shell: Snail shells are mainly composed of a mixture of proteins called conchin, and calcium carbonate. Conchin is the main component in the outer layer of the shell, known as the periostracum. The inner layers of the shell are composed of a network of calcium carbonate, conchin, and different mineral salts. The mantle produces the shell through addition around a central axis called the columella, causing a spiraling pattern.{{Cite web |title=compound coil |url=https://dnr.illinois.gov/education/wildaboutpages/wildaboutinvertebrates/wildaboutillinoissnails/walscompoundcoil.html |access-date=2024-10-07 |website=dnr.illinois.gov |language=en}} The spiraling patterns on a snail’s shell are known as coils or whorls. Whorl size generally increases as the snail ages. Size differences in shell size are believed to be mainly influenced by genetic and environmental components. Moister conditions often correlate with larger snails. In larger populations, adult snails attain smaller shell sizes due to the effects of pheromones on growth rate.{{Cite journal |last=Goodfriend |first=Glenn |date=1986 |title=Variation in Land-snail Shell form and Size and its Causes: a Review |url=https://academic.oup.com/sysbio/article-abstract/35/2/204/1649641 |journal=Systematic Biology |volume=35 |issue=2 |pages=204–223 |doi=10.1093/sysbio/35.2.204 |via=Oxford Academic}}

Radula: The radula is an anatomical structure used by most species of Mollusca for feeding. Gastropods are morphologically highly variable and have diverse feeding strategies. Snails can be herbivores, detritivores, scavengers, parasites, ciliary feeders, or have highly specialized predation.{{Cite journal |last1=Krings |first1=Wencke |last2=Faust |first2=Taissa |last3=Kovalev |first3=Alexander |last4=Neiber |first4=Marco Thomas |last5=Glaubrecht |first5=Matthias |last6=Gorb |first6=Stanislav |date=July 2019 |title=In slow motion: radula motion pattern and forces exerted to the substrate in the land snail Cornu aspersum (Mollusca, Gastropoda) during feeding |journal=Royal Society Open Science |language=en |volume=6 |issue=7 |pages=190222 |doi=10.1098/rsos.190222 |issn=2054-5703 |pmc=6689628 |pmid=31417728|bibcode=2019RSOS....690222K }} Nearly all snails utilize a feeding apparatus including the oral structures of one or more jaws and the radula. The radula comprises a chitinous ribbon with teeth arranged in transverse and longitudinal rows.{{Cite journal |last=Vortsepneva |first=Elena |date=2023 |title=Main patterns of radula formation and ontogeny in Gastropoda |url=https://onlinelibrary.wiley.com/doi/abs/10.1002/jmor.21538 |journal=Journal of Morphology |volume=284 |issue=1 |pages=e21538 |doi=10.1002/jmor.21538 |pmid=36426387 |via=Wiley Online Library}} The radula continually renews itself during the entire lifespan of a mollusk. The teeth and membrane are continuously synthesized in the radular sac and then shifted forward towards the working zone of the radula. The teeth harden and mineralize during their travel to the working zone. The presence of the radula is common throughout most snail species, but often differs in many characteristics, like the shape, size, and number of odontoblasts that form a tooth.

The average snail's diet varies greatly depending on the species, including different feeding styles from herbivores to highly specialized feeders and parasites.{{Cite journal |last1=Krings |first1=Wencke |last2=Faust |first2=Taissa |last3=Kovalev |first3=Alexander |last4=Neiber |first4=Marco Thomas |last5=Glaubrecht |first5=Matthias |last6=Gorb |first6=Stanislav |date=July 2019 |title=In slow motion: radula motion pattern and forces exerted to the substrate in the land snail Cornu aspersum (Mollusca, Gastropoda) during feeding |journal=Royal Society Open Science |language=en |volume=6 |issue=7 |pages=190222 |doi=10.1098/rsos.190222 |issn=2054-5703 |pmc=6689628 |pmid=31417728|bibcode=2019RSOS....690222K }} Some snails like the Euglandina rosea, or rosy wolfsnail, are carnivorous and prey on other snails.{{Cite journal |last1=Clifford |first1=Kavan T. |last2=Gross |first2=Liaini |last3=Johnson |first3=Kwame |last4=Martin |first4=Khalil J. |last5=Shaheen |first5=Nagma |last6=Harrington |first6=Melissa A. |date=2003 |title=Slime-trail tracking in the predatory snail, Euglandina rosea. |url=https://doi.apa.org/doi/10.1037/0735-7044.117.5.1086 |journal=Behavioral Neuroscience |language=en |volume=117 |issue=5 |pages=1086–1095 |doi=10.1037/0735-7044.117.5.1086 |pmid=14570557 |issn=1939-0084}} However, most land snails are herbivores or omnivores.{{Cite web |title=Mollusks : Carnegie Museum of Natural History |url=https://www.carnegiemnh.org/science/mollusks/dietbehavior.html |access-date=2024-12-04 |website=www.carnegiemnh.org}} Among land snails, there is also a large variation in preference for specific food. For example, Cepaea nemoralis, or the grove snail, prefers dead plant material over fresh herbs or grasses. Age may also impact food preference, with adult grove snails showing a significantly larger preference for dead plant material than juvenile grove snails.{{Cite journal |last1=Williamson |first1=P. |last2=Cameron |first2=R. A. D. |date=1976 |title=Natural Diet of the Landsnail Cepaea Nemoralis |url=https://www.jstor.org/stable/3543468 |journal=Oikos |volume=27 |issue=3 |pages=493–500 |doi=10.2307/3543468 |jstor=3543468 |bibcode=1976Oikos..27..493W |issn=0030-1299}} Other snails, like the generalist herbivore Arianta arbustorum, or copse snail, choose their meals based on availability, consuming a mix of arthropods, wilted flowers, fresh and decayed plant material, and soil.{{Cite journal |last1=Speiser |first1=Bernhard |last2=Rowell-Rahier |first2=Martine |date=1991 |title=Effects of Food Availability, Nutritional Value, and Alkaloids on Food Choice in the Generalist Herbivore Arianta arbustorum (Gastropoda: Helicidae) |url=https://www.jstor.org/stable/3545495 |journal=Oikos |volume=62 |issue=3 |pages=306–318 |doi=10.2307/3545495 |jstor=3545495 |bibcode=1991Oikos..62..306S |issn=0030-1299}}

Generally, land snails are most active at night due to the damp weather. The humid nighttime air minimizes water evaporation and is beneficial to land snails because their movement requires mucus, which is mostly composed of water. In addition to aiding movement, mucus plays a vital role in transporting food from the gill to the mouth, cleansing the mantle cavity, and trapping food before ingestion.{{Citation |last1=Davies |first1=Mark S. |title=Mucus from Marine Molluscs |date=1998-01-01 |volume=34 |pages=1–71 |editor-last=Blaxter |editor-first=J. H. S. |url=https://linkinghub.elsevier.com/retrieve/pii/S0065288108602102 |access-date=2024-12-04 |publisher=Academic Press |doi=10.1016/s0065-2881(08)60210-2 |last2=Hawkins |first2=S. J. |series=Advances in Marine Biology |isbn=978-0-12-026134-5 |editor2-last=Southward |editor2-first=A. J. |editor3-last=Tyler |editor3-first=P. A.}}

{{Main article|Land snail|Freshwater snail|Sea snail}}

{{Main article|Slug}}

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File:Slug 02.jpg

Gastropods that lack a conspicuous shell are commonly called slugs rather than snails.{{cite web|url=https://www.diffen.com/difference/Slug_vs_Snail|title=Slug vs Snail - Difference and Comparison - Diffen|access-date=29 March 2018}} Some species of slug have a maroon-brown shell, some have only an internal vestige that serves mainly as a calcium lactate repository, and others have some to no shell at all. Other than that there is little morphological difference between slugs and snails. There are however important differences in habitats and behavior.

A shell-less animal is much more maneuverable and compressible, so even quite large land slugs can take advantage of habitats or retreats with very little space, retreats that would be inaccessible to a similar-sized snail. Slugs squeeze themselves into confined spaces such as under loose bark on trees or under stone slabs, logs or wooden boards lying on the ground. In such retreats they are in less danger from either predators or desiccation. Those are often suitable places for laying their eggs.

Slugs as a group are far from monophyletic; scientifically speaking "slug" is a term of convenience with little taxonomic significance. The reduction or loss of the shell has evolved many times independently within several very different lineages of gastropods. The various taxa of land and sea gastropods with slug morphology occur within numerous higher taxonomic groups of shelled species; such independent slug taxa are not in general closely related to one another.{{Cite web |title=TUATARA: VOLUME 25, ISSUE 2, JANUARY 1982 HOW TO BE SLUGGISH |url=https://ndhadeliver.natlib.govt.nz/webarchive/20210104000423/http://nzetc.victoria.ac.nz/tm/scholarly/tei-Bio25Tuat02-t1-body-d2.html |access-date=2024-12-27 |website=ndhadeliver.natlib.govt.nz}}

Snails can also be associated with parasitic diseases such as schistosomiasis, angiostrongyliasis, fasciolopsiasis, opisthorchiasis, fascioliasis, paragonimiasis and clonorchiasis, which can be transmitted to humans.

{{More citations needed section|date=March 2018}}

File:Snail-wiki-120-Zachi-Evenor.jpg – garden snail]]

Land snails are known as an agricultural and garden pest but some species are an edible delicacy and occasionally household pets. In addition, their mucus can also be used for skin care products.{{Cite web |last=Howley |first=Elaine |date=18 July 2022 |title=Snail Mucin for Skin Care: Products and Uses |url=https://health.usnews.com/wellness/articles/snail-mucin-for-skin-care |url-status=live |archive-url=https://web.archive.org/web/20230718220458/https://health.usnews.com/wellness/articles/snail-mucin-for-skin-care |archive-date=18 July 2023 |access-date=29 July 2023 |website=U.S. News}}

There is a variety of snail-control measures that gardeners and farmers use in an attempt to reduce damage to valuable plants. Traditional pesticides are still used, as are many less toxic control options such as concentrated garlic or wormwood solutions. Copper metal is also a snail repellent, and thus a copper band around the trunk of a tree will prevent snails from climbing up and reaching the foliage and fruit. A layer of a dry, finely ground, and scratchy substance such as diatomaceous earth can also deter snails.Sanders, April. [http://homeguides.sfgate.com/egg-shells-stop-snails-eating-plants-87576.html "Do Egg Shells Help Stop Snails From Eating Plants?"] Home Guides | SF Gate. Accessed 01 July 2019.

The decollate snail (Rumina decollata) will capture and eat garden snails, and because of this it has sometimes been introduced as a biological pest control agent. However, this is not without problems, as the decollate snail is just as likely to attack and devour other gastropods that may represent a valuable part of the native fauna of the region.

Certain varieties of snails, notably the family Muricidae, produce a secretion that is a color-fast natural dye. The ancient Tyrian purple was made in this way as were other purple and blue dyes.{{cite journal |last=Ziderman |first=I. I. |year=1986 |title=Purple dye made from shellfish in antiquity |journal=Review of Progress in Coloration |issn=1472-3581 |volume=16 |pages=46–52|doi=10.1111/j.1478-4408.1986.tb03743.x }}{{cite journal |title=Whelks and purple dye in Anglo-Saxon England |last=Biggam |first=Carole P. |journal=The Archeo+Malacology Group Newsletter |issue=9 |date=March 2006 |url=http://triton.anu.edu.au/MalacGp09.pdf }}{{cite book |last=Moorey |first=Peter |year=1999 |title=Ancient Mesopotamian Materials and Industries: The Archaeological Evidence |url=https://archive.org/details/ancientmesopotam00moor |url-access=limited |publisher=Eisenbrauns |location=Winona Lake, Indiana |isbn=978-1-57506-042-2 |page=[https://archive.org/details/ancientmesopotam00moor/page/n81 138]}} The extreme expense of extracting this secretion is sufficient quantities limited its use to the very wealthy. It is such dyes as these that led to certain shades of purple and blue being associated with royalty and wealth.{{cite book |last=Nuttall |first=Zelia |author-link=Zelia Nuttall |editor1-last=Boas |editor1-first=Franz |year=1909 |chapter=A Curious Survival in Mexico of the Use of the Purpura Shell-fish for Dyeing |title=Putnam Anniversary Volume |volume=Anthropological Essays Presented to Fredrick Ward Putnam in Honor of his Seventieth Birthday, by his Friends and Associates |publisher=G. E. Stechert & Co. |location=New York |lccn=10011191 |pages=368–384 |chapter-url=https://archive.org/details/cu31924029880774}}

Throughout history, snails have been kept as pets. There are many famous snails such as Lefty (Born Jeremy) and within fiction, Gary and Brian the snail.{{Cite web|url=https://www.bbc.co.uk/programmes/articles/J1y1vyNcTlCsfp7CmQTz03/seven-of-the-worlds-most-famous-snails|title=BBC Radio 3 – Slow Radio – Seven of the world's most famous snails|website=BBC|language=en-GB|access-date=2020-02-14}}

{{Main article|Snails as food}}

File:Cooked snails.JPG]]

In French cuisine, edible snails are served for instance in Escargot à la Bourguignonne. The practice of rearing snails for food is known as heliciculture. For purposes of cultivation, the snails are kept in a dark place in a wired cage with dry straw or dry wood. Coppiced wine-grape vines are often used for this purpose. During the rainy period, the snails come out of hibernation and release most of their mucus onto the dry wood/straw. The snails are then prepared for cooking. Their texture when cooked is slightly chewy and tender.

As well as being eaten as gourmet food, several species of land snails provide an easily harvested source of protein to many people in poor communities around the world. Many land snails are valuable because they can feed on a wide range of agricultural wastes, such as shed leaves in banana plantations. In some countries, giant African land snails are produced commercially for food.

Land snails, freshwater snails and sea snails are all eaten in many countries. In certain parts of the world snails are fried. For example, in Indonesia, they are fried as satay, a dish known as sate kakul. The eggs of certain snail species are eaten in a fashion similar to the way caviar is eaten.{{cite news | url=https://www.reuters.com/article/us-france-snails-idUSL2055417620071220 | title=The height of slow-food, France serves snail caviar | newspaper=Reuters | date=20 December 2007 }}

In Bulgaria, snails are traditionally cooked in an oven with rice or fried in a pan with vegetable oil and red paprika powder. Before they are used for those dishes, however, they are thoroughly boiled in hot water (for up to 90 minutes) and manually extracted from their shells. The two species most commonly used for food in the country are Helix lucorum and Helix pomatia.{{citation needed|date=January 2020}}

Snails and slug species that are not normally eaten in certain areas have occasionally been used as famine food in historical times. A history of Scotland written in the 1800s recounts a description of various snails and their use as food items in times of plague.{{cite book |last=Chambers |first=Robert |author-link=Robert Chambers (publisher born 1802) |year=1858|title=Domestic annals of Scotland, from the reformation to the revolution |publisher=W. & R. Chambers |url=https://archive.org/details/domesticannalsof02chamiala}} (Also quoted [http://www.electricscotland.com/history/domestic/vol2ch2c.htm here].

File:Mochesnail.jpg sp.) on a Moche pot, 200 AD, Larco Museum Collection, Lima, Peru]]

Because of its slowness, the snail has traditionally been seen as a symbol of laziness. In Christian culture, it has been used as a symbol of the deadly sin of sloth.{{cite book |last=de Vries |first=Ad |year=1976 |title=Dictionary of Symbols and Imagery |publisher=North-Holland Publishing Company |location=Amsterdam |isbn=978-0-7204-8021-4 |page=[https://archive.org/details/dictionaryofsymb0000vrie/page/430 430] |url=https://archive.org/details/dictionaryofsymb0000vrie/page/430 }}{{cite book |last=Tresidder |first=Jack |year=2006 |title=Symbols and Their Meanings |publisher=Barnes & Noble |location=New York |isbn=978-0-7607-8164-7 |page=41 |url=https://books.google.com/books?id=tlVsHBc443gC&q=The+snail+was+also+a+Christian+emblem+of+sloth.+...}} In Mayan mythology, the snail is associated with sexual desire, being personified by the god Uayeb.Susan Milbrath, Star Gods of the Maya: Astronomy in Art, Folklore, and Calendars, University of Texas Press, 01/01/2010

Snails were widely noted and used in divination. The Greek poet Hesiod wrote that snails signified the time to harvest by climbing the stalks, while the Aztec moon god Tecciztecatl bore a snail shell on his back. This symbolised rebirth; the snail's penchant for appearing and disappearing was analogised with the moon.{{cite book |last=Cooper |first=J. C. |year=1992 |title=Symbolic and Mythological Animals |publisher=Aquarian Press |location=London |isbn=978-1-85538-118-6 |page=213}}

File:Dewi Sekartaji Keong Emas.JPG

Keong Emas (Javanese and Indonesian for Golden Snail) is a popular Javanese folklore about a princess magically transformed and contained in a golden snail shell. The folklore is a part of popular Javanese Panji cycle telling the stories about the prince Panji Asmoro Bangun (also known as Raden Inu Kertapati) and his consort, princess Dewi Sekartaji (also known as Dewi Chandra Kirana).

In contemporary speech, the expression "a snail's pace" is often used to describe a slow, inefficient process. The phrase "snail mail" is used to mean regular postal service delivery of paper messages as opposed to the delivery of email, which can be virtually instantaneous.

• Pasilalinic-sympathetic compass

{{Reflist}}

File:The snail 1 నత్త.jpg|A snail

File:Snail climbing stone slabs.png|Snail climbing stone slabs at Servia, Greece. The shell is so heavy that its bottom side becomes visible.

File:SeaSnails.ogv|Snails feeding on the sea floor (50 sec)

File:Snail Melc melc codobelc.ogv|Video of snail after rain (31 sec)

File:Snail moving on ground.webm|Snail moving on a wet ground

File:Snail moving across leaves.webm|Snail moving across leaves.

File:Lumache nei monti Iblei - Pantalica.jpg|Snails in the Hyblaean Mountains.

{{Commons category|Snails}}

• [http://www.snail-world.com/ Introduction to Snails], Infoqis Publishing, Co.

{{Edible molluscs}}

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Category:Mollusc common names

Category:Paraphyletic groups