Alethinophidia

{{short description|Clade of snakes}}

{{Automatic taxobox

| taxon = Alethinophidia

| image = Natrix natrix (Marek Szczepanek).jpg

| image_caption = Grass snake, Natrix natrix

| fossil_range = {{Fossil range|94|0}} Cenomanian to PresentJ.-C. Rage and C. Werner. 1999.[https://www.academia.edu/33304223/MID-CRETACEOUS_CENOMANIAN_SNAKES_FROM_WADI_ABU_HASHIM_SUD_AN_THE_EARLIEST_SNAKE_ASSEMBLAGE "Mid-Cretaceous (Cenomanian) snakes from Wadi Abu Hashim, Sudan: The earliest snake assemblage"]. 35, 85-110

| authority = Nopcsa, 1923

| subdivision_ranks = Subclades

| subdivision = * Afrophidia

}}

:Common names: advanced snakes.

The Alethinophidia are an infraorder of snakes that includes all snakes other than blind snakes and thread snakes. Snakes have long been grouped into families within Alethinophidia based on their morphology, especially that of their teeth. More modern phylogenetic hypotheses using genetic data{{ITIS |id=634390 |taxon=Alethinophidia |access-date=14 May 2018}}{{cite journal|last1=Pyron|first1=R. A.|last2=Burbrink|first2=F.|last3=Wiens|first3=J. J.|title=A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes|journal=BMC Evolutionary Biology|date=2013|volume=13|issue=1 |pages=93|doi=10.1186/1471-2148-13-93|pmid=23627680|pmc=3682911 |bibcode=2013BMCEE..13...93P |doi-access=free }}{{cite journal|last1=Reynolds|first1=RG|last2=Niemiller|first2=ML|last3=Revell|first3=LJ|title=Toward a Tree-of-Life for the boas and pythons: multilocus species-level phylogeny with unprecedented taxon sampling|journal=Molecular Phylogenetics and Evolution|date=2014|volume=71|pages=201–213|url=http://www.rgrahamreynolds.info/wp-content/uploads/2014/01/Reynolds_etal_2014_MPE.pdf|doi=10.1016/j.ympev.2013.11.011|access-date=2018-05-14|archive-url=https://web.archive.org/web/20151202212953/http://www.rgrahamreynolds.info/wp-content/uploads/2014/01/Reynolds_etal_2014_MPE.pdf|archive-date=2015-12-02|url-status=dead|pmid=24315866|bibcode=2014MolPE..71..201G }}{{cite journal|last1=Streicher|first1=J. W.|last2=Ruane|first2=S.|title=Phylogenomics of Snakes|journal=eLS|date=2018|doi=10.1002/9780470015902.a0027476|pages=1–8|isbn=9780470015902}}{{cite journal|last1=Figueroa|first1=A.|last2=McKelvy|first2=A. D.|last3=Grismer|first3=L. L.|last4=Bell|first4=C. D.|last5=Lailvaux|first5=S. P.|title=A species-level phylogeny of extant snakes with description of a new colubrid subfamily and genus|journal=PLOS ONE|date=2016|volume=11|issue=9|pages=e0161070|pmc=5014348|doi=10.1371/journal.pone.0161070|pmid=27603205|bibcode=2016PLoSO..1161070F|doi-access=free}}{{cite journal|last1=Zheng|first1=Y|last2=Wiens|first2=JJ|title=Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species|journal=Molecular Phylogenetics and Evolution|date=2016|volume=94|issue=Pt B|pages=537–547|url=http://www.wienslab.com/Publications_files/Zheng_Wiens_2015b_MPE.pdf|doi=10.1016/j.ympev.2015.10.009|pmid=26475614|bibcode=2016MolPE..94..537Z}}{{cite book|last1=Scanlon|first1=J. D.|last2=Lee|first2=M. S. Y.|editor1-last=Aldridge|editor1-first=R. D.|editor2-last=Sever|editor2-first=D. M.|title=The Major Clades of Living Snakes: Morphological Evolution, Molecular Phylogeny, and Divergence Dates in Reproductive Biology and Phylogeny of Snakes|date=2011|publisher=Science Publishers|location=Enfield, NH|pages=55–95}}{{cite book|last1=Vidal|first1=N.|last2=Delmas|first2=A. S.|last3=Hedges|first3=S. B.|editor1-last=Henderson|editor1-first=R. W.|editor2-last=Powell|editor2-first=R.|title=The higher-level relationships of alethinophidian snakes inferred from seven nuclear and mitochondrial genes|date=2007|publisher=Eagle Mountain Publishing|location=Eagle Mountain, Utah, USA|pages=27–33}}{{cite book|last1=Vitt|first1=L. J.|last2=Caldwell|first2=J. P.|title=Herpetology: an introductory biology of amphibians and reptiles|date=2014|publisher=Academic Press|location=Burlington|pages=108–109|edition=4th}}{{cite web|last1=Uetz|first1=Peter|title=Serpentes at The Reptile Database|url=http://reptile-database.reptarium.cz/advanced_search?taxon=Serpentes&submit=Search|website=The Reptile Database|publisher=EMBL|access-date=14 May 2018}}{{overcite|date=March 2024}} support the recognition of 19 extant families (see below), although the taxonomy of alethinophidian snakes has long been debated, and ultimately the decision whether to assign a particular clade to a particular Linnaean rank (such as a superfamily, family, or subfamily) is arbitrary.

Etymology

The infraorder name Alethinophidia derives from the two Ancient Greek words {{wikt-lang|grc|ἀληθινός}} ({{grc-transl|ἀληθινός}}), meaning "truthful, genuine", and {{wikt-lang|grc|ὄφις}} ({{grc-transl|ὄφις}}), meaning "snake".{{Cite book |title=Abrégé du dictionnaire grec français |last=Bailly |first=Anatole |date=1981-01-01 |publisher=Hachette |isbn=978-2010035289 |location=Paris |oclc=461974285 }}{{Cite web |url=http://www.tabularium.be/bailly/ |title=Greek-french dictionary online |last=Bailly |first=Anatole |website=www.tabularium.be |access-date=January 7, 2019}}

Fossil record

Fossils of alethinophidians were found in Cenomanian (Middle Cretaceous) sites of Wadi Milk Formation in Wadi Abu Hashim, Sudan. Coniophis presents the vertebral morphology similar to modern-day Aniliidae. Two extinct families from the same location, the Anomalophiidae and Russellophiidae, also belong to the Alethinophidia. Krebsophis is the earliest russellophiid. The family Nigerophiidae includes both aquatic Nubianophis from Wadi Abu Hashim and Nigerophis from the Palaeocene of Niger. The genus Eoanilius (belongs to Aniliidae) appeared in the Eocene. It is also existed in Oligocene and early Miocene. The extinct marine Simoliophidae are known from the Cenomanian of North Africa, the Middle East, and Eastern Europe, indicating a Tethyan distribution; they are notable for preserving evidence of vestigial hindlimbs.{{Cite journal |last1=Hsiang |first1=Allison Y. |last2=Field |first2=Daniel J. |last3=Webster |first3=Timothy H. |last4=Behlke |first4=Adam DB |last5=Davis |first5=Matthew B. |last6=Racicot |first6=Rachel A. |last7=Gauthier |first7=Jacques A. |date=2015-05-20 |title=The origin of snakes: revealing the ecology, behavior, and evolutionary history of early snakes using genomics, phenomics, and the fossil record |journal=BMC Evolutionary Biology |volume=15 |issue=1 |pages=87 |doi=10.1186/s12862-015-0358-5 |issn=1471-2148 |pmc=4438441 |pmid=25989795 |bibcode=2015BMCEE..15...87H |doi-access=free }}

Systematics

= Extant taxa =

= Fossil taxa =

Based on Gower & Zaher (2022):{{Cite book |last1=Gower |first1=David J. |url=http://dx.doi.org/10.1017/9781108938891.027 |title=The Origin and Early Evolutionary History of Snakes |last2=Zaher |first2=Hussam |date=2022-08-11 |publisher=Cambridge University Press |pages=473–476 |doi=10.1017/9781108938891.027|isbn=978-1-108-93889-1 }}

See also

References

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