Cambridge Greensand

{{confuse|Cambridge Formation}}

{{Infobox rockunit

| name = Cambridge Greensand

| image =

| caption =

| type = Member

| age = Earliest Cenomanian ~{{fossilrange|100}}

| period = Cenomanian

| prilithology = Glauconitic marl

| otherlithology = Phosphorite

| namedby =

| region = England

| country = {{flag|UK}}

| coordinates = {{coord|52.2|N|0.1|E|display=inline,title}}

| paleocoordinates = {{coord|41.0|N|1.8|E|display=inline}}

| unitof = West Melbury Marly Chalk Formation

| subunits =

| underlies = Chalk Group

| overlies = Gault Formation

| thickness = {{convert|0.1|-|1|m|ft|abbr=on}}

| extent = North Hertfordshire and Cambridgeshire

| area =

| map = {{Location map+ | England

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | England

| lat_deg = 52.2

| lon_deg = 0.1

| mark = Lightgreen pog.svg

| marksize = 12

}}

}}

| map_caption =

| namedfor = Cambridge

| location_ts = Arlesey Brickpit

}}

The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age.{{Cite journal|last1=Hart|first1=Malcolm B.|last2=Fox|first2=Lyndsey R.|date=2020|title=Micropalaeontology and stratigraphical setting of the Cambridge Greensand|url=http://sp.lyellcollection.org/lookup/doi/10.1144/SP498-2018-144|journal=Geological Society, London, Special Publications|language=en|volume=498|issue=1|pages=147–163|doi=10.1144/SP498-2018-144|bibcode=2020GSLSP.498..147H|s2cid=210304478|issn=0305-8719}} It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation.[http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=CBG Cambridge Greensand] at BGS It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.

Description

The lithology is made out of glauconitic marl, described as a "chalk mud", containing abundant ostracod, coccolith and foram remains, with a concentration of phosphatic nodules and bones at the base.

Vertebrate paleofauna

= Birds =

class="wikitable"

|+

! colspan="6" |Birds

Genus

!Species

!Locality

!Material

!Notes

!Images

rowspan="3" |Enaliornis

|E. barrette

|

|Braincases, vertebrae, pelvis [and] limb elements"Table 11.1," in Weishampel, et al. (2004). Page 215.

| rowspan="3" |This genus is the oldest known hesperornithine

|

E sedgwicki

|

|Hindlimb elements

|

E. seeleyi

|

|Assorted cranial and postcranial elements

|

= Dinosaurs =

== Ornithischians ==

{{Paleobiota-key-compact}}

class="wikitable"

! colspan="6" |Ornithischians

Genus

!Species

!Locality

!Material

!Notes

!Images

rowspan="2" |Anoplosaurus

|A. curtonotus

|

|"Partial postcranium""Table 17.1," in Weishampel, et al. (2004). Page 367.

|

|

A. major

|

|"Cervical vertebrae." "Vertebrae.""Table 19.1," in Weishampel, et al. (2004). Page 417.

|

|

rowspan="4" |Acanthopholis

|style="background:#E6E6E6;"| A. eucercus

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| "[Two] caudal centra."

|style="background:#E6E6E6;"| Nomen dubium

|style="background:#E6E6E6;"|

style="background:#E6E6E6;"| A. platypus

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| "Phalanx, caudal centra."

|style="background:#E6E6E6;"| Nomen dubium

|style="background:#E6E6E6;"|

style="background:#fbdddb;" |A. macrocercus

|style="background:#fbdddb;" |

|style="background:#fbdddb;" |"Osteoderms." "Vertebrae, fragmentary skeleton elements."

|style="background:#fbdddb;" |Reassigned to Syngonosaurus

|style="background:#fbdddb;" |

style="background:#E6E6E6;"| A. stereocercus

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| "Osteoderms." "Vertebrae."

|style="background:#E6E6E6;"| Nomen dubium

|style="background:#E6E6E6;"|

Eucercosaurus

|E. tanyspondylus

|

|"Vertebrae."

|

|

style="background:#E6E6E6;"| Trachodon

|style="background:#E6E6E6;"| T. cantabrigiensis

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| "Dentary tooth.""Table 20.1," in Weishampel, et al. (2004). Page 443.

|style="background:#E6E6E6;"| Nomen dubium

|style="background:#E6E6E6;"|

== Saurischians ==

class="wikitable"

! colspan="6" |Saurischians

Genus

!Species

!Locality

!Material

!Notes

!Images

Macrurosaurus

|M. semnus

|

|"Caudal vertebrae""Table 13.1," in Weishampel, et al. (2004). Page 270.

|Titanosauriform sauropod or indeterminate macronarian

|

= Pterosaurs =

{{Paleobiota-key-compact}}

class="wikitable"

! colspan="6" |Pterosaurs

Genus

!Species

!Locality

!Material

!Notes

!Images

Aerodraco{{cite journal |last1=Holgado |first1=B. |last2=Pêgas |first2=R.V. |year=2020 |title=A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae |journal=Acta Palaeontologica Polonica |volume=65 |doi=10.4202/app.00751.2020|doi-access=free }}

| A. sedgwickii

|

| Anterior rostrum

|

|

rowspan="2" |Amblydectes

|A. crassidens

|

|Jaw fragments

|

|

style="background:#E6E6E6;"| A. eurygnathus

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| Jaw fragments

|style="background:#E6E6E6;"| Possibly a junior synonym of A. crassidens{{Cite journal |last=Holgado |first=Borja |date=2021 |title=On the validity of the genus Amblydectes Hooley 1914 (Pterodactyloidea, Anhangueridae) and the presence of Tropeognathinae in the Cambridge Greensand |url=http://www.scielo.br/scielo.php?script=sci_arttext&pid=S0001-37652021000401220&tlng=en |journal=Anais da Academia Brasileira de Ciências |volume=93 |issue=suppl 2 |doi=10.1590/0001-3765202120201658 |issn=1678-2690}}

|style="background:#E6E6E6;"|

rowspan=2 |Camposipterus

|C. colorhinus

|

|Anterior rostra fragments

|

|

C. nasutus

|

|Partial rostrum

|

|

Draigwenia

|D. platystomus

|

|Anterior rostrum fragments

|Formerly Amblydectes platystomus

|

rowspan="2" |Lonchodraco

|L. machaerorhynchus

|

|

|May or may not{{Cite journal |last1=Pêgas |first1=Rodrigo V. |date=2024-06-10 |title=A taxonomic note on the tapejarid pterosaurs from the Pterosaur Graveyard site (Caiuá Group, ?Early Cretaceous of Southern Brazil): evidence for the presence of two species |journal=Historical Biology |language=en |pages=1–22 |url=https://www.researchgate.net/publication/381306918 |doi=10.1080/08912963.2024.2355664 |issn=0891-2963}} be referrable to Ikrandraco

|

style="background:#E6E6E6;"| L. microdon

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| Junior synonym of L. machaerorhynchus{{cite journal|first=A.O. |last=Averianov |year=2020 |title=Taxonomy of the Lonchodectidae (Pterosauria, Pterodactyloidea) |journal=Proceedings of the Zoological Institute RAS |volume=324 |issue=1 |pages=41–55 |doi=10.31610/trudyzin/2020.324.1.41|doi-access=free }}

|style="background:#E6E6E6;"|

Nicorhynchus

| N. capito{{cite journal |last1=Holgado |first1=B. |last2=Pêgas |first2=R.V. |year=2020 |title=A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae |journal=Acta Palaeontologica Polonica |volume=65 |doi=10.4202/app.00751.2020|doi-access=free }}

|

| Jaw fragments

| Possibly synonymous with Coloborhynchus{{Cite journal |last1=Smith |first1=Roy E. |last2=Ibrahim |first2=Nizar |last3=Longrich |first3=Nicholas |last4=Unwin |first4=David M. |last5=Jacobs |first5=Megan L. |last6=Williams |first6=Cariad J. |last7=Zouhri |first7=Samir |last8=Martill |first8=David M. |date=September 2023 |title=The pterosaurs of the Cretaceous Kem Kem Group of Morocco |journal=PalZ |language=en |volume=97 |issue=3 |pages=519–568 |doi=10.1007/s12542-022-00642-6 |issn=0031-0220|doi-access=free |bibcode=2023PalZ...97..519S }}

|style="background:#E6E6E6;"|

rowspan="3" |Ornithocheirus

|O. simus{{cite journal |last1=Rodrigues |first1=Taissa |last2=Kellner |first2=Alexander Wilhelm Armin |name-list-style=amp |year=2013 |title=Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England |journal=ZooKeys |issue=308 |pages=1–112 |doi=10.3897/zookeys.308.5559 |pmc=3689139 |pmid=23794925 |doi-access=free|bibcode=2013ZooK..308....1R }}Unwin D.M., 2001, "An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England", Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189–221

|

|

|

|

style="background:#E6E6E6;"| "Ornithocheirus" denticulatus

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

style="background:#E6E6E6;"| "Ornithocheirus" polyodon

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

Ornithostoma{{cite journal |author=Averianov A.O. |year=2012 |title=Ornithostoma sedgwicki – valid taxon of azhdarchoid pterosaurs |journal=Proceedings of the Zoological Institute RAS |volume=316 |issue=1 |pages=40–49|doi=10.31610/trudyzin/2012.316.1.40 }}{{Cite journal |last1=Smith |first1=Roy E. |last2=Martill |first2=David M. |last3=Unwin |first3=David M. |last4=Steel |first4=Lorna |date=February 2021 |title=Edentulous pterosaurs from the Cambridge Greensand (Cretaceous) of eastern England with a review of Ornithostoma Seeley, 1871 |url=https://linkinghub.elsevier.com/retrieve/pii/S0016787820300997 |journal=Proceedings of the Geologists' Association |language=en |volume=132 |issue=1 |pages=110–126 |doi=10.1016/j.pgeola.2020.10.004|bibcode=2021PrGA..132..110S }}

|O. sedgwicki

|

|Jaw fragments

|Azhdarchoid pterosaur

|

Indeterminate azhdarchoid

|

|

| Jaw fragments

| Likely represents a distinct taxon from Ornithostoma{{cite journal|last1=Smith|first1=Roy E.|last2=Martill|first2=David M.|last3=Zouhri|first3=Samir|year=2023|title=Distinctive azhdarchoid pterosaur jaws from the mid-Cretaceous Cambridge Greensand of eastern England and the Kem Kem Group of Morocco|journal=Proceedings of the Geologists' Association|volume=134|issue=3|pages=269–275|doi=10.1016/j.pgeola.2023.03.002 |doi-access=free}}

= Ichthyosaurs =

class="wikitable"

! colspan="6" |Ichthyosaurs

Genus

!Species

!Locality

!Material

!Notes

!Images

Cetarthrosaurus

|C. walkeri{{cite journal |author=Valentin Fischer |author2=Nathalie Bardet |author3=Myette Guiomar |author4=Pascal Godefroit |name-list-style=amp |year=2014 |title=High Diversity in Cretaceous Ichthyosaurs from Europe Prior to Their Extinction |journal=PLOS ONE |volume=9 |issue=1 |pages=e84709 |bibcode=2014PLoSO...984709F |doi=10.1371/journal.pone.0084709 |pmc=3897400 |pmid=24465427 |doi-access=free}}

|

|

|

|

Maiaspondylus

|M. cantabrigiensis

|

|

|

|

Pervushovisaurus

|P. campylodon

|

|

|

|

Sisteronia

|S. seeleyi

|

|

|

|

= Lepidosauria =

{{Paleobiota-key-compact}}

class="wikitable"

! colspan="6" |Lepidosauria

Genus

!Species

!Locality

!Material

!Notes

!Images

style="background:#E6E6E6;"| Patricosaurus

|style="background:#E6E6E6;"| P. merocratusP. M. Barrett and S. E. Evans. 2002. A reassessment of the Early Cretaceous reptile 'Patricosaurus merocratus' Seeley from the Cambridge Greensand, Cambridgeshire, UK. Cretaceous Research 23:231-240

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"| Chimeric specimen

|style="background:#E6E6E6;"|

Invertebrates

= Ammonites =

  • Salaziceras salazacenseC. W. Wright and W. J. Kennedy. 1979. Origin and evolution of the Cretaceous micromorph ammonite family Flickiidae. Palaeontology 22:685-704

See also

References