Clavatula
{{Short description|Genus of gastropods}}
{{Automatic taxobox
|fossil_range =
|image = Clavatula muricata 004.jpg
|image_caption = Apertural view of a shell of Clavatula muricata
|taxon = Clavatula
|authority = Lamarck, 1801
|type_species= Clavatula coronata
|type_species_authority = Lamarck, 1801
|synonyms_ref =
|synonyms =
- Clavatula (Clavatula) Lamarck, 1801 ·
- Pleurotoma (Clavatula) Lamarck, 1801
|display_parents = 3
}}
Clavatula is a genus of sea snails, marine gastropod mollusks in the family Clavatulidae.{{WRMS species|204726|Clavatula Lamarck, 1801||6 April 2011}}{{cite journal |author1=P. Bouchet |author2=Yu. I. Kantor |author3=A. Sysoev |author4=N. Puillandre |year=2011 |title=A new operational classification of the Conoidea (Gastropoda) |journal=Journal of Molluscan Studies |volume=77 |issue=3 |pages=273–308 |doi=10.1093/mollus/eyr017|doi-access=free }}
Description
In this genus, the shell is fusiform, with a well-produced spire. The whorls are coronated with tubercles or short spines at the suture;. The aperture is oval. The columellar lip is smooth, arcuate, and callous behind. The outer lip is thin and arcuated, with the anal sinus situated below the sutural coronal. The operculum is semioval with its nucleus about the middle, on the inner side.
The foot is large, short and obtuse behind. The eyes are placed externally near the extremity of the tentacles. The radula contains a small unicuspid central tooth with single sharp laterals (formula :1-1-1).
The species mostly inhabits the west coast of Africa.George Washington Tryon, Manual of Conchology vol. VI p. 157; 1884
Species
Species within the genus Clavatula include:
{{div col|colwidth=20em}}
- † Clavatula agassizi (Bellardi, 1847)
- † Clavatula agatensis Bellardi, 1877
- Clavatula ahuiri Cossignani & Ardovini, 2014
- Clavatula albasulcata R. Aiken, 2021
- † Clavatula albucianensis Bellardi, 1877
- † Clavatula anaglypta (Millet, 1865)
- † Clavatula apenninica Bellardi, 1877
- † Clavatula apolloniae (R. Hoernes & Auinger, 1891)
- †Clavatula aralica (Luković, 1924) (an extinct species from the Eocene in western Kazakhstan (Tshegan Formation) ) Luković, M.T. (1924) Eocenska fauna molusaka iz oblasti izmedju aralskog i čalkarskoga jezera i njen značaj. Spomenik Srpske Kraljevske Akademije, 63, 1–74, 3 pls
- † Clavatula ariejansseni Harzhauser, Landau & R. Janssen, 2022
- Clavatula asamusiensis Nomura & Zinbo, 1940{{WRMS species|433246|Clavatula asamusiensis Nomura & Zinbo, 1940||15 April 2010}}
- †Clavatula asperulata (Lamarck, 1822) [http://www.fossilshells.nl/taxon_clavatulinae.html Fossilshells: Clavatula asperulata]
- † Clavatula atatuerki Harzhauser, Landau & R. Janssen, 2022
- † Clavatula auingeri Finlay, 1927
- † Clavatula barnabasi Kovács, Leél-Őssy & Vicián, 2023
- † Clavatula basilica Bellardi, 1877
- † ''Clavatula beauforti'' Oostingh, 1938
- † Clavatula berauensis Beets, 1941
- † Clavatula bicarinata Bellardi, 1877
- Clavatula bimarginata (Lamarck, 1822){{WRMS species|224384|Clavatula bimarginata (Lamarck, 1822)||15 April 2010}}
- † Clavatula boreoromana Kautsky, 1925
- † Clavatula burdigalensis Cossmann, 1895
- Clavatula caerulea (Weinkauff & Kobelt, 1875){{WRMS species|433247|Clavatula caerulea (Weinkauff & Kobelt, 1875)||15 April 2010}}
- † Clavatula camillae (R. Hoernes & Auinger, 1891)
- †Clavatula carinifera Grateloup, 1832 [http://www.fossilshells.nl/taxon_clavatulinae.html Fossilshells: Clavatula carinifera]
- † Clavatula circumclusa Bellardi, 1877
- Clavatula colini Von Maltzan, 1883{{WRMS species|433249|Clavatula colini Von Maltzan, 1883||15 April 2010}}
- † Clavatula complanata Bellardi, 1877
- †Clavatula concatenata (Grateloup, 1832) [http://www.fossilshells.nl/taxon_clavatulinae.html Fossilshells: Clavatula concatenata]
- Clavatula congoensis Nolf & Verstraeten, 2008{{WRMS species|433250|Clavatula congoensis Nolf & Verstraeten, 2008||15 April 2010}}
- † Clavatula consimilis Bellardi, 1877
- † Clavatula coppii Bellardi, 1877
- Clavatula coronata Lamarck, 1801{{WRMS species|433251|Clavatula coronata Lamarck, 1801||15 April 2010}}
- Clavatula cossignanii Ardovini, 2004{{WRMS species|388588|Clavatula cossignanii Ardovini, 2004||15 April 2010}}
- † Clavatula danuvii Vicián, Z. Kovács & Stein, 2019
- Clavatula debilis (R.B. Hinds, 1843) (taxon inquirendum)
- Clavatula decorata Sowerby III, 1916{{WRMS species|433252|Clavatula decorata Sowerby III, 1916||15 April 2010}}
- † Clavatula defossa Bellardi, 1877
- † Clavatula defrancii Bellardi, 1877
- Clavatula diadema (Kiener, 1840){{WRMS species|225503|Clavatula diadema (Kiener, 1840)||15 April 2010}}
- † Clavatula djocdjocartae (K. Martin, 1885)
- † Clavatula dodecapunctata Vera-Peláez & Lozano-Francisco, 2001
- † Clavatula dujardini Peyrot, 1938
- † Clavatula eichwaldi Bellardi, 1877
- † Clavatula excavata Bellardi, 1877
- Clavatula filograna Odhner, 1923{{WRMS species|433254|Clavatula filograna Odhner, 1923||15 April 2010}}
- Clavatula flammulata Knudsen, 1952{{WRMS species|225505|Clavatula flammulata Knudsen, 1952||15 April 2010}}
- † Clavatula flexicosta Bellardi, 1877
- † Clavatula francisci (Toula, 1901) (accepted > unreplaced junior homonym)
- Clavatula gabonensis Melvill, 1923{{WRMS species|433255|Clavatula gabonensis Melvill, 1923||15 April 2010}}
- † Clavatula geniculata Bellardi, 1877
- † Clavatula glaberrima J.P.S. Grateloup, 1832 https://www.bagniliggia.it/WMSD/HtmSpecies/8038000021.htm WMSDB: Clavatula glaberrima]
- †Clavatula glaberrima praecedens (Peyrot, 1931) [http://www.fossilshells.nl/taxon_clavatulinae.html Fossilshells: Clavatula glaberrima praecedens]
- † Clavatula gothica Bellardi, 1877
- Clavatula gracilior Sowerby II, 1870{{WRMS species|433256|Clavatula gracilior Sowerby II, 1870||15 April 2010}}
- † Clavatula gracilis Erünal-Erentöz, 1958
- † Clavatula gradata (Defrance, 1826)
- Clavatula hattenbergeri Nolf & Verstraeten, 2008{{WRMS species|433257|Clavatula hattenbergeri Nolf & Verstraeten, 2008||15 April 2010}}
- Clavatula helena Bartsch, 1915{{WRMS species|215594|Clavatula helena Bartsch, 1915||15 April 2010}}
- †Clavatula helwerdae Ceulemans, Van Dingenen & Landau, 2018
- † Clavatula heros (Mayer, 1858)
- † Clavatula hirmetzli Z. Kovács & Vicián, 2021
- †Clavatula iberoechinata Vera-Peláez & Lozano-Francisco, 2001
- Clavatula imperialis Lamarck, 1816{{WRMS species|224445|Clavatula imperialis Lamarck||15 April 2010}}
- † Clavatula implexa Bellardi, 1877
- † Clavatula inedita Bellardi, 1877
- † Clavatula inornata Bellardi, 1877
- † Clavatula interrupta (Brocchi, 1814) (accepted > unreplaced junior homonym)
- † Clavatula irisae Harzhauser, Landau & R. Janssen, 2022
- † Clavatula isseli Bellardi, 1877
- † Clavatula istvani Kovács, Leél-Őssy & Vicián, 2023
- † Clavatula jarzynkae Harzhauser, Landau & R. Janssen, 2022
- † Clavatula juliae (R. Hoernes & Auinger, 1891)
- † Clavatula justinae (R. Hoernes & Auinger, 1891)
- Clavatula knudseni Nolf & Verstraeten, 2007{{WRMS species|433258|Clavatula knudseni Nolf & Verstraeten, 2007||15 April 2010}}
- Clavatula kraepelini (Strebel, 1914){{WRMS species|225508|Clavatula kraepelini (Strebel, 1914)||15 April 2010}}
- † Clavatula labiolirata Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013
- † Clavatula laciniata Bellardi, 1877
- † Clavatula landaui Vera-Peláez & Lozano-Francisco, 2001
- Clavatula lelieuri (Récluz, 1851){{WRMS species|225510|Clavatula lelieuri (Récluz, 1851)||15 April 2010}}
- † Clavatula ligeriana (Peyrot, 1938)
- Clavatula malala Bozzetti, 2018
- † Clavatula manzonii Bellardi, 1877
- † Clavatula mariae (R. Hoernes & Auinger, 1891)
- Clavatula martensi Von Maltzan, 1883{{WRMS species|433260|Clavatula martensi Von Maltzan, 1883||15 April 2010}}
- † Clavatula martinelli Vera-Peláez & Lozano-Francisco, 2001
- Clavatula matthiasi Nolf, 2008{{WRMS species|433261|Clavatula matthiasi Nolf, 2008||15 April 2010}}
- Clavatula milleti (Petit de la Saussaye, 1851){{WRMS species|433262|Clavatula milleti (Petit de la Saussaye, 1851)||15 April 2010}}
- Clavatula muricata (Lamarck, 1822){{WRMS species|225513|Clavatula muricata (Lamarck, 1822)||15 April 2010}}
- Clavatula mystica (Reeve, 1843){{WRMS species|345762|Clavatula mystica (Reeve, 1843)||15 April 2010}}
- Clavatula nathaliae Nolf, 2006{{WRMS species|390482|Clavatula nathaliae Nolf, 2006||15 April 2010}}
- † Clavatula neogradata Glibert, 1954
- † Clavatula nodosa Bellardi, 1877
- † Clavatula obtruta (Millet, 1865)
- † Clavatula olgae (R. Hoernes & Auinger, 1891)
- † Clavatula oostinghi Beets, 1942
- † Clavatula orientoromana Báldi, 1960
- † Clavatula palatina Strausz, 1954
- † Clavatula pettkoi Biskupič, 2023
- Clavatula petzyae Boyer & Ryall, 2006{{WRMS species|390427|Clavatula petzyae Boyer & Ryall, 2006||15 April 2010}}
- † Clavatula pusilla Bellardi, 1877
- † Clavatula pyramidum Cuvillier, 1933 [https://science.mnhn.fr/institution/mnhn/collection/f/item/b44040?listIndex=12&listCount=19 MNHN, Paris: Clavatula pyramidum]
- Clavatula regia (Röding, 1798)
- †Clavatula reginae Hoernes & Auinger, 1891
- Clavatula retusa R.B. Hinds, 1843 (taxon inquirendum)
- † Clavatula romana (Defrance, 1826)
- Clavatula rubrifasciata (Reeve, 1845){{WRMS species|433265|Clavatula rubrifasciata (Reeve, 1845)||15 April 2010}}
- † Clavatula rugata Bellardi, 1877
- † Clavatula ruida Bellardi, 1877
- † Clavatula rusticula (Mayer-Eymar, 1886)
- Clavatula santhai Z. Kovács & Vicián, 2021 †
- † Clavatula sceauxensis Landau, Van Dingenen & Ceulemans, 2020
- † Clavatula seminuda Bellardi, 1877
- † Clavatula seyithasanensis Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013
- Clavatula smithi Knudsen, 1952{{WRMS species|433267|Clavatula smithi Knudsen, 1952||15 April 2010}}
- Clavatula solangeae Bozzetti, 2006{{WRMS species|390484|Clavatula solangeae Bozzetti, 2006||15 April 2010}}
- † Clavatula sophiae (R. Hoernes & Auinger, 1891)
- † Clavatula sorini Harzhauser, Landau & R. Janssen, 2022
- † Clavatula sotterii (Michelotti, 1847)
- Clavatula strebeli Knudsen, 1952{{WRMS species|225507|Clavatula strebeli Knudsen, 1952||15 April 2010}}
- Clavatula taxea (Röding, 1798){{WRMS species|215593|Clavatula taxea (Röding, 1798)||15 April 2010}}
- † Clavatula theodori (Toula, 1901)
- Clavatula tripartita (Weinkauff & Kobelt, 1876){{WRMS species|215592|Clavatula tripartita (Weinkauff & Kobelt, 1876)||15 April 2010}}
- † Clavatula turriculata (Grateloup, 1846)
- † Clavatula turriculoides Bellardi, 1877
- † Clavatula unicostata Bellardi, 1877
- † Clavatula veronicae (R. Hoernes & Auinger, 1891)
- Clavatula virgineus (Dillwyn, 1817){{WRMS species|433268|Clavatula virgineus (Dillwyn, 1817)||15 April 2010}}
- † Clavatula vitalisi Strausz, 1955
- Clavatula xanteni Nolf & Verstraeten, 2006{{WRMS species|390485|Clavatula xanteni Nolf & Verstraeten, 2006||15 April 2010}}
{{div col end}}
- Clavatula erecta W. H. Turton, 1932
;Species brought into synonymy :
- Clavatula albicans Hinds, 1843: synonym of Splendrillia albicans (Hinds, 1843)
- Clavatula amabilis Hinds, 1843: synonym of Tritonoturris amabilis (Hinds, 1843)
- Clavatula argillacea Hinds, 1843: synonym of Etrema argillacea (Hinds, 1843)
- Clavatula auriculifera Lamarck, J.B.P.A. de M. 1816: synonym of Clavus canalicularis (Röding, 1798)
- Clavatula bella Hinds, 1843: synonym of Bellacythara bella (Hinds, 1843)
- Clavatula boreointerrupta Kautsky, 1925:[http://www.fossilshells.nl/taxon_clavatulinae.html Fossilshells: Clavatula boreointerrupta] synonym of † Granulatocincta boreointerrupta (Kautsky, 1925)
- Clavatula candida Hinds, 1843: synonym of Glyphostoma candida (Hinds, 1843)
- Clavatula candida (Philippi, 1848): synonym of Pusionella valida (Dunker, 1852)
- Clavatula christianae Nolf, 2011: synonym of Paraclavatula christianae (Nolf, 2011) (original combination)
- Clavatula coelata Hinds, 1843: synonym of Crassispira coelata (Hinds, 1843)
- Clavatula crenularis Lamarck, 1816: synonym of Ptychobela nodulosa (Gmelin, 1791)
- Clavatula delphinae Nolf, 2008:{{WRMS species|433253|Clavatula delphinae Nolf, 2008||15 April 2010}} synonym of Paraclavatula delphinae (Nolf, 2008) (original combination)
- Clavatula echinata Lamarck, 1816: synonym of Clavus flammulatus Montfort, 1810
- Clavatula ericea Hinds, 1843: synonym of Lioglyphostoma ericea (Hinds, 1843)
- Clavatula fimbriata Hinds, 1843: synonym of Daphnellopsis fimbriata (Hinds, 1843)
- Clavatula flammea Hinds, 1843: synonym of Daphnella flammea (Hinds, 1843)
- Clavatula flavidula (Lamarck, 1822): synonym of Clathrodrillia flavidula (Lamarck, 1822)
- Clavatula fulva Hinds, 1843: synonym of Clavus fulva (Hinds, 1843)
- Clavatula gravis (Hinds, 1843): synonym of Makiyamaia gravis (Hinds, 1843)
- Clavatula griffithii Gray, 1834: synonym of Ptychobela griffithii (Gray, 1834)
- Clavatula haliplex Bartsch, 1915: synonym of Toxiclionella haliplex (Bartsch, 1915)
- Clavatula halistrepta Bartsch, 1915: synonym of Clionella halistrepta (Bartsch, 1915)
- Clavatula hottentota (E. A. Smith, 1882): synonym of Clavus hottentotus (E. A. Smith, 1882)
- Clavatula impressa Hinds, 1843: synonym of Kylix impressa (Hinds, 1843)
- Clavatula kraussii (E. A. Smith, 1877): synonym of Clionella kraussii (E. A. Smith, 1877)
- Clavatula laeta Hinds, 1843: synonym of Splendrillia laeta (Hinds, 1843)
- Clavatula lineata Lamarck, 1816: synonym of Perrona lineata (Lamarck, 1816)
- Clavatula lobatopsis Barnard, 1963: synonym of Clionella lobatopsis (Barnard, 1963)
- Clavatula luctuosa (Hinds, 1843):* [http://www.geokniga.org/bookfiles/geokniga-dali1909peru.pdf W.H. Dall (1909), Report on the collection of shells from Peru ;Proceedings of the United States National Museum, Vol. 37, pages 147-294, with Plates 20—28] synonym of Crassispira kluthi E. K. Jordan, 1936
- Clavatula marmarina (Watson, 1881):{{WRMS species|433259|Clavatula marmarina (Watson, 1881)||15 April 2010}} synonym of Fenimorea marmarina (Watson, 1881)
- Clavatula merita Hinds, 1843: synonym of Tenaturris merita (Hinds, 1843)
- Clavatula metula Hinds, 1843: synonym of Anarithma metula (Hinds, 1843)
- Clavatula micans Hinds, 1843: synonym of Globidrillia micans (Hinds, 1843)
- Clavatula neglecta Hinds, 1843: synonym of Glyphostoma neglecta (Hinds, 1843)
- Clavatula nitens Hinds, 1843: synonym of Agladrillia nitens (Hinds, 1843)
- Clavatula obesa Reeve:{{WRMS species|225090|Clavatula obesa Reeve||15 April 2010}} synonym of Perrona obesa (Reeve, 1842)
- Clavatula occata Hinds, 1843: synonym of Diptychophlia occata (Hinds, 1843)
- Clavatula pardalis Hinds, 1843: synonym of Anachis pardalis (Hinds, 1843)
- Clavatula parilis E. A. Smith, 1902: synonym of Clavatula tripartita (Weinkauff, 1876)
- Clavatula perronii (Reeve, 1843):{{WRMS species|433263|Clavatula perronii (Reeve, 1843)||15 April 2010}} synonym of Perrona perron (Gmelin, 1791)
- Clavatula pfefferi (Strebel, 1912): synonym of Fusiturris pfefferi (Strebel, 1912)
- Clavatula plumbea Hinds, 1843: synonym of Kurtziella plumbea (Hinds, 1843)
- Clavatula pseudomystica Nolf, 2008:{{WRMS species|433264|Clavatula pseudomystica Nolf, 2008||15 April 2010}} synonym of Paraclavatula pseudomystica (Nolf, 2008) (original combination)
- Clavatula pudica Hinds, 1843: synonym of Agladrillia pudica (Hinds, 1843)
- Clavatula pungens Gould, 1860: synonym of Veprecula pungens (Gould, 1860)
- Clavatula pyramis Hinds, 1843: synonym of Pseudorhaphitoma pyramis (Hinds, 1843)
- Clavatula quinteni Nolf & Verstraeten, 2006:{{WRMS species|390483|Clavatula quinteni Nolf & Verstraeten, 2006||15 April 2010}} synonym of Perrona quinteni (Nolf & Verstraeten, 2006)
- Clavatula quisqualis Hinds, 1843: synonym of Leptadrillia quisqualis (Hinds, 1843)
- Clavatula rava Hinds, 1843: synonym of Clathurella rava (Hinds, 1843)
- Clavatula rigida Hinds, 1843: synonym of Clathurella rigida (Hinds, 1843)
- Clavatula robusta Hinds, 1843: synonym of Cheungbeia robusta (Hinds, 1843)
- Clavatula rubida Hinds, 1843: synonym of Lienardia rubida (Hinds, 1843)
- Clavatula rubiginosa Hinds, 1843: synonym of Otitoma rubiginosa (Hinds, 1843)
- Clavatula rubrofasciata Reeve: synonym of Clavatula rubrifasciata (Reeve, 1845)
- Clavatula sacerdos (Reeve, 1845):{{WRMS species|433266|Clavatula sacerdos (Reeve, 1845)||15 April 2010}} synonym of Clavatula mystica (Reeve, 1843)
- Clavatula scalaris Hinds, 1843: synonym of Tritonoturris scalaris (Hinds, 1843)
- Clavatula sculpta Hinds, 1843: synonym of Rimosodaphnella sculpta (Hinds, 1843)
- Clavatula semicostata (Kiener, 1840): synonym of Clionella semicostata (Kiener, 1840)
- Clavatula sinensis Hinds, 1843: synonym of Crassispira sinensis (Hinds, 1843)
- Clavatula sinuata (Born, 1778): synonym of Clionella sinuata (Born, 1778)
- Clavatula spicata Hinds, 1843: synonym of Inquisitor spicata (Hinds, 1843)
- Clavatula spirata Lamarck:{{WRMS species|224447|Clavatula spirata Lamarck||15 April 2010}} synonym of Perrona spirata (Lamarck, 1816)
- Clavatula spurca Hinds, 1843: synonym of Etrema spurca (Hinds, 1843)
- Clavatula striata Gray, 1826: synonym of Epideira striata (Gray, 1826)
- Clavatula subspirata (von Martens, 1902):{{WRMS species|432627|Clavatula subspirata (Martens, 1902)||15 April 2010}} synonym of Perrona subspirata (von Martens, 1902)
- Clavatula subventricosa (E. A. Smith, 1877): synonym of Clionella subventricosa (E. A. Smith, 1877)
- Clavatula taxus : synonym of Clavatula taxea (Röding, 1798) (incorrect subsequent spelling)
- Clavatula tessellata Hinds, 1843: synonym of Kermia tessellata (Hinds, 1843)
- Clavatula tumida Sowerby II, 1870: synonym of Toxiclionella tumida (Sowerby II, 1870)
- Clavatula turriplana (G.B. Sowerby III, 1903): synonym of Turricula turriplana (Sowerby III, 1903)
References
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