Clavatulidae
{{Short description|Family of gastropods}}
{{Use dmy dates|date=January 2024}}
{{Automatic taxobox
| taxon = Clavatulidae
| image = Perrona semimarginata 01.jpg
| image_caption = Five views of a shell of † Tomellana semimarginata
| authority = Gray, 1853Gray J. E. (1853). "On the division of Ctenobranchous Gasteropodous Mollusca into larger groups and families". Annals and Magazine of Natural History (2)11: 124-133. Clavatulidae (spelled as Clavatulina) is on the page [https://archive.org/stream/annalsmagazineof11anna#page/128/mode/2up/ 128].
| synonyms =
- Pusionellinae Gray, 1853
- Clionellidae Stimpson, 1865
- Melatomidae Gill, 1871
- Turriculinae Powell, 1942 (inv.)
| subdivision_ranks = Genera
| subdivision = See text
}}
Clavatulidae is a taxonomic family of sea snails, marine gastropod mollusks in the superfamily Conoidea.{{Cite journal
|author1=Bouchet, P. |author2=Rocroi, J.-P.
| year = 2005
| title = Classification and Nomenclator of Gastropod Families
| journal = Malacologia
| volume =47
| issue = 1–2
}} The family is not well differentiated morphologically.
Clavatulidae was raised, based on cladistic analysis, from subfamily to the family level by Rosenberg in 1998. It is no longer regarded as a subfamily of Turridae by several malacologists (Kantor, Sysoev).[http://www.marinespecies.org/aphia.php?p=taxdetails&id=411813 World Register of Marine Species: Clavatulidae]. Retrieved 1 April 2010.{{cite journal |author1=P. Bouchet |author2=Yu. I. Kantor |author3=A. Sysoev |author4=N. Puillandre |year=2011 |title=A new operational classification of the Conoidea (Gastropoda) |journal=Journal of Molluscan Studies |volume=77 |issue=3 |pages=273–308 |doi=10.1093/mollus/eyr017|doi-access=free |url=https://hal.archives-ouvertes.fr/hal-02458082/file/Bouchet%20et%20al%202011%20J.%20Moll.%20Stud.pdf }}Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). {{ISBN|0-915826-22-4}}. XII, 195 pp. This family has no subfamilies.
General characteristics
This family consists of species with a medium-sized to rather large, fusiform shell. The oblong, pointed spire is rather high. The aperture is oval and mostly white. The operculum has a medio-lateral nucleus. The siphonal canal varies between rather short (e.g. Pusionella compacta) to moderately long and slightly incurved (e.g. Fusiturris undatiruga). The anal sinus varies from very shallow to rather deep. The outer lip can be slightly incurved and serrated on its side. The subsutural ramp is usually well developed. The sculpture of the shell in this family shows various forms, going from a rather smooth surface (e.g. Gemmuloborsonia colorata) to being finely ribbed longitudinally and striated transversally. The stenoglossan radula has the formula 1-(1-R-1)-1 George Washington Tryon, Manual of Conchology vol. VI Clavatulinae p. 228 ; 1884
Genera
Genera in the family Clavatulidae include:
- Benthoclionella Kilburn, 1974
- Caliendrula Kilburn, 1985
- Clavatula Lamarck, 1801
- Clionella Gray, 1847
- † Granulatocincta Harzhauser, Landau & R. Janssen, 2022
- † Hemisurcula Casey, 1904
- Makiyamaia Kuroda, 1961
- † Megaclavatula Harzhauser, Landau & R. Janssen, 2022
- † Neoperrona Harzhauser, Landau & R. Janssen, 2022
- † Orthosurcula T. L. Casey, 1904
- Pagodaturris Kantor, Fedosov & Puillandre, 2018
- Paraclavatula Kantor, Horro, Rolán & Puillandre, 2018
- Perrona Schumacher, 1817
- Pusionella Gray, 1847
- Scaevatula Gofas, 1990
- † Striopusionella Harzhauser, Landau & R. Janssen, 2022
- Tomellana Wenz, 1943
- Toxiclionella Powell, 1966
- † Trachelochetus Cossmann, 1889
- Trachydrillia Nolf & Swinnen, 2010
- Turricula Schumacher, 1817
; Genera moved to other families:
- Danilacarina Bozzetti, 1997, moved to Cochlespiridae
- Iwaoa Kuroda, 1953, moved to Horaiclavidae
;Genera brought into synonymy:
- Melatoma Swainson, 1840: synonym of Clionella Gray, 1847
- Netrum Philippi, 1850: synonym of Pusionella Gray, 1847
- Surcula H. Adams & A. Adams, 1853: synonym of Turricula Schumacher, 1817
- Tomella Swainson, 1840: synonym of Tomellana Wenz, 1943
- †Trachelochetus Cossmann, 1889:[https://www.researchgate.net/profile/Jean-Michel_Pacaud/publication/259676619_Revision_des_Mollusques_paleogenes_du_Bassin_de_Paris_IV_Liste_systematique_actualisee/links/0f31752d5081f30819000000.pdf Pacaud, Jean—Michel, and Jacques Le Renard. "Révision des mollusques Paléogènes du Bassin de Paris. IV-Liste systématique actualisée." Cossmanniana 3.4 (1995): 155-187.] synonym of †Clavatula (Trachelochetus) Cossmann, 1889 represented as Clavatula Lamarck, 1801
- Tyrrhenoturris Coen, 1929: synonym of Fusiturris Thiele, 1929
Ecology
Species from family Clavatulidae are omnivores, predators and scavengers.WoRMS (2009). Clavatulinae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2009) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=153900 on 2010-04-01
References
{{Reflist}}
External links
{{Commons category|Clavatulidae}}
{{Wikispecies}}
- [http://www.bagniliggia.it/WMSD/HtmFamily/CLAVATULIDAELS.htm Worldwide Mollusc Species Data Base: family Clavatulidae]
{{Taxonbar|from=Q5129591}}