Helicoidea

{{Short description|Superfamily of gastropods}}

{{Use dmy dates|date=November 2021}}

{{Automatic taxobox

| image = Bradybaena similaris 60096358.jpg

| image_caption = Bradybaena similaris

| taxon = Helicoidea

| authority = Rafinesque, 1815

| subdivision_ranks = Families

| subdivision = See text.

}}

Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Stylommatophora.

Taxonomy

= 2017 taxonomy and latest developments<ref name=":0">{{Cite journal |last1=Bouchet |first1=Philippe |last2=Rocroi |first2=Jean-Pierre |last3=Hausdorf |first3=Bernhard |last4=Kaim |first4=Andrzej |last5=Kano |first5=Yasunori |last6=Nützel |first6=Alexander |last7=Parkhaev |first7=Pavel |last8=Schrödl |first8=Michael |last9=Strong |first9=Ellen E. |date=2017 |title=Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families |url=http://www.bioone.org/doi/10.4002/040.061.0201 |journal=[[Malacologia]] |language=en |volume=61 |issue=1–2 |pages=1–526 |doi=10.4002/040.061.0201 |issn=0076-2997|url-access=subscription }}</ref><ref>{{Cite journal |last1=Sei |first1=Makiri |last2=Robinson |first2=David G |last3=Geneva |first3=Anthony J |last4=Rosenberg |first4=Gary |date=2017-11-23 |title=Doubled helix: Sagdoidea is the overlooked sister group of Helicoidea (Mollusca: Gastropoda: Pulmonata) |url=http://academic.oup.com/biolinnean/article/122/4/697/4210279 |journal=Biological Journal of the Linnean Society |language=en |volume=122 |issue=4 |pages=697–728 |doi=10.1093/biolinnean/blx082 |issn=0024-4066|doi-access=free }}</ref><ref>{{Cite journal |date=2019-03-29 |title=A New Species of the Genus Bunnya H. B. Baker, 1942 (Helicoidea) from MÉXico |url=https://bioone.org/journals/malacologia/volume-62/issue-2/040.062.0204/A-New-Species-of-the-Genus-Bunnya-H-B-Baker/10.4002/040.062.0204.full |journal=Malacologia |volume=62 |issue=2 |pages=237 |doi=10.4002/040.062.0204 |issn=0076-2997|url-access=subscription }}</ref><ref>{{Cite journal |last1=Saadi |first1=Ahmed J. |last2=Wade |first2=Christopher M. |date=2019 |title=Resolving the basal divisions in the stylommatophoran land snails and slugs with special emphasis on the position of the Scolodontidae |url=https://linkinghub.elsevier.com/retrieve/pii/S1055790319301939 |journal=[[Molecular Phylogenetics and Evolution]] |language=en |volume=139 |pages=106529 |doi=10.1016/j.ympev.2019.106529|pmid=31176967 }}</ref> =

The Bouchet et al. 2017 nomenclator provides an up to date system of Helicoidea. The system is in some parts preliminary, as the authors relied on unpublished (as of 2023) phylogenomic study, which did not include all New World taxa. They classified Epiphragmophoridae, Helminthoglyptidae, Humboldtianidae, Monadeniidae and Xanthonychidae (provisionally also Lysinoidae and Echinichidae) as subfamilies of the last taxon, because there are no deep splits between them in the cited unpublished study (see also Koene & Schulenburg 2005{{Cite journal |last1=Koene |first1=Joris M |last2=Schulenburg |first2=Hinrich |date=2005 |title=Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails |journal=BMC Evolutionary Biology |volume=5 |issue=1 |pages=25 |doi=10.1186/1471-2148-5-25 |pmc=1080126 |pmid=15799778 |doi-access=free }}).

Cepolidae, Labyrinthidae and Thysanophoridae constitute the sister group of the remaining Helicoidea.

=2015 taxonomy=

Razkin et al. (2015)Razkin O., Gómez-Moliner B. J., Prieto C. E., Martínez-Ortí A., Arrébola J. R., Chueca L. J. & Madeira M. J. (2015). "Molecular phylogeny of the western Palearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution 83: 99–117. {{doi|10.1016/j.ympev.2014.11.014}}. reorganized classification into monophyletic taxa according to the molecular phylogeny. This study is focus on Western Palaearctic species. The Hygromiidae s.l. family was divided into three families, Canariellidae, Geomitridae and Hygromiidae. Moreover, the family Cochlicellidae was including within the Geomitridae family as a tribe (Cochlicellini).

The classification proposed for Western Palaearctic Helicoidea is as follows:

{{expand section|date=April 2019}}

=2012 taxonomy=

Thompson & Naranjo-García (2012) described a new family Echinichidae and placed it to the superfamily Xanthonychoidea.Thompson F. G. & Naranjo-García E. (2012). "Echinichidae, a new family of dart-bearing helicoid slugs from Mexico, with the description of a new genus and three new species (Gastropoda: Pulmonata: Xanthonychoidea)". Archiv für Molluskenkunde 141(2): 197–208. {{doi|10.1127/arch.moll/1869-0963/141/197-208}}. [http://www.schweizerbart.de/resources/downloads/paper_previews/79370.pdf preview]. Therefore, the family Xanthonychidae was moved from Helicoidea to Xanthonychoidea.

=2005 taxonomy=

There are 19 families within the superfamily Helicoidea according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.{{Bouchet 2005}}Poppe G. T. & Tagaro S. P. (2006). "The new classification of Gastropods according to Bouchet & Rocroi, 2005". Visaya, février 2006: 10 pp. [http://www.journal-malaco.fr/bouchet&rocroi_2005_Visaya.pdf PDF] {{webarchive|url=https://web.archive.org/web/20070927101354/http://www.journal-malaco.fr/bouchet%26rocroi_2005_Visaya.pdf|date=27 September 2007}}.

According to H. Nordsieck[http://www.clausilia.de/media//DIR_64901/04_2006_helicoidea.pdf Higher classification of the Helicoidea and the molecular analyses of their phylogeny (by Hartmut Nordsieck)] {{webarchive|url=https://web.archive.org/web/20151129213842/http://www.clausilia.de/media//DIR_64901/04_2006_helicoidea.pdf|date=29 November 2015}} the family Xanthonychidae (sensu Hausdorf & Bouchet) is probably polyphyletic (contains several different lineages) and therefore should be divided into several families.

File:Helicella_macedonica.jpg

References