Hieracium#Species

{{short description|Genus of flowering plants}}

{{Automatic taxobox

|image=Yellow Hawkweed.jpg

|image_caption=Hieracium caespitosum

|display_parents=3

|taxon=Hieracium

|authority=L.

|synonyms=

  • Chlorocrepis{{cite web |url=http://www.bgbm.org/IOPI/GPC/PTaxonDetail.asp?PTRefFk=&NameCache=Hieracium#1 |title=Vascular Plants of Russia and Adjacent Countries as of 26.10.96 |access-date=2007-12-17 |author=International Organization for Plant Information (IOPI) |author-link=IPNI |work=Provisional Global Plant Checklist |publisher=International Organization for Plant Information (IOPI) |url-status=dead |archive-url=https://web.archive.org/web/20131213223601/http://www.bgbm.org/IOPI/GPC/PTaxonDetail.asp?PTRefFk=&NameCache=Hieracium#1 |archive-date=2013-12-13}}

  • Crepidopsis Arv.-Touv.
  • Pilosella Vaill.{{cite web |url=http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?5670 |title=Genus: Hieracium L. |access-date=2007-12-23 |author=Germplasm Resources Information Network |author-link=Germplasm Resources Information Network |work=(GRIN) Online Database |publisher=USDA, ARS, National Genetic Resources Program |archive-url=https://web.archive.org/web/20131213222733/http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?5670 |archive-date=2013-12-13 |url-status=dead}}

}}

Hieracium ({{IPAc-en|h|aɪ|.|ə|ˈ|r|æ|s|i|əm}}),{{cite web |url=http://www.calflora.net/bloomingplants/pronunciationguide.html |title=Guide to the Pronunciation of Specific, Generic and Family Names |access-date=2007-12-22 |work=Southern California Wildflowers |archive-date=2007-11-02 |archive-url=https://web.archive.org/web/20071102051547/http://www.calflora.net/bloomingplants/pronunciationguide.html |url-status=live }}

known by the common name hawkweed{{cite web |url=https://plants.usda.gov/java/ClassificationServlet?source=profile&symbol=HIERA&display=31 |title=Classification for Kingdom Plantae Down to Genus Hieracium L. |access-date=2007-12-18 |author=Natural Resources Conservation Service |author-link=Natural Resources Conservation Service |year=2007 |work=The PLANTS Database |publisher=USDA, Baton Rouge, LA 70874-4490 USA. |archive-date=2019-06-21 |archive-url=https://web.archive.org/web/20190621042744/https://plants.usda.gov/java/ClassificationServlet?source=profile&symbol=HIERA&display=31 |url-status=live }} and classically as {{Transliteration|grc|hierakion}} (from ancient Greek ἱέραξ, {{Transliteration|grc|hierax}} 'hawk'),{{cite web |url=http://www.calflora.net/botanicalnames/pageHI-HY.html |title=HI-HY |access-date=2007-12-26 |last=Charters |first=Michael L. |work=California Plant Names: Latin and Greek Meanings and Derivations A Dictionary of Botanical Etymology |archive-date=2016-04-16 |archive-url=https://web.archive.org/web/20160416061442/http://www.calflora.net/botanicalnames/pageHI-HY.html |url-status=live }}

is a genus of flowering plant in the family Asteraceae, and closely related to dandelion (Taraxacum), chicory (Cichorium), prickly lettuce (Lactuca) and sow thistle (Sonchus),

{{cite web |url=http://www.montana.edu/wwwpb/pubs/mt9816.html |title=Orange and Meadow Hawkweed, 199816 |access-date=2007-12-22 |last=Cooperative extension service |first=Matthew J. Rinella and Roger L. Sheley |author-link=Cooperative extension service |date=December 2002 |edition=Reprint |work=MontGuide fact sheet |publisher=Department of Land Resources and Environmental Sciences, Montana State University - Bozeman |url-status=dead |archive-url=https://web.archive.org/web/20070814175819/http://www.montana.edu/wwwpb/pubs/mt9816.html |archive-date=2007-08-14}}

which are part of the tribe Cichorieae. Hawkweeds, with their 10,000+ recorded species and subspecies,{{cite web |url=http://www.ipni.org/ipni/advPlantNameSearch.do?find_family=&find_genus=Hieracium |title=Plant Name Search Results |access-date=2007-12-17 |author=International Organization for Plant Information (IOPI) |author-link=IPNI |work=International Plant Names Index |archive-date=2012-10-10 |archive-url=https://web.archive.org/web/20121010145839/http://www.ipni.org/ipni/advPlantNameSearch.do?find_family=&find_genus=Hieracium |url-status=live }} do their part to make Asteraceae the second largest family of flowering plants.

{{cite book |first=Theodore F. |last=Niehaus |others=Illustrations by Charles L. Ripper |title=Pacific States Wildflowers |year=1976 |series=Peterson Field Guides |publisher=Houghton Mifflin Company |location=New York |isbn=0-395-91095-1 |pages=[https://archive.org/details/fieldguidetopaci00theo_0/page/102 102, 220] |url=https://archive.org/details/fieldguidetopaci00theo_0/page/102 }}

Some botanists group all these species or subspecies into approximately 800 accepted species,

{{cite web |url=http://www.bgbm.org/IOPI/GPC/PTaxonDetail.asp?PTRefFk=&NameCache=Hieracium |title=Details for: Hieracium |access-date=2007-12-17 |author=International Organization for Plant Information (IOPI) |author-link=IPNI |work=Provisional Global Plant Checklist |url-status=dead |archive-url=https://web.archive.org/web/20131213223601/http://www.bgbm.org/IOPI/GPC/PTaxonDetail.asp?PTRefFk=&NameCache=Hieracium |archive-date=2013-12-13}}

while others prefer to accept several thousand species. Since most hawkweeds reproduce exclusively asexually by means of seeds that are genetically identical to their mother plant (apomixis or agamospermy), clones or populations that consist of genetically identical plants are formed and some botanists (especially in UK, Scandinavia and Russia) prefer to accept these clones as good species (arguing that it is impossible to know how these clones are interrelated) whereas others (mainly in Central Europe and USA) try to group them into a few hundred more broadly defined species. What is here treated as the single genus Hieracium is now treated by most European experts as two different genera, Hieracium and Pilosella, with species such as Hieracium pilosella, Hieracium floribundum and Hieracium aurantiacum referred to the latter genus. Many members of the genus Pilosella reproduce both by stolons (runners like those of strawberries) and by seeds, whereas true Hieracium species reproduce only by seeds. In Pilosella, many individual plants are capable of forming both normal sexual and asexual (apomictic) seeds, whereas individual plants of Hieracium only produce one kind of seeds. Another difference is that all species of Pilosella have leaves with smooth (entire) margins whereas most species of Hieracium have distinctly dentate to deeply cut or divided leaves.

{{cquote|A dry roadside dotted with small, ¾ inch red orange flowers, interspersed with very similar yellow ones, and often the white of daisies, is a good sign that you are in Hawkweed country.|author=Marion Edsall

{{cite book |last=Edsall |first=Marion |title=Roadside Plants and Flowers A Traveler's Guide to the Midwest and Great Lakes Area |url=https://archive.org/details/roadsideplantsfl0000edsa |url-access=registration |orig-year=1985 |others=Cover design: Bruce Gore |date=2007-12-15 |publisher=University of Wisconsin Press |location=Madison, Wisconsin |isbn=978-0-299-09704-2 |id=Dewey 582.0977 |pages=[https://archive.org/details/roadsideplantsfl0000edsa/page/46 46] |quote=A dry roadside dotted with small, 3/4 inch red orange flowers, interspersed with very similar yellow ones, and often the white of daisies, is a good sign that you are in Hawkweed country.}} }}

Description

=Flowers and flower-heads=

Hieracium or hawkweeds, like others in the family Asteraceae, mostly have yellow,{{cite web |url=http://www.botanical.com/botanical/mgmh/h/hawwee07.html |title=Hawkweed, Wood |access-date=2007-12-16 |author=Mrs M. Grieve |author-link=Maud Grieve |year=1933 |work=A Modern Herbal |publisher=botanical.com |archive-date=2008-05-10 |archive-url=https://web.archive.org/web/20080510181619/http://www.botanical.com/botanical/mgmh/h/hawwee07.html |url-status=live }}

tightly packed flower-heads of numerous small flowers

but, unlike daisies and sunflowers in the same family, they have not two kinds of florets but only strap-shaped (spatulate) florets, each one of which is a complete flower in itself, not lacking stamens,

and joined to the stem by leafy bracts. As in other members of the tribe Cichorieae, each ray corolla is tipped by 3 to 5 teeth.

=Bracts, stems and leaves=

Erect single, glabrous or hairy stems, sometimes branched away from the point of attachment, sometimes branched throughout.

The hairiness of hawkweeds can be very complex: from surfaces with scattered to crowded, tapered, whiplike, straight or curly, smooth to setae; "stellate-pubescent" or surfaces with scattered to crowded, dendritically branched (often called, but seldom truly, "stellate") hairs; and "stipitate-glandular" or surfaces with scattered to crowded gland-tipped hairs mostly. Surfaces of stems, leaves, peduncles, and phyllaries may be glabrous or may bear one, two, or all three of the types of hairs mentioned above.{{cite web |url=http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=115448 |title=Hieracium in Flora of North America |access-date=2007-12-16 |author=John L. Strother |work=FNA Vol. 19, 20 and 21 |publisher=efloras.org |pages=Page 219, 278, 279 |archive-date=2008-04-21 |archive-url=https://web.archive.org/web/20080421045557/http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=115448 |url-status=live }}

Like the other members of the Chicory tribe, hawkweeds contain a milky latex.

Ecology

The large yellow underwing moth (Noctua pronuba) feeds on Hieracium species.

Distribution

Hieracium species are native to Africa, Asia, Europe, North America,{{cite web |url=https://plants.usda.gov/java/profile?symbol=HIERA |title=PLANTS Profile for Hieracium L. |access-date=2007-12-18 |author=Natural Resources Conservation Service |author-link=Natural Resources Conservation Service |year=2007 |work=The PLANTS Database |publisher=USDA, Baton Rouge, LA 70874-4490 USA. |archive-date=2008-10-10 |archive-url=https://web.archive.org/web/20081010020343/http://plants.usda.gov/java/profile?symbol=HIERA |url-status=live }} Central America and South America.

Species

{{main|List of Hieracium species}}

The classification of Hieracium into species is notoriously difficult. One reason is the apomictic reproduction (in which plants asexually produce seeds), which tends to produce a lot of minor geographical variation. Over 9000 species names have been published in Hieracium but some botanists regard many of those as synonyms of larger species.

=Europe=

=North America=

The list below is a selection of species that have been accepted by the USDA Natural Resources Conservation Service and Canada.[http://www.for.gov.bc.ca/hfp/publications/00230/Hawkweed%20key_PNW_R3-June06.pdf Key to Identification of Invasive and Native Hawkweeks (Hieracium spp.) in the Pacific Northwest] {{Webarchive|url=https://web.archive.org/web/20130515233942/http://www.for.gov.bc.ca/hfp/publications/00230/Hawkweed%20key_PNW_R3-June06.pdf |date=2013-05-15 }}, BC Ministry of Forests

A more complete list is given in the list of Hieracium species.

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Some species are now placed in the genus Pilosella:{{Cite POWO|title=Pilosella Hill|id=30065694-2|access-date=2023-06-13|mode=cs1}}

Plant pest

All species of the genus Hieracium are classed as invasive species throughout New Zealand. They are banned from sale, propagation and distribution under the National Pest Plant Accord. Hieracium is a pasture weed that reduces available feed for livestock and displaces the indigenous plants.{{cite web |url=http://www.weedbusters.co.nz/weed_info/detail.asp?WeedID=119 |title=Hieracium species Detailed information sheet |publisher=The Weedbusters Management Committee (www.weedbusters.co.nz) |access-date=2010-05-02 |url-status=dead |archive-url=https://web.archive.org/web/20100515032635/http://weedbusters.co.nz/weed_info/detail.asp?WeedID=119 |archive-date=2010-05-15}} It is a particular threat in alpine ecosystems previously dominated by native tussocks, though it will colonise habitats from bare ground, to exotic pine forest, to native Southern Beech forest.{{cite web |url=http://www.doc.govt.nz/documents/science-and-technical/DSIS109.pdf |title=Genetic diversity in tussock hawkweed (Hieracium lepidulum) and use of allele frequencies for identifying patterns of spread |access-date=2013-09-06 |archive-date=2016-03-05 |archive-url=https://web.archive.org/web/20160305035152/http://www.doc.govt.nz/documents/science-and-technical/DSIS109.pdf |url-status=live }}

In the United States, many species of Hieracium have been introduced and all species present are considered noxious weeds in one or more states.{{cite web |url=http://plants.usda.gov/plant-profile?symbol=HIERA |title=Hieracium L._hawkweed_USDA NRCS |access-date=2013-09-06 |archive-date=2014-04-24 |archive-url=https://web.archive.org/web/20140424233413/http://plants.usda.gov/core/profile?symbol=HIERA |url-status=live }}

In Australia, hawkweeds are invasive pests in alpine regions, all species of Hieracium are listed or declared under various State Acts.{{Cite web |url=http://invasives.org.au/blog/hawkweeds-a-recent-discovery-in-victorias-alps-and-a-taxonomic-name-change/ |title=Hawkweeds: A recent discovery in Victoria's Alps and a taxonomic name change |date=10 July 2013 |access-date=6 March 2016 |archive-date=5 March 2016 |archive-url=https://web.archive.org/web/20160305132142/http://invasives.org.au/blog/hawkweeds-a-recent-discovery-in-victorias-alps-and-a-taxonomic-name-change/ |url-status=live }}

References

{{Reflist}}

Further reading

  • {{cite book |last=Espie |first=Peter |title=Hieracium in New Zealand: ecology and management |publisher=AgResearch |location=Mosgiel |year=2001 |isbn=0-478-20900-2}}
  • McCosh, D. and Rich, T.C.G. 209. Hieracium proximum (Caithness Hawkweed) in Ireland. Ir. Nat J. 30: 54.
  • Rich, T.C.G., Cotton, D.C.F., Hood, R.L.I.B., Houston, L., McCosh, J. and Jackson, M.B.W. 2009. Conservation of Ireland's biodiversity: status of the Irish endemic Hieracium basalticola Pugsley (Basalt Hawkweed) (Asteraceae). Ir. Nat J. 30: 79–89.