Paleobiota of the Djadochta Formation

{{Short description|Geological formation in Mongolia}}

File:Protoceratops specimen MPC-D 100 519.jpg

The Djadochta Formation (sometimes spelled Djadokhta, Djadokata, or Dzhadokhtskaya) is a geological formation in the Gobi Desert of Mongolia. It dates to the Campanian stage of the Late Cretaceous and is famous for its dinosaur fossils including Oviraptor, Protoceratops, and Velociraptor. It is also known for a high diversity of mammal and lizard fossils, and a complete catalogue of its fossil contents is shown below.

{{Paleobiota-key-compact}}

Vertebrates

=Amphibians=

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Gobiates

| Indeterminate

| Udyn Sayr

| "Partial skeleton with partial urostyle."{{cite journal|last1=Roček|first1=Z.|date=2008|title=The Late Cretaceous frog Gobiates from Central Asia: its evolutionary status and possible phylogenetic relationships|journal=Cretaceous Research|volume=29|issue=4|pages=577–591|doi=10.1016/j.cretres.2008.01.005|bibcode=2008CrRes..29..577R |url=http://rocek.gli.cas.cz/Publications/Gobiates.pdf}}

| A frog.

|

=Crocodylomorphs=

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Artzosuchus

| A. brachicephalus

| Udyn Sayr

| "Partial skull."{{cite journal|last1=Efimov|first1=N. B.|date=1983|title=Review of crocodilians from Mongolia|journal=Transactions of the Joint Soviet Mongolian Paleontological Expedition|volume=24|pages=79–96|language=ru}}

| A crocodylomorph.

|

rowspan="2" | Gobiosuchus

| G. kielanae

| Bayn Dzak

| "Multiple specimens with partial skulls and skeletons."{{cite journal|last1=Osmólska|first1=H.|date=1972|title=Preliminary note on a crocodilian from the Upper Cretaceous of Mongolia|journal=Palaeontologia Polonica|volume=27|pages=43–47|url=http://www.palaeontologia.pan.pl/Archive/1972-27_43-47_5-6.pdf}}{{cite journal|last1=Osmólska|first1=H.|last2=Hua|first2=S.|last3=Buffetaut|first3=E.|date=1997|title=Gobiosuchus kielanae (Protosuchia) from the Late Cretaceous of Mongolia: anatomy and relationships|journal=Acta Palaeontologica Polonica|volume=42|issue=2|pages=257–289|url=https://www.app.pan.pl/archive/published/app42/app42-257.pdf}}

| A gobisuchid.

| 200px

G.? parvus

| Udyn Sayr

| "Partial skull and skeleton."{{cite journal|last1=Efimov|first1=M. B.|date=193|title=Revision of fossil crocodiles of Mongolia|journal=Transactions of the Joint Soviet-Mongolian Paleontological Expedition|volume=24|pages=76–94|language=ru}}{{cite journal|last1=Efimov|first1=M. B.|date=1988|title=The fossil crocodiles and champsosaurides of Mongolia and USSR|journal=Transactions of the Joint Soviet-Mongolian Paleontological Expedition|volume=36|pages=5–105|language=ru}}

| A gobisuchid.

|

Shamosuchus

| S. djadochtaensis

| Bayn Dzak, Ukhaa Tolgod

| "Two skulls and partial skeleton."{{cite journal|last1=Pol|first1=D.|last2=Turner|first2=A. H.|last3=Norell|first3=M. A.|date=2009|title=Morphology of the late Cretaceous crocodylomorph Shamosuchus djadochtaensis and a discussion of neosuchian phylogeny as related to the origin of Eusuchia|journal=Bulletin of the American Museum of Natural History|number=324|pages=1–103|doi=10.1206/0003-0090-324.1.1|hdl=2246/5977|s2cid=83551788|hdl-access=free|oclc=421118402|url=https://www.researchgate.net/publication/230203413}}{{cite journal|last1=Turner|first1=A. H.|date=2015|title=A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia|journal=PLOS ONE|volume=10|issue=2|page=e0118116|bibcode=2015PLoSO..1018116T|doi=10.1371/journal.pone.0118116|doi-access=free|pmc=4340866|pmid=25714338}}

| A paralligatorid.

|

=Lizards=

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Adamisaurus

| A. magnidentatus

| Bayn Dzak, Tugriken Shireh, Ukhaa Tolgod

| "Skulls and skeletons from multiple specimens."{{cite journal|last1=Sulimski|first1=A.|date=1978|title=New data on the genus Adamisaurus Sulimski 1972 (Sauria) from the Upper Cretaceous of Mongolia|journal=Palaeontologica Polonica|volume=58|pages=43–56|url=http://www.palaeontologia.pan.pl/Archive/1978_38_43-56_12-14.pdf}}{{cite journal|last1=Keqin|first1=G.|last2=Norell|first2=M. A.|date=2000|title=Taxonomic composition and systematics of late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert|journal=American Museum Novitates|number=249|pages=1–118|hdl=2246/1596|hdl-access=free|doi=10.1206/0003-0090(2000)249<0001:TCASOL>2.0.CO;2|s2cid=129367764 |url=https://www.researchgate.net/publication/283992599}}

| A teiid. Also present in the Barun Goyot Formation and Bayan Mandahu Formation.

|

Aiolosaurus

| A. oriens

| Ukhaa Tolgod

| "Incomplete skull and partial skeleton."

| A varanoid.

|

Carusia

| C. intermedia

| Bayn Dzak, Ukhaa Tolgod

| "Skulls from multiple specimens."

| A carusiid. Also present in the Barun Goyot Formation and Bayan Mandahu Formation.

|

Cherminotus

| C. longifrons

| Tugriken Shireh, Ukhaa Tolgod

| "Skulls and partial skeleton."

| A varanoid. Also present in the Barun Goyot Formation.

|

Ctenomastax

| C. parva

| Zos

| "Incomplete skull."

| An iguanid. Also present in the Barun Goyot Formation.

|

Dzhadochtosaurus

| D. giganteus

| Tugriken Shireh

| "Partially complete skull."{{cite journal|last1=Alifanov|first1=V. R.|date=1993|title=Новые ящерицы семейства Macrocephalosauridae (Sauria) из верхнего мела Монголии и критические замечания о системе Teiidae (sensu Estes, 1983)|trans-title=New lizards of the family Macrocephalosauridae (Sauria) from the Upper Cretaceous of Mongolia, critical remarks on the systematics of the Teiidae (sensu Estes, 1983)|journal=Paleontological Journal|volume=27|issue=1|pages=57–74|language=ru|url=https://www.geokniga.org/bookfiles/geokniga-paleontologicaljournal1993-1.pdf}}

| A macrocephalosaur.

|

Eoxanta

| E. lacertifrons

| Ukhaa Tolgod

| "Incomplete skull."

| A scincomorph. Also present in the Barun Goyot Formation.

|

Estesia

| E. mongoliensis

| Bayn Dzak, Ukhaa Tolgod

| "Partial skulls and teeth."

| A monstersaur. Also present in the Barun Goyot Formation.

|

Flaviagama

| F. dzerzhinskii

| Tugriken Shireh

| "Skull and two vertebrae."{{cite journal|last1=Alifanov|first1=V. R.|date=1989|title=New priscagamas (Lacertilia) from the Upper Cretaceous of Mongolia and their systematic position in the Iguania|journal=Paleontological Journal|volume=4|pages=73–87|language=ru}}

| A priscagamid.

|

Globaura

| G. venusta

| Bayn Dzak, Ukhaa Tolgod

| "Partial skulls."

| A scincomorph. Also present in the Barun Goyot Formation.

|

Gobiderma

| G. pulchrum

| Udyn Sayr, Ukhaa Tolgod

| "Skulls and skin impressions."

| A monstersaur. Also present in the Barun Goyot Formation.

|

Gobinatus

| G. arenosus

| Ukhaa Tolgod

| "Partial skull."

| A teiid. Also present in the Barun Goyot Formation.

|

Hymenosaurus

| H. clarki

| Ukhaa Tolgod

| "Partial skull."

| A scincomorph.

|

Isodontosaurus

| I. gracilis

| Bayn Dzak, Tugriken Shireh, Ukhaa Tolgod, Zos

| "Numerous skulls and a partial skeleton."

| An iguanian. Also present in the Bayan Mandahu Formation.

|

Macrocephalosaurus

| Indeterminate

| Ukhaa Tolgod

| "Partial skull and skeleton."

| A teiid. Also present in the Barun Goyot Formation.

|

Mimeosaurus

| M. crassus

| Bayn Dzak, Ukhaa Tolgod, Zos Wash

| "Partially complete skulls."

| An acrodont. Also present in the Bayan Mandahu Formation.

|

Myrmecodaptria

| M. microphagosa

| Ukhaa Tolgod

| "Single skull."

| A gekkotan.

|

Ovoo

| O. gurval

| Little Ukhaa Tolgod

| "Partial skull."{{cite journal|last1=Norell|first1=M. A.|last2=Gao|first2=K.-Q.|last3=Conrad|first3=J. L.|date=2008|title=A new platynotan lizard (Diapsida: Squamata) from the Late Cretaceous Gobi Desert (Ömnögov), Mongolia|journal=American Museum Novitates|number=3605|pages=1–22|hdl=2246/5910|hdl-access=free|doi=10.1206/0003-0082(2008)3605[1:ANPLDS]2.0.CO;2|s2cid=86423607 |url=https://digitallibrary.amnh.org/bitstream/handle/2246/5910//v3/dspace/updateIngest/pdfs/N3605.pdf?sequence=1&isAllowed=y}}

| A varanid.

|

Parmeosaurus

| P. scutatus

| Ukhaa Tolgod

| "Articulated skull and skeleton."

| A scincomorph.

|

Phrynosomimus

| P. asper

| Ukhaa Tolgod

| "Two partial skulls."

| An acrodont. Also present in the Barun Goyot Formation.

|

Priscagama

| P. gobiensis

| Bayn Dzak, Ukhaa Tolgod

| "Incomplete skulls."

| An priscagamid. Also present in the Barun Goyot Formation.

|

Saichangurvel

| S. davidsoni

| Ukhaa Tolgod

| "Complete skull and skeleton in articulation."{{cite journal|last1=Conrad|first1=J. L.|last2=Norell|first2=M. A.|date=2007|title=A Complete Late Cretaceous Iguanian (Squamata, Reptilia) from the Gobi and Identification of a New Iguanian Clade|journal=American Museum Novitates|number=3584|pages=1–19|hdl=2246/5877|hdl-access=free|doi=10.1206/0003-0082(2007)3584[1:ACLCIS]2.0.CO;2|s2cid=83407564 |url=https://digitallibrary.amnh.org/bitstream/handle/2246/5877//v3/dspace/updateIngest/pdfs/N3584.pdf?sequence=1&isAllowed=y}}

| An iguanian.

|

Slavoia

| S. darevskii

| Ukhaa Tolgod

| "Skulls and skeleton."

| A scincomorph. Also present in the Barun Goyot Formation.

|

Telmasaurus

| T. grangeri

| Bayn Dzak

| "Partial skull and skeleton."{{cite journal|last1=Gilmore|first1=C. W.|year=1943|title=Fossil lizards of Mongolia|journal=Bulletin of the American Museum of Natural History|volume=81|pages=361–384}}

| A varanid.

|

Temujinia

| T. ellisoni

| Tugriken Shireh, Ukhaa Tolgod

| "Several partial skulls."

| An iguanid.

|

Tchingisaurus

| T. multivagus

| Ukhaa Tolgod

| "Partial skull."

| A teiid.

|

Unnamed scincomorph

| Indeterminate

| Ukhaa Tolgod

| "Partial skull."

| A scincomorph.

|

Varanoidea indet.

| Indeterminate

| Ukhaa Tolgod

| "Partial maxilla and vertebra."

| A varanoid.

|

Zapsosaurus

| Z. sceliphros

| Tugriken Shireh

| "Two partial skulls."

| An iguanid.

|

=Mammals=

==Multituberculates==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Bulganbaatar

| B. nemegtbaataroides

| Bayn Dzak, Ukhaa Tolgod

| "Partial skull, and other remains."Kielan-Jaworowska (1974). "Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia)". Palaeontologica Polonica 30: 23–44.{{cite journal|last1=Dashzeveg|first1=D.|last2=Novacek|first2=M. J.|last3=Norell|first3=M. A.|last4=Clark|first4=J. M.|last5=Chiappe|first5=L. M.|last6=Davidson|first6=A.|last7=McKenna|first7=M. C.|last8=Dingus|first8=L.|last9=Swisher|first9=C.|last10=Perle|first10=A.|date=1995|title=Extraordinary preservation in a new vertebrate assemblage from the Late Cretaceous of Mongolia|journal=Nature|volume=374|issue=6521|pages=446–449|bibcode=1995Natur.374..446D|doi=10.1038/374446a0|s2cid=4245878|url=https://www.researchgate.net/publication/238822891}}

| A djadochtatherioid.

|

Catopsbaatar

| C. catopsaloides

| Ukhaa Tolgod

| Not specified.

| A djadochtatheriid. Also present in the Barun Goyot Formation.

| 200px

Chulsanbaatar

| C. vulgaris

| Ukhaa Tolgod

| "Skull and partial skeleton."

| A djadochtatherioid. Also present in the Barun Goyot Formation.

|

Djadochtatherium

| D. matthewi

| Bayn Dzak, Tugriken Shireh

| "Partial skulls."{{cite journal|last1=Simpson|first1=G. G.|date=1925|title=A Mesozoic mammal skull from Mongolia|journal=American Museum Novitates|number=201|pages=1–12|hdl=2246/3185|hdl-access=free|url=http://digitallibrary.amnh.org/bitstream/handle/2246/3185//v2/dspace/ingest/pdfSource/nov/N0201.pdf?sequence=1&isAllowed=y}}{{cite journal|last1=Kielan-Jaworowska|first1=Z.|last2=Hurum|first2=J. H.|date=1997|title=Djadochtatheria: a new suborder of multituberculate mammals|journal=Acta Palaeontologica Polonica|volume=42|issue=2|pages=201–242|url=https://www.app.pan.pl/archive/published/app42/app42-201.pdf}}

| A djadochtatheriid.

|

Deltatheroides

| D. cretacicus

| Bayn Dzak

| "Partial skull."

| A djadochtatheriid.

|

Kamptobaatar

| K. kuczynskii

| Bayn Dzak, Ukhaa Tolgod

| "Partial skull, and other remains."{{cite journal|last1=Kielan-Jaworowska|first1=Z.|date=1969|title=New Upper Cretaceous multituberculate genera from Bayn Dzak, Gobi Desert|journal=Palaeontologica Polonica|volume=21|pages=35–49|url=http://www.palaeontologia.pan.pl/Archive/1969-21_35-52_10-17.pdf}}

| A sloanbaatarid.

|

Kryptobaatar

| K. dashzevegi

| Bayn Dzak, Tugriken Shireh, Ukhaa Tolgod

| "Skulls and skeleton remains from several specimens."{{cite journal|last1=Kielan-Jaworowska|first1=Z.|last2=Hurum|first2=J. H.|last3=Badamgarav|first3=D.|date=2003|title=An extended range of the multituberculate Kryptobaatar and distribution of mammals in the Upper Cretaceous of the Gobi Desert|journal=Acta Palaeontologica Polonica|volume=48|issue=2|pages=273–278|url=https://app.pan.pl/archive/published/app48/app48-273.pdf}}

| A djadochtatheriid. Gobibaatar and Tugrigbaatar are considered synonyms of this taxon.{{cite book|last1=Kielan-Jaworowska|first1=Z.|last2=Cifelli|first2=R. L.|last3=Luo|first3=Z.-X.|year=2004|title=Mammals from the Age of Dinosaurs: Origins, Evolution, and Structure|publisher=Columbia University Press|location=New York|doi=10.7312/kiel11918|jstor=10.7312/kiel11918|isbn=9780231119184}}

|

Mangasbaatar

| M. udanii

| Udyn Sayr

| "Skulls and partial skeleton from two specimens."{{cite journal|last1=Rougier|first1=G. W.|last2=Sheth|first2=A. S.|last3=Spurlin|first3=B. K.|last4=Bolortsetseg|first4=M.|last5=Novacek|first5=M. J.|date=2016|title=Craniodental anatomy of a new Late Cretaceous multituberculate mammal from Udan Sayr, Mongolia|journal=Palaeontologica Polonica|volume=67|issue=3|pages=197–248|doi=10.18297/ETD/1317|doi-access=free|url=http://www.palaeontologia.pan.pl/PP67/Rougier.pdf}}

| A djadochtatheriid.

|

Nemegtbaatar

| N. gobiensis

| Ukhaa Tolgod

| Not specified.

| A djadochtatherioid. Also present in the Barun Goyot Formation.

| File:Nemegtbaatar restoration.jpg

Sloanbaatar

| S. mirabilis

| Bayn Dzak, Ukhaa Tolgod

| "Complete skull, and other remains."

| A sloanbaatarid.

|

Tombaatar

| T. sabuli

| Ukhaa Tolgod

| "Partial skull."{{cite journal|last1=Rougier|first1=G. W.|last2=Novacek|first2=M. J.|last3=Dashzeveg|first3=D.|date=1997|title=A New Multituberculate from the Late Cretaceous Locality Ukhaa Tolgod, Mongolia. Considerations on Multituberculate Interrelationships|journal=American Museum Novitates|number=3191|pages=1–26|hdl=2246/3591|hdl-access=free|url=https://digitallibrary.amnh.org/bitstream/handle/2246/3591//v2/dspace/ingest/pdfSource/nov/N3191.pdf?sequence=1&isAllowed=y}}

| A djadochtatheriid.

|

==Therians==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Asiatherium

| A. reshetovi

| Udyn Sayr

| "Articulated skull and skeleton."{{cite journal|last1=Trofimov|first1=B. A|last2=Szalay|first2=F. S.|date=1994|title=New Cretaceous marsupial from Mongolia and the early radiation of Metatheria|journal=Proceedings of the National Academy of Sciences of the United States of America|volume=91|number=26|pages=12569−12573|doi=10.1073/pnas.91.26.12569|doi-access=free|jstor=2366383|pmc=45480|pmid=7809079|bibcode=1994PNAS...9112569T|url=https://www.pnas.org/content/pnas/91/26/12569.full.pdf}}

| A metatherian.

| 200px

Deltatheridium

| D. pretrituberculare

| Bayn Dzak, Ukhaa Tolgod

| "Partial skull and skeleton remains."{{cite journal|last1=Gregory|first1=W. K.|last2=Simpson|first2=G. G.|date=1926|title=Cretaceous Mammal Skulls from Mongolia|journal=American Museum Novitates|number=225|pages=698–699| doi=10.1038/118698a0|bibcode=1926Natur.118..698G|hdl=2246/3193|s2cid=38785853|hdl-access=free|url=http://digitallibrary.amnh.org/bitstream/handle/2246/3193//v2/dspace/ingest/pdfSource/nov/N0225.pdf?sequence=1&isAllowed=y}}{{cite journal|last1=Rougier|first1=G. W.|last2=Wible|first2=J. R.|last3=Novacek|first3=M. J.|date=1998|title=Implications of Deltatheridium specimens for early marsupial history|journal=Nature|volume=396|number=6710|pages=459–463|doi=10.1038/24856|pmid=9853752|bibcode=1998Natur.396..459R|s2cid=204996216|url=https://www.researchgate.net/publication/13433535}}

| A metatherian.

|

rowspan="2" | Hyotheridium

| H. dobsoni

| Bayn Dzak

| "Partial skull."

| A therian.

|

Indeterminate

| Ukhaa Tolgod

| Not specified.

| A therian.

|

Kennalestes

| K. gobiensis

| Bayn Dzak, Ukhaa Tolgod

| "Nearly complete skull, and other remains."{{cite journal|last1=Kielan-Jaworowska|first1=Z.|date=1968|title=Preliminary data on the Upper Cretaceous eutherian mammals from Bayn Dzak, Gobi Desert|journal=Palaeontologica Polonica|volume=19|pages=171–191|url=http://www.palaeontologia.pan.pl/Archive/1968-19_171-191_22-27.pdf}}

| An eutherian.

|

Maelestes

| M. gobiensis

| Ukhaa Tolgod

| "Partial skull with skeleton."{{cite journal|last1=Wible|first1=J. R.|last2=Rougier|first2=G. W.|last3=Novacek|first3=M. J.|last4=Ashek|first4=R. J.|date=2007|title=Cretaceous eutherians and Laurasian origin for placental mammals near the K/T boundary|journal=Nature|volume=447|issue=7147|pages=1003–1006|bibcode=2007Natur.447.1003W|doi=10.1038/nature05854|pmid=17581585|s2cid=4334424|url=https://www.researchgate.net/publication/6254868}}

| A cimolestid.

|

Ukhaatherium

| U. nessovi

| Ukhaa Tolgod

| "Partial to nearly complete skeletons from several specimens."{{cite journal|last1=Novacek|first1=M. J.|last2=Rougier|first2=G. W.|last3=Wible|first3=J. R.|last4=McKenna|first4=M. C.|last5=Dashzeveg|first5=D.|last6=Horovitz|first6=I.|date=1997|title=Epipubic bones in eutherian mammals from the Late Cretaceous of Mongolia|journal=Nature|volume=389|issue=6650|pages=483–486|bibcode=1997Natur.389..483N|doi=10.1038/39020|pmid=9333234|s2cid=205026882|url=https://www.researchgate.net/publication/13894490}}{{cite journal|last1=Horovitz|first1=I.|date=2000|title=The tarsus of Ukhaatherium nessovi (Eutheria, Mammalia) from the Late Cretaceous of Mongolia: an appraisal of the evolution of the ankle in basal therians|journal=Journal of Vertebrate Paleontology|volume=20|issue=3|pages=547–560|doi=10.1671/0272-4634(2000)020[0547:TTOUNE]2.0.CO;2|jstor=4524126|s2cid=86031996 }}{{cite journal|last1=Horovitz|first1=I.|date=2003|title=Postcranial skeleton of Ukhaatherium nessovi (Eutheria, Mammalia) from the Late Cretaceous of Mongolia|journal=Journal of Vertebrate Paleontology|volume=23|issue=4|pages=857–868|doi=10.1671/2399-10|jstor=4524387|bibcode=2003JVPal..23..857H |s2cid=85809847}}

| An eutherian.

|

Zalambdalestes

| Z. lechei

| Bayn Dzak, Tugriken Shireh

| "Skulls and skeletons from several specimens."{{cite journal|last1=Wible|first1=J. R.|last2=Novacek|first2=M. J.|last3=Rougier|first3=G. W.|date=2004|title=New data on the skull and dentition in the Mongolian Late Cretaceous eutherian mammal Zalambdalestes|journal=Bulletin of the American Museum of Natural History|volume=281|number=281|pages=1–144|doi=10.1206/0003-0090(2004)281<0001:NDOTSA>2.0.CO;2|hdl=2246/449|s2cid=83491778 |hdl-access=free|url=http://digitallibrary.amnh.org/dspace/bitstream/handle/2246/449/B281.pdf?sequence=1}}

| An eutherian.

| File:Zalambdalestes.jpg

=Pterosaurs=

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Azhdarchidae indet.

| Indeterminate

| Tugriken Shireh

| "Indeterminate bone inside the gut cavity of a Velociraptor."{{cite journal | last1 = Hone | first1 = D. | last2 = Tsuihiji | first2 = T. | last3 = Watabe | first3 = M. | last4 = Tsogtbaatr | first4 = K. | title = Pterosaurs as a food source for small dromaeosaurs | doi = 10.1016/j.palaeo.2012.02.021 | journal = Palaeogeography, Palaeoclimatology, Palaeoecology | volume = 331-332 | page = 27 | year = 2012 | bibcode = 2012PPP...331...27H }}

| An azhdarchid.

|

=Turtles=

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
rowspan="3" | Nanhsiungchelyidae indet.

| Indeterminate

| Abdrant Nuru

| "Three shell fragments."{{cite journal|last1=Danilov|first1=I. G.|last2=Syromyatnikova|first2=E.|date=2008|title=New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the nanhsiungchelyid record in Asia|journal=Proceedings of the Zoological Institute RAS|volume=312|number=1–2|pages=3–25|doi=10.31610/trudyzin/2008.312.1-2.3 |s2cid=130642652 |url=http://www.zin.ru/journals/trudyzin/doc/vol_312_1_2/tz_312_1_2_danilov.pdf}}

| A nanhsiungchelyid.

|

Indeterminate

| Bayn Dzak

| "Partial shells."

| A nanhsiungchelyid.

|

Indeterminate

| Udyn Sayr

| "Two shell fragments."

| A nanhsiungchelyid.

|

rowspan="2" | Zangerlia

| Z. dzamynchondi

| Zamyn Khondt

| "Partial shell."{{cite journal|last1=Sukhanov|first1=V. B.|last2=Narmandakh|first2=P.|date=2006|title=New taxa of Mesozoic turtles from Mongolia|journal=Fossil Turtle Research|volume=1|pages=119–127}}

| A nanhsiungchelyid.

|

Z. ukhaachelys

| Ukhaa Tolgod

| "Partial skull and skeleton."{{cite journal|last1=Joyce|first1=W. G.|last2=Norell|first2=M. A.|date=2005|title=Zangerlia ukhaachelys, New Species, a Nanhsiungchelyid Turtle from the Late Cretaceous of Ukhaa Tolgod, Mongolia|journal=American Museum Novitates|number=3481|pages=1–19|doi=10.1206/0003-0082(2005)481[0001:ZUNSAN]2.0.CO;2|hdl=2246/5672|s2cid=54948634 |hdl-access=free|url=https://digitallibrary.amnh.org/bitstream/handle/2246/5672/N3481.pdf?sequence=1&isAllowed=y}}

| A nanhsiungchelyid.

|

=Dinosaurs=

== Ankylosaurids ==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Minotaurasaurus

| M. ramachandrani

| Ukhaa Tolgod

| [Two] complete skulls, mandibles, and first cervical half-ring.{{cite journal|last1=Alicea|first1=J.|last2=Loewen|first2=M.|date=2013|title=New Minotaurasaurus material from the Djodokta Formation establishes new taxonomic and stratigraphic criteria for the taxon|journal=Journal of Vertebrate Paleontology|volume=Program and Abstracts|pages=76|url=https://vertpaleo.org/wp-content/uploads/2021/03/SVP-2013-merged-book-10-15-2013.pdf}}{{cite journal|last1=Penkalski|first1=P.|last2=Tumanova|first2=T.|date=2017|title=The cranial morphology and taxonomic status of Tarchia (Dinosauria: Ankylosauridae) from the Upper Cretaceous of Mongolia|journal=Cretaceous Research|volume=70|pages= 117–12|doi=10.1016/j.cretres.2016.10.004|bibcode=2017CrRes..70..117P }}

| An ankylosaurid previously thought to be a junior synonym of Tarchia, but is now considered to be a valid and distinct taxon.

| 200px

Pinacosaurus

| P. grangeri

| Bayn Dzak, Ukhaa Tolgod

| [Three] skulls, mandibles, predentary, cervical vertebrae, dorsal vertebrae, caudal vertebrae, ribs, scapula, coracoids, humerus, radius, ulna, ilium, femora, tibia, fibula, pelvis, manus, tail club handles, cervical half-rings, osteoderms, and a nearly complete skeleton lacking a skull.{{cite journal|last=Gilmore|first=C. W.|title=Two new dinosaurian reptiles from Mongolia with notes on some fragmentary specimens|journal=American Museum Novitates|issue=679|pages=1–20|date=1933|hdl=2246/2076}}{{cite journal|last1=Hill|first1=R. V.|last2=Witmer|first2=L. W.|last3=Norell|first3=M. A.|date=2003|title=A New Specimen of Pinacosaurus grangeri (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia: Ontogeny and Phylogeny of Ankylosaurs|journal=American Museum Novitates|issue=3395|pages=1–29|doi=10.1206/0003-0082(2003)395<0001:ANSOPG>2.0.CO;2|hdl=2246/2821|s2cid=3253690 |hdl-access=free|url=https://digitallibrary.amnh.org/bitstream/handle/2246/2821//v2/dspace/ingest/pdfSource/nov/N3395.pdf?sequence=1&isAllowed=y}}{{cite journal|last1=Currie|first1=P. J.|last2=Badamgarav|first2=D.|last3=Koppelhus|first3=E. B.|last4=Sissons|first4=R.|last5=Vickaryous|first5=M. K.|date=2011|title=Hands, feet and behaviour in Pinacosaurus (Dinosauria: Ankylosauridae)|journal=Acta Palaeontologica Polonica|volume=56|issue=3|pages=489–504|doi=10.4202/app.2010.0055|s2cid=129291148|url=http://www.app.pan.pl/archive/published/app56/app20100055.pdf|doi-access=free}}

| An ankylosaurid also known from the Alagteeg Formation and Bayan Mandahu Formation.

| 200px

Ankylosauridae indet.

| Indeterminate

| Zamyn Khondt

| Partially complete postcranial skeleton with in situ osteoderms.{{cite journal|last1=Arbour|first1=V. M.|last2=Currie|first2=P. J.|date=2013|title=The taxonomic identity of a nearly complete ankylosaurid dinosaur skeleton from the Gobi Desert of Mongolia|journal=Cretaceous Research|volume=46|pages=24–30|doi=10.1016/j.cretres.2013.08.008|bibcode=2013CrRes..46...24A }}

| Previously referred to Saichania, but is now referred to as Ankylosauridae indet., or cf. Pinacosaurus.

| 200px

== Marginocephalians ==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
style="background:#e6e6e6;" | Bainoceratops

| style="background:#e6e6e6;" | B. efremovi

| style="background:#e6e6e6;" | Bayn Dzak

| style="background:#e6e6e6;" | "Partial vertebrae."{{cite journal|last1=Tereshchenko|first1=V.|last2=Alifanov|first2=V. R.|date=2003|title=Bainoceratops efremovi, a New Protoceratopid Dinosaur (Protoceratopidae, Neoceratopsia) from the Bain-Dzak Locality (South Mongolia)|journal=Paleontological Journal|volume=37|issue=3|pages=293–302|url=https://www.researchgate.net/publication/288557787}}

| style="background:#e6e6e6;" | A protoceratopsid. May be synonymous with Protoceratops.{{cite journal|last1=Makovicky|first1=P. J.|last2=Norell|first2=M. A.|date=2006|title=Yamaceratops dorngobiensis, a New Primitive Ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia|journal=American Museum Novitates|number=3530|pages=1–42|doi=10.1206/0003-0082(2006)3530[1:YDANPC]2.0.CO;2|hdl=2246/5808|hdl-access=free|url=http://digitallibrary.amnh.org/dspace/bitstream/2246/5808/1/N3530.pdf}}

| style="background:#e6e6e6;" |

Bagaceratops

| Indeterminate

| Udyn Sayr

| "Skull with partial skeleton."{{cite journal|last1=Czepiński|first1=Ł.|date=2020|title=New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata|journal=Acta Palaeontologica Polonica|volume=65|issue=3|pages=481–497|doi=10.4202/app.00701.2019|doi-access=free|url=http://www.app.pan.pl/archive/published/app65/app007012019.pdf}}

| A protoceratopsid. Indeterminate between Bagaceratops and Protoceratops.

| 200px

Goyocephale

| G. lattimorei

| Boro Khovil

| "Partial skull and skeleton."{{cite journal|last1=Perle|first1=A.|last2=Maryańska|first2=T.|last3=Osmólska|first3=H.|date=1982|title=Goyocephale lattimorei gen. et sp. n., a new flat-headed pachycephalosaur (Ornithischia, Dinosauria) from the Upper Cretaceous of Mongolia|journal=Acta Palaeontologica Polonica|volume=27|issue=1–4|pages=115–127|url=https://www.app.pan.pl/archive/published/app27/app27-115.pdf}}

| A pachycephalosaurid. Locality sediments may belong to this formation.{{cite journal|last1=Sullivan|first1=R. M.|date=2006|title=A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)|journal=New Mexico Museum of Natural History and Science Bulletin|issue=35|pages=347–365|url=https://www.researchgate.net/publication/240625747}}

| 200px

rowspan="2" | Protoceratops

| P. andrewsi

| Bayn Dzak, Tugriken Shireh, Udyn Sayr, Zamyn Khondt

| "Multiple partial to complete specimens."{{cite journal|last1=Brown|first1=B.|last2=Schlaikjer|first2=E. M.|date=1940|title=The Structure and Relationships of Protoceratops|journal=Annals of the New York Academy of Sciences|volume=40|number=3|pages=133–266|bibcode=1940NYASA..40..133B|doi=10.1111/j.2164-0947.1940.tb00068.x|oclc=1673730|url=https://drive.google.com/file/d/1EM9A8LG5eqbQ5_Ho16QJx0_MebAqLjLQ/view}}{{cite journal|last1=Fastovsky|first1=D. E.|last2=Weishampel|first2=D. B.|last3=Watabe|first3=M.|last4=Barsbold|first4=R.|last5=Tsogtbaatar|first5=K.|last6=Narmandakh|first6=P.|date=2011|title=A nest of Protoceratops andrewsi (Dinosauria, Ornithischia)|journal=Journal of Paleontology|volume=85|issue=6|page=1035−1041|doi=10.1666/11-008.1|jstor=41409110|s2cid=129085129|url=https://www.researchgate.net/publication/261971168}}{{cite journal|last1=Handa|first1=N.|last2=Watabe|first2=M.|last3=Tsogtbaatar|first3=K.|date=2012|title=New Specimens of Protoceratops (Dinosauria: Neoceratopsia) from the Upper Cretaceous in Udyn Sayr, Southern Gobi Area, Mongolia|journal=Paleontological Research|volume=16|issue=3|pages=179–198|doi=10.2517/1342-8144-16.3.179|s2cid=130903035}}{{cite journal|last1=Hone|first1=D. W. E.|last2=Farke|first2=A. A.|last3=Watabe|first3=M.|last4=Shigeru|first4=S.|last5=Tsogtbaatar|first5=K.|date=2014|title=A New Mass Mortality of Juvenile Protoceratops and Size-Segregated Aggregation Behaviour in Juvenile Non-Avian Dinosaurs|journal=PLOS ONE|volume=9|issue=11|pages=e113306|doi=10.1371/journal.pone.0113306|doi-access=free|pmc=4245121|pmid=25426957|bibcode=2014PLoSO...9k3306H}}

| A protoceratopsid.

| 200px

P. hellenikorhinus

| Bor Tolgoi, Udyn Sayr

| "Partial cranial remains."{{cite journal|last1=Chiba|first1=K.|last2=Ryan|first2=M. J.|last3=Saneyoshi|first3=M.|last4=Konishi|first4=S.|last5=Yamamoto|first5=Y.|last6=Mainbayar|first6=B.|last7=Tsogtbaatar|first7=K.|date=2020|title=Taxonomic re-evaluation of Protoceratops (Dinosauria: Ceratopsia) specimens from Udyn Sayr, Mongolia|journal=Journal of Vertebrate Paleontology|volume=Program and Abstracts|url=https://vertpaleo.org/wp-content/uploads/2021/03/SVP_2020_Program-Abstracts-Volume-FINAL-for-Publishing-1.27.2021.pdf}}

| A protoceratopsid.

| 200px

Protoceratopsidae indet.

| Indeterminate

| Ukhaa Tolgod

| "Numerous skulls and remains."{{cite journal|last1=Czepiński|first1=Ł.|date=2019|title=Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert|journal=Historical Biology|volume=32|issue=10|pages=1394–1421|doi=10.1080/08912963.2019.1593404|s2cid=132780322|url=http://dinosaurmailinglist.cmnh.org/2019Apr/pdfzmfpMk1aO4.pdf|access-date=2021-07-03|archive-date=2021-07-08|archive-url=https://web.archive.org/web/20210708144840/http://dinosaurmailinglist.cmnh.org/2019Apr/pdfzmfpMk1aO4.pdf|url-status=dead}}{{cite journal|last1=Prieto-Márquez|first1=A.|last2=Garcia-Porta|first2=J.|last3=Joshi|first3=S. H.|last4=Norell|first4=M. A.|last5=Makovicky|first5=P. J.|date=2020|title=Modularity and heterochrony in the evolution of the ceratopsian dinosaur frill|journal=Ecology and Evolution|volume=10|issue=13|pages=6288–6309|doi=10.1002/ece3.6361|doi-access=free|pmc=7381594|pmid=32724514|bibcode=2020EcoEv..10.6288P }}

| A protoceratopsid.

| 200px

Udanoceratops

| U. tschizhovi

| Udyn Sayr

| "Skull and fragmented skeleton elements."{{cite journal|last1=Kurzanov|first1=S. M.|date=1992|title=A giant protoceratopsid from the Upper Cretaceous of Mongolia|journal=Paleontological Journal|pages=81–93|language=ru}}

| A giant leptoceratopsid.

| 200px

==Ornithopods==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Hadrosauroidea indet.

| Indeterminate

| Tugriken Shireh

| "Fragmented remains from juveniles."{{cite journal|last1=Barsbold|first1=R.|last2=Perle|first2=A.|date=1983|title=On taphonomy of joint burial of juvenile dinosaurs and some aspects of their ecology|journal=Transactions of the Joint Soviet-Mongolian Paleontological Expedition|volume=24|pages=121–125|language=ru}}

| A hadrosauroid.

|

style="background:#f3e9f3;" | Plesiohadros

| style="background:#f3e9f3;" | P. djadokhtaensis

| style="background:#f3e9f3;" | Alag Teeg

| style="background:#f3e9f3;" | "Skull and partial body elements."{{cite book|last1=Tsogtbaatar|first1=K.|last2=Weishampel|first2=D.|last3=Evans|first3=D. C.|last4=Watabe|first4=M.|year=2014|chapter=A New Hadrosauroid (Plesiohadros djadokhtaensis) from the Late Cretaceous Djadokhtan Fauna of Southern Mongolia|chapter-url=https://www.researchgate.net/publication/290319937|editor-last1=Eberth|editor-first1=D. A.|editor-last2=Evans|editor-first2=D. C.|title=Hadrosaurs|pages=108–135|publisher=Indiana University Press|isbn=978-0-253-01385-9}}

| style="background:#f3e9f3;" | A hadrosauroid. Actually hails from the Alagteeg Formation.{{cite journal|last1=Hasegawa|first1=H.|last2=Tada|first2=R.|last3=Ichinnorov|first3=N.|last4=Minjin|first4=C.|date=2009|title=Lithostratigraphy and depositional environments of the Upper Cretaceous Djadokhta Formation, Ulan Nuur basin, southern Mongolia, and its paleoclimatic implication|journal=Journal of Asian Earth Sciences|volume=35|issue=1|pages=13–26|doi=10.1016/j.jseaes.2008.11.010|bibcode=2009JAESc..35...13H|url=https://linkinghub.elsevier.com/retrieve/pii/S1367912008001880}}

| style="background:#f3e9f3;" |

==Sauropods==

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Sauropoda indet.

| Indeterminate

| Not specified.{{cite journal|last1=Jerzykiewicz|first1=T.|last2=Russell|first2=D. A.|date=1991|title=Late Mesozoic stratigraphy and vertebrates of the Gobi Basin|journal=Cretaceous Research|volume=12|number=4|pages=345–377|doi=10.1016/0195-6671(91)90015-5|bibcode=1991CrRes..12..345J }}

| Not specified.

| A sauropod.

== Theropods ==

=== Alvarezsaurs ===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Alvarezsauridae indet.

| Indeterminate

| Tugriken Shireh

| "Partial skull, braincase, and skeleton elements of two specimens."Suzuki, Chiappe, Dyke, Watabe, Barsbold and Tsogtbaatar, 2002. A new specimen of Shuvuuia deserti Chiappe et al., 1998, from the Mongolian Late Cretaceous with a discussion of the relationships of alvarezsaurids to other theropod dinosaurs. Contributions in Science. 494, 1–18.Longrich & Currie (2009). Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae. Cretaceous Research. 30(1), 239–252.

| An alvarezsaurid also known as the Tugriken Shireh alvarezsaur. Uncertainly referred as Parvicursor sp.

| rowspan = 100| File:Kol ghuva.jpg]] File:Shuvuuia.jpg]]

Kol

| K. ghuva

| Ukhaa Tolgod

| "A well-preserved right foot."{{cite journal |first1=A.H. |last1=Turner |first2=S.J. |last2=Nesbitt |first3=M.A. |last3=Norell |title=A Large Alvarezsaurid from the Cretaceous of Mongolia |journal=American Museum Novitates |issue=3648 |pages=1–14 |year=2009 |doi=10.1206/639.1|hdl=2246/5967 |s2cid=59459861 |url=https://zenodo.org/record/4712967 }}

| A large alvarezsaurid. Its classification has been criticized.{{cite journal|author=Federico L. Agnolin | author2=Jaime E. Powell | author3=Fernando E. Novas | author4=Martin Kundrát |date=2012|title=New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs| journal=Cretaceous Research | volume=35 | pages=33–56 | doi=10.1016/j.cretres.2011.11.014| bibcode=2012CrRes..35...33A |url=https://www.researchgate.net/publication/257047862}}

Shuvuuia

| S. deserti

| Ukhaa Tolgod

| "Multiple specimens with skull and skeletons."Chiappe, L.M., Norell, M. A., and Clark, J. M. (1998). "The skull of a relative of the stem-group bird Mononykus." Nature, 392 (6673): 275–278.Dufeau, 2003. The cranial anatomy of the theropod dinosaur Shuvuuia deserti (Coelurosauria: Alvarezsauridae), and its bearing upon coelurosaurian phylogeny. Masters Thesis, The University of Texas at Austin. 275 pp.Pei, 2015. New paravian fossils from the Mesozoic of east Asia and their bearing on the phylogeny of the Coelurosauria. PhD thesis, Columbia University. 545 pp.Saitta, E., Fletcher, I., Martin, P.G., Pittman, M., Kaye, T., True, L., Norell, M., Abbott, G., Summons, R., Penkman, K., & Vinther, J. (2018). Preservation of feather fibers from the Late Cretaceous dinosaur Shuvuuia deserti raises concern about immunohistochemical analyses on fossils. Organic Geochemistry. 125: 142–151.

| An alvarezsaurid.

rowspan="2" | Undescribed Alvarezsauridae

| Indeterminate

| Bayn Dzak

| "Partial pelvic girdle and hindlimb."Norell, Chiappe and Clark, (1993). New limb on the avian family tree. Natural History. September, 38–43.

| An alvarezsaurid.

Indeterminate

| Gilvent Wash

| Not given.

| An alvarezsaurid.

=== Birds ===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Apsaravis

| A. ukhaana

| Ukhaa Tolgod

| "Partial postcranial skeleton."Clarke, Julia A., Norell, Mark A. (2002). "The morphology and phylogenetic position of Apsaravis ukhaana from the Late Cretaceous of Mongolia". American Museum Novitates, No. 3387, American Museum of Natural History, New York, NY.

| A basal ornithurine bird.

|

Elsornis

| E. keni

| Tugriken Shireh

| "Partial articulated skeleton lacking the skull."{{cite journal|last1=Chiappe|first1=L. M.|last2=Suzuki|first2=S.|last3=Dyke|first3=G. J.|last4=Watabe|first4=M.|last5=Tsogtbaatar|first5=K.|last6=Barsbold|first6=R.|date=2007|title=A new Enantiornithine bird from the Late Cretaceous of the Gobi desert|journal=Journal of Systematic Palaeontology|volume=5|issue=2|pages=193–208|doi=10.1017/S1477201906001969|bibcode=2007JSPal...5..193C |s2cid=85391743}}

| An enantiornithe.

|

Gobipteryx

| G. minuta

| Ukhaa Tolgod

| "Partial skull."Chiappe, L, M.; Norell M. A.; Clark, J. M. (2001). A new skull of Gobipteryx minuta (Aves: Enantiornithes) from the Cretaceous of the Gobi Desert. American Museum Novitates. 3346, 1–15.

| An enantiornithine. Also present in the Barun Goyot Formation.

|

rowspan="3"; style="background:#e3f5ff;" | Protoceratopsidovum

| style="background:#e3f5ff;" | P. fluxuosum

| style="background:#e3f5ff;" | Bayn Dzak

| style="background:#e3f5ff;" | "Partial eggs."{{cite journal|last1=Mikhailov|first1=K. E.|date=1994|title=Theropod and protoceratopsian dinosaur eggs from the Cretaceous of Mongolia and Kazakhstan|journal=Paleontological Journal|volume=28|issue=2|pages=101–120|url=https://www.researchgate.net/publication/285873142}}

| style="background:#e3f5ff;" | Eggs probably laid by a bird.

| style="background:#e3f5ff;" |

style="background:#e3f5ff;" | P. minimum

| style="background:#e3f5ff;" | Baga Tariach, Tugriken Shireh

| style="background:#e3f5ff;" | "Clutch of four eggs and one pole of egg."

| style="background:#e3f5ff;" | Eggs probably laid by a bird.

| style="background:#e3f5ff;" |

style="background:#e3f5ff;" | P. sincerum

| style="background:#e3f5ff;" | Bayn Dzak, Tugriken Shireh

| style="background:#e3f5ff;" | "Multiple eggs and shells."

| style="background:#e3f5ff;" | Eggs probably laid by a bird.

| style="background:#e3f5ff;" |

style="background:#e3f5ff;" | Styloolithus

| style="background:#e3f5ff;" | S. sabathi

| style="background:#e3f5ff;" | Bayn Dzak

| style="background:#e3f5ff;" | "Partial to complete eggs."{{cite journal | last1 = Varricchio | first1 = D.J. | last2 = Barta | first2 = D.E. | year = 2015 | title = Revisiting Sabath's "Larger Avian Eggs" from the Gobi Cretaceous | url = https://www.app.pan.pl/article/item/app000852014.html | journal = Acta Palaeontologica Polonica | volume = 60 | issue = 1| pages = 11–25 }}

| style="background:#e3f5ff;" | Eggs probably laid by a bird.

| style="background:#e3f5ff;" |

===Dromaeosaurs===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Halszkaraptor

| H. escuilliei

| Ukhaa Tolgod

| "Partial skeleton with complete skull."{{cite journal|last1=Cau|first1=A.|last2=Beyrand|first2=V.|last3=Voeten|first3=D. F. A. E.|last4=Fernandez|first4=V.|last5=Tafforeau|first5=P.|last6=Stein|first6=K.|last7=Barsbold|first7=R.|last8=Tsogtbaatar|first8=K.|last9=Currie|first9=P. J.|last10=Godefroit|first10=P.|date=2017|title=Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs|journal=Nature|volume=552|issue=7685|pages=395–399|bibcode=2017Natur.552..395C|doi=10.1038/nature24679|pmid=29211712|s2cid=4471941|url=https://www.scienceinschool.org/sites/default/files/teaserMaterial/issue43_nature_cau2017.pdf|access-date=2021-07-03|archive-date=2021-07-08|archive-url=https://web.archive.org/web/20210708144831/https://www.scienceinschool.org/sites/default/files/teaserMaterial/issue43_nature_cau2017.pdf|url-status=dead}}

| A halszkaraptorine.

| 200px

Mahakala

| M. omnogovae

| Tugriken Shireh

| "Fragmented skull and skeleton."{{cite journal|last1=Turner|first1=A. H.|last2=Pol|first2=D.|last3=Clarke|first3=J. A.|last4=Erickson|first4=G. M.|last5=Norell|first5=M. A.|date=2007|title=A Basal Dromaeosaurid and Size Evolution Preceding Avian Flight|journal=Science|volume=317|issue=5843|pages=1378–1381|bibcode=2007Sci...317.1378T|doi=10.1126/science.1144066|doi-access=free|pmid=17823350}}

| A halszkaraptorine.

| 200px

Tsaagan

| T. mangas

| Ukhaa Tolgod

| "Skull and partial skeleton."{{cite journal | last1 = Norell | first1 = M.A. | last2 = Clark | first2 = J.M. | last3 = Turner | first3 = A.H. | last4 = Makovicky | first4 = P.J. | last5 = Barsbold | first5 = R. | last6 = Rowe | first6 = T. | year = 2006 | title = A new dromaeosaurid theropod from Ukhaa Tolgod (Omnogov, Mongolia) | journal = American Museum Novitates | issue = 3545 | pages = 1–51 | doi = 10.1206/0003-0082(2006)3545[1:ANDTFU]2.0.CO;2 | issn = 0003-0082 | hdl = 2246/5823 | url = https://www.biodiversitylibrary.org/bibliography/169121 }}

| A dromaeosaurid

| 200x200px

Velociraptor

| V. mongoliensis

| Bayn Dzak, Chimney Buttes, Gilvent Wash, Tugriken Shireh, Udyn Sayr, Ukhaa Tolgod

| "Multiple partial to complete specimens."Norell and Makovicky, 1997. Important features of the dromaeosaur skeleton: Information from a new specimen. American Museum Novitates. 3215, 28 pp.Barsbold and Osmólska, 1999. The skull of Velociraptor (Theropoda) from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica. 44(2), 189–219.Norell and Makovicky, 1999. Important features of the dromaeosaurid skeleton II: Information from newly collected specimens of Velociraptor mongoliensis. American Museum Novitates. 3282, 45 pp.Watabe and Tsogtbaatar, 2004. Report on the Japan – Mongolia Joint Paleontological Expedition to the Gobi desert, 2000. Hayashibara Museum of Natural Sciences Research Bulletin. 2, 45–67.Turner, Makovicky and Norell, 2007. Feather quill knobs in the dinosaur Velociraptor. Science. 317, 1721.David Hone; Jonah Choiniere; Corwin Sullivan; Xing Xu; Michael Pittman; Qingwei Tan (2010). New evidence for a trophic relationship between the dinosaurs Velociraptor and Protoceratops, 291(3–4), 0–492. doi:10.1016/j.palaeo.2010.03.028.

| A dromaeosaurid.

| 200px

rowspan="2" | Undescribed Dromaeosauridae

| Indeterminate

| Abdrant Nuru

| "Claw."Watabe and Suzuki, (2000). Report on the Japan – Mongolia Joint Paleontological Expedition to the Gobi desert, 1996. Hayashibara Museum of Natural Sciences Research Bulletin. 1, 58–68.

| A dromaeosaurid.

|

Indeterminate

| Zos Wash

| "Frontal region."

| A dromaeosaurid. Differs from Tsaagan.

|

===Non-maniraptoran theropods===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Aepyornithomimus

| A. tugrikinensis

| Tugriken Shireh

| "Nearly complete foot."{{cite journal|last1=Chinzorig|first1=T.|last2=Kobayashi|first2=Y.|last3=Tsogtbaatar|first3=K.|last4=Currie|first4=P. J.|last5=Watabe|first5=M.|last6=Barsbold|first6=R.|date=2017|title=First Ornithomimid (Theropoda, Ornithomimosauria) from the Upper Cretaceous Djadokhta Formation of Tögrögiin Shiree, Mongolia|journal=Scientific Reports|volume=7|issue=5835|page=5835|bibcode=2017NatSR...7.5835C|doi=10.1038/s41598-017-05272-6|doi-access=free|pmc=5517598|pmid=28724887}}

| An ornithomimid.

| 200px

Ornithomimosauria indet.

| Indeterminate

| Ukhaa Tolgod

| "Partial braincase, jaw tips, ribs, and vertebral fragments".{{cite journal | last1 = Makovicky | first1 = P. J. | last2 = Norell | first2 = Mark A. | year = 1998 | title=A partial ornithomimid briancase from Ukhaa Tolgod (Upper Cretaceous, Mongolia) | journal = American Museum Novitates | issue = 3247 | pages = 1–16}}{{cite journal | last1 = Ksepka | first1 = Daniel T. | last2 = Norell | first2 = Mark A. | year = 2004 | title=Ornithomimosauria cranial material from Ukhaa Tolgod (Omnogov, Mongolia) | journal = American Museum Novitates | issue = 3448 | pages = 1–4 | issn = 0003-0082 | doi=10.1206/0003-0082(2004)448<0001:ocmfut>2.0.co;2| hdl = 2246/2813 | s2cid = 55019144 | url = https://www.biodiversitylibrary.org/item/292382 }}

| An ornithomimid.

|

rowspan="4" |

Tyrannosauridae indet.

| Indeterminate

| Bayn Dzak

| Not specified.{{cite journal|last1=Longrich|first1=N. R.|last2=Currie|first2=P. J.|last3=Dong|first3=Z.|date=2010|title=A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia|journal=Palaeontology|volume=53|issue=5|pages=945–960|doi=10.1111/j.1475-4983.2010.00968.x|doi-access=free|bibcode=2010Palgy..53..945L }}

| A tyrannosaurid.

|

Indeterminate

| Khongil

| "Supraorbital, vertebra, rib, femur and metatarsals."{{cite journal|last1=Watabe|first1=M.|last2=Tsogtbaatar|first2=K.|date=2004|title=Report on the Japan – Mongolia Joint Paleontological Expedition to the Gobi desert, 2000|journal=Hayashibara Museum of Natural Sciences Research Bulletin|volume=2|pages=45–67}}

| A tyrannosaurid.

|

Indeterminate

| Not specified.

| "Partial right ilium."{{cite journal|last1=Gilmore|first1=C. W.|date=1933|title=Two new dinosaurian reptiles from Mongolia with notes on some fragmentary specimens|journal=American Museum Novitates|issue=679|pages=1–20|hdl=2246/2076|hdl-access=free}}

| A tyrannosaurid.

|

Indeterminate

| Not specified.

| "Teeth."{{cite journal|last1=Jerzykiewicz|first1=T.|last2=Currie|first2=P. J.|last3=Eberth|first3=D. A.|last4=Johnston|first4=P. A.|last5=Koster|first5=E. H.|last6=Zheng|first6=J.-J.|date=1993|title=Djadokhta Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi|journal=Canadian Journal of Earth Sciences|volume=30|issue=10|pages=2180–2195|bibcode=1993CaJES..30.2180J|doi=10.1139/e93-190}}

| A tyrannosaurid.

|

===Oviraptorosaurs===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Avimimus

| A. portentonsus

| Udyn Sayr

| "Partial skull and skeleton."{{cite journal|last1=Kurzanov|first1=S. M.|date=1981|title=An unusual theropod from the Upper Cretaceous of Mongolia|journal=Transactions, Joint Soviet–Mongolian Palaeontological Expedition|volume=15|pages=39–49|language=ru}}{{cite journal|last1=Funston|first1=G. F.|last2=Mendonca|first2=S. E.|last3=Currie|first3=P. J.|last4=Barsbold|first4=R.|last5=Barsbold|first5=R.|date=2018|title=Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin|journal=Palaeogeography, Palaeoclimatology, Palaeoecology|volume=494|pages=101–120|bibcode=2018PPP...494..101F|doi=10.1016/j.palaeo.2017.10.023|url=https://www.researchgate.net/publication/320986075}}

| An avimimid. Its locality origin has been disputed and may hail from the Nemegt Formation.{{cite journal|last1=Watabe|first1=M.|last2=Suzuki|first2=S.|last3=Tsogtbaatar|first3=K.|date=2006|title=Geological and geographical distribution of bird-like theropod, Avimimus in Mongolia|journal =Journal of Vertebrate Paleontology|volume=26|issue=supp. 003|pages=136A−137A|doi=10.1080/02724634.2006.10010069|s2cid=220413406}}

| 200px

Citipati

| C. osmolskae

| Ukhaa Tolgod

| "Multiple specimens with partial to nearly complete skeletons, an embryo, eggs and nesting individuals."{{cite journal|last1=Norell|first1=M. A.|last2=Clark|first2=J. M.|last3=Dashzeveg|first3=D.|last4=Barsbold|first4=R.|last5=Chiappe|first5=L. M.|last6=Davidson|first6=A. R.|last7=McKenna|first7=M. C.|last8=Altangerel|first8=P.|last9=Novacek|first9=M. J.|date=1994|title=A theropod dinosaur embryo and the affinities of the Flaming Cliffs Dinosaur eggs|journal=Science|volume=266|issue=5186|pages=779–782|bibcode=1994Sci...266..779N|doi=10.1126/science.266.5186.779|pmid=17730398|jstor=2885545|s2cid=22333224|url=https://www.researchgate.net/publication/6110932}}{{cite journal|last1=Clark|first1=J. M.|last2=Norell|first2=M. A.|last3=Barsbold|first3=R.|date=2001|title=Two new oviraptorids (Theropoda: Oviraptorosauria) from the Late Cretaceous Djadokta Formation, Ukhaa Tolgod|journal=Journal of Vertebrate Paleontology|volume=21|issue=2|pages=209–213|doi=10.1671/0272-4634(2001)021[0209:TNOTOU]2.0.CO;2|jstor=20061948|s2cid=86076568 |url=https://www.researchgate.net/publication/232685671}}{{cite journal|last1=Clark|first1=J. M.|last2=Norell|first2=M. A.|last3=Rowe|first3=T.|date=2002|title=Cranial Anatomy of Citipati osmolskae (Theropoda, Oviraptorosauria), and a Reinterpretation of the Holotype of Oviraptor philoceratops|journal=American Museum Novitates|number=3364|pages=1–24|doi=10.1206/0003-0082(2002)364<0001:CAOCOT>2.0.CO;2|hdl=2246/2853|s2cid=52247789 |hdl-access=free|url=http://digitallibrary.amnh.org/bitstream/handle/2246/2853/v2/dspace/ingest/pdfSource/nov/N3364.pdf?sequence=1}}{{cite journal|last1=Norell|first1=M. A.|last2=Balanoff|first2=A. M.|last3=Barta|first3=D. E.|last4=Erickson|first4=G. M.|date=2018|title=A second specimen of Citipati osmolskae associated with a nest of eggs from Ukhaa Tolgod, Omnogov Aimag, Mongolia|journal=American Museum Novitates|number=3899|pages=1–44|doi=10.1206/3899.1|hdl=2246/6858|s2cid=53057001|hdl-access=free|url=https://www.researchgate.net/publication/324784729}}

| An oviraptorid.

| 200px

rowspan="2"; style="background:#e3f5ff;" | Elongatoolithus

| style="background:#e3f5ff;" | E. frustrabilis

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

style="background:#e3f5ff;" | E. subtitectorius

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

| style="background:#e3f5ff;" |

Khaan

| K. mckennai

| Ukhaa Tolgod

| "Several specimens with partial to complete skeletons and skulls."{{cite journal|last1=Balanoff|first1=A. M.|last2=Norell|first2=M. A.|date=2012|title=Osteology of Khaan mckennai (Oviraptorosauria, Theropoda)|journal=Bulletin of the American Museum of Natural History|volume=372|number=372|pages=1–77|doi=10.1206/803.1|hdl=2246/6413|s2cid=86067306|hdl-access=free|oclc=884738416|url=https://www.researchgate.net/publication/270678811}}{{cite journal|last1=Scott Persons IV|first1=W.|last2=Funston|first2=G. F.|last3=Currie|first3=P. J.|last4=Norell|first4=M. A.|date=2015|title=A possible instance of sexual dimorphism in the tails of two oviraptorosaur dinosaurs|journal=Scientific Reports|volume=5|number=9472|pages=16247|bibcode=2015NatSR...5E9472P|doi=10.1038/srep09472|doi-access=free|pmc=4379468|pmid=25824625}}

| An oviraptorid.

| 200px

style="background:#c9ecff;" | Macroolithus

|style="background:#e3f5ff;" | M. mutabilis

|style="background:#e3f5ff;" |

|style="background:#e3f5ff;" | "Eggs."

|style="background:#e3f5ff;" | Eggs probably laid by an oviraptorid

|style="background:#e3f5ff;" |

Oviraptor

| O. philoceratops

| Bayn Dzak

| "Partial skeleton with skull, associated with a nest and juvenile."{{cite journal|last1=Osborn|first1=H. F.|date=1924|title=Three new Theropoda, Protoceratops zone, central Mongolia|journal=American Museum Novitates|number=144|pages=1–12|hdl=2246/3223|hdl-access=free|oclc=40272928}}

| An oviraptorid.

| 200px

rowspan="4" | Oviraptoridae indet.

| Indeterminate

| Zamyn Khondt

| "Nearly complete skeleton with skull."{{cite journal|last1=Barsbold|first1=R.|date=1981|title=Bezzubyye khishchnyye dinozavry Mongolii|trans-title=Toothless carnivorous dinosaurs of Mongolia|journal=Transactions, Joint Soviet–Mongolian Palaeontological Expedition|volume=15|pages=28–39|s2cid=195060070|language=ru}} [https://paleoglot.org/files/Barsbold_81.pdf Translated paper]

| An oviraptorid also known as the Zamyn Khondt oviraptorid. Uncertainly referred to Citipati.

| 200px

Indeterminate

| Zamyn Khondt

| "Nearly complete skull with atlas and axis."{{cite thesis|last1=Funston|first1=G. F.|date=2019|title=Anatomy, systematics, and evolution of Oviraptorosauria (Dinosauria, Theropoda)|type=PhD thesis|chapter=Chapter 4 – Oviraptoridae|publisher=University of Alberta|pages=446–556|chapter-url=https://era.library.ualberta.ca/items/32edf6a6-9791-478f-9b95-7deef3c6fd09/view/f4dada67-a5ad-439c-bf57-db37333f786a/Funston_Gregory_F_201909_PhD.pdf}}

| An oviraptorid.

| 200px

Indeterminate

| Udyn Sayr

| "Assemblage of individuals."

| An oviraptorid.

|

Indeterminate

| Not specified.

| "Two skulls with characteristic high crest."{{cite thesis|last1=Jansen|first1=S. O. K.|date=2008|title=Beak morphology in oviraptorids, based on extant birds and turtles|publisher=University of Oslo|pages=1–48|hdl=10852/11785|hdl-access=free|url=https://www.duo.uio.no/bitstream/handle/10852/11785/Jansenx2008x-xBeakxmorphologyxinxoviraptoridsxxbasedxonxextantxbirdsxandxturtlesxxCompressedxpicturesx.pdf?sequence=1&isAllowed=y}}{{cite web|date=2012|title=Oviraptor philoceratops|website=WitmerLab Collections|url=https://people.ohio.edu/witmerl/collections/Theropods/oviraptor.htm}}{{cite web|date=2021|title=Oviraptor philoceratops, skull and hand|website=Prehistoric Planet Store|url=http://www.prehistoricstore.com/item.php?item=1840}}

| An oviraptorid.

| 200px

===Troodontids===

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Almas

| A. ukhaa

| Ukhaa Tolgod

| "Skull with partial skeleton."{{Cite journal | doi = 10.1206/3889.1 | last1 = Pei | first1 = R. | last2 = Norell | first2 = M.A. |last3 = Barta |first3 = D.E | last4 = Bever| first4 = G.S. | last5 = Pittman | first5 = M. | last6 = Xu | first6 = Xing | year = 2017 | title = Osteology of a New Late Cretaceous Troodontid Specimen from Ukhaa Tolgod, Ömnögovi Aimag, Mongolia | journal = American Museum Novitates | issue = 3889 | pages = 1–47 | s2cid = 90883541 | url = https://www.biodiversitylibrary.org/item/267827 }}

| A troodontid.

| 200px

Archaeornithoides

| A. deinosauriscus

| Bayn Dzak

| "Partial skull."{{cite journal|last1=Elżanowski|first1=A.|last2=Wellnhofer|first2=P.|date=1992|title=A new link between theropods and birds from the Cretaceous of Mongolia|journal=Nature|volume=359|issue=6398|pages=821–823|doi=10.1038/359821a0|bibcode=1992Natur.359..821E|s2cid=4350813}}

| A troodontid? Uncertain relationships among coelurosaurs.

| File:Archaeornithoides.jpg

Byronosaurus

| B. jaffei

| Ukhaa Tolgod

| "Skull and fragmentary skeleton."{{cite journal|last1=Makovicky|first1=P. J.|last2=Norell|first2=M. A.|last3=Clark|first3=J. M.|last4=Rowe|first4=T. E.|date=2003|title=Osteology and Relationships of Byronosaurus jaffei (Theropoda: Troodontidae)| journal = American Museum Novitates|number=3402|pages=1–32|doi=10.1206/0003-0082(2003)402<0001:oarobj>2.0.co;2|hdl=2246/2828|s2cid=51824767 |hdl-access=free|url=https://digitallibrary.amnh.org/bitstream/handle/2246/2828//v2/dspace/ingest/pdfSource/nov/N3402.pdf?sequence=1&isAllowed=y}}

| A troodontid.

| 200px

Gobivenator

| G. mongoliensis

| Zamyn Khondt

| "Almost complete skeleton."{{Cite journal | last1 = Tsuihiji | first1 = T. | last2 = Barsbold | first2 = R. | last3 = Watabe | first3 = M. | last4 = Tsogtbaatar | first4 = K. | last5 = Chinzorig | first5 = T. | last6 = Fujiyama | first6 = Y. | last7 = Suzuki | first7 = S. | doi = 10.1007/s00114-014-1143-9 | title = An exquisitely preserved troodontid theropod with new information on the palatal structure from the Upper Cretaceous of Mongolia | journal = Naturwissenschaften | year = 2014 | pmid = 24441791| volume=101 | issue = 2 | pages=131–142| bibcode = 2014NW....101..131T | s2cid = 13920021 }}

| A troodontid.

| 200px

Saurornithoides

| S. mongoliensis

| Bayn Dzak

| "Skull with fragmentary skeleton."{{cite journal|last1=Norell|first1=M. A.|last2=Makovicky|first2=P. J.|last3=Bever|first3=G. S.|last4=Balanoff|first4=A. M.|last5=Clark|first5=J. M.|last6=Barsbold|first6=R.|last7=Rowe|first7=T.|date=2009|title=A review of the Mongolian Cretaceous dinosaur Saurornithoides (Troodontidae, Theropoda)|journal=American Museum Novitates|number=3654|pages=1–63|doi=10.1206/648.1|hdl=2246/5973|hdl-access=free|url=https://www.biodiversitylibrary.org/itempdf/280747}}

| A troodontid.

| 200px

rowspan="4" | Troodontidae indet.

| Indeterminate

| Ukhaa Tolgod

| "Partial skeleton."Prieto-Marquez, Bolortsetseg and Horner, 2011. A diminutive deinonychosaur (Dinosauria: Theropoda) from the Early Cretaceous of Oosh (Ovorkhangai, Mongolia). Alcheringa. 1–20.

| A troodontid.

|

Indeterminate

| Ukhaa Tolgod

| "Juvenile skulls, skeleton, and one nest."{{cite thesis|last1=Grellet-Tinner|first1=G.|date=2005|title=A phylogenetic analysis of oological characters: A case study of saurischian dinosaur relationships and avian evolution|chapter=CHAPTER VII: An Egg Clutch of the Troodontid Byronosaurus jaffei from the Gobi Desert: Novel Perspectives on the Origin of the Avian Reproductive Physiology|type=PhD thesis|publisher=University of Southern California|page=105|chapter-url=http://digitallibrary.usc.edu/digital/collection/p15799coll16/id/331907}}{{cite journal|last1=Bever|first1=G. S.|last2=Norell|first2=M. A.|date=2009|title=The Perinate Skull of Byronosaurus (Troodontidae) with Observations on the Cranial Ontogeny of Paravian Theropods|journal=American Museum Novitates|number=3657|pages=1–51|doi=10.1206/650.1|hdl=2246/5980|hdl-access=free|url=https://www.researchgate.net/publication/229181568}}

| A troodontid. Referred to either Almas,{{cite journal|last1=Pei|first1=R.|last2=Norell|first2=M. A.|date=2011|title=A new troodontid (Dinosauria: Theropoda) from the Late Cretaceous Djadokhta Formation of Mongolia|journal=Journal of Vertebrate Paleontology|volume=31|issue=supp. 002|pages=172A|doi=10.1080/02724634.2011.10635174|s2cid=210915628 }} or Byronosaurus.

| 200px

Indeterminate

| Ukhaa Tolgod

| "Fragmented skull and skeleton remains."{{cite journal|last1=Norell|first1=M. A.|last2=Hwang|first2=S. H.|date=2004|title=A Troodontid Dinosaur from Ukhaa Tolgod (Late Cretaceous Mongolia)|journal=American Museum Novitates|number=3446|pages=1–9|doi=10.1206/0003-0082(2004)446<0001:ATDFUT>2.0.CO;2|hdl=2246/2811|s2cid=10547745 |hdl-access=free|url=https://digitallibrary.amnh.org/bitstream/handle/2246/2811//v2/dspace/ingest/pdfSource/nov/N3446.pdf?sequence=1&isAllowed=y}}

| A troodontid. Provisionally referred to Saurornithoides, but now excluded.

|

Indeterminate

| Ukhaa Tolgod

| "Partial skull and skeletons from two specimens."

| A troodontid.

|

Flora

class="wikitable sortable"
GenusSpeciesLocationMaterialNotesImages
Radicites

| R. gobiensis

| Bayn Dzak

| "Twenty plant roots."{{cite journal |last1=Naugolnykh|first1=S. V.|date=2016|title=Upper Cretaceous Paleosols of the Bain Dzak Section, Southern Mongolia|journal=Paleontological Journal|volume=50|issue=12|pages=1451–1469|doi=10.1134/s0031030116120133|bibcode=2016PalJ...50.1451N |s2cid=133243515|url=https://www.researchgate.net/publication/313543951}}

| A tracheophyte, likely conifer.

|

See also

References