paramo ground tyrant
{{Short description|Species of bird}}
{{Use American English|date=April 2025}}
{{Speciesbox
| name = Paramo ground tyrant
| image = Plain-capped ground-tyrant (Muscisaxicola alpinus) on Senecio niveo-aureus Chingaza.jpg
| image_caption = Chingaza National Natural Park, Colombia
on Senecio niveo-aureus
| status = LC
| status_system = IUCN3.1
| genus = Muscisaxicola
| species = alpinus
| authority = (Jardine, 1849)
| synonyms = Muscisaxicola alpina (Jardine, 1849) [orthographic error]
| range_map = Muscisaxicola alpinus map.svg
}}
The paramo ground tyrant (Muscisaxicola alpinus), more widely known as the plain-capped ground-tyrant, is a species of bird in the family Tyrannidae, the tyrant flycatchers.{{cite web |url=https://www.worldbirdnames.org/new/bow/flycatchers/ |title=Tyrant flycatchers |website=IOC World Bird List |version =v 15.1 | editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela Rasmussen |date=March 2025 |access-date=March 3, 2025 }} It is found in Colombia, Ecuador, and possibly Peru.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 30 March 2025. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved 30 March 2025
Taxonomy and systematics
The paramo ground tyrant was originally described in 1849 as Taenioptera alpina.{{cite book | last=Jardine |first=William | date= 1849 | title=Contributions to ornithology for 1849 | publisher=W.H. Lizars |page=47-9 | url=https://www.biodiversitylibrary.org/page/51703070 |access-date=April 20, 2025 }} It was eventually moved to genus Muscisaxicola that had been erected in 1837.{{cite journal | last=d'Orbigny |first=Alcide |last2=de Lafresnaye |first2=Frédéric |title=Synopsis Avium | date= 1837 | journal=Magasin de zoologie | volume=7 |page=65 | language=Latin |url=https://www.biodiversitylibrary.org/page/37087510 |access-date=April 20, 2025 }} Until the 2020 publication of a molecular phylogenetic study it had been widely treated as conspecific with the cinereous ground tyrant (M. cinereus) and Taczanowski's ground tyrant (M. griseus).Chesser, R. T. (2000). Evolution in the High Andes: The phylogenetics of Muscisaxicola ground-tyrants. Molecular Phylogenetics and Evolution 15(3):369–380 Of the major taxonomic systems, only the IOC calls it the "paramo ground tyrant"; Wikipedia generally follows IOC nomenclature. The South American Classification Committee of the American Ornithological Society (SACC), the Clements taxonomy, and BirdLife International's Handbook of the Birds of the World call it the "plain-capped ground-tyrant".Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 30 March 2025. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved 30 March 2025Clements, J. F., P.C. Rasmussen, T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, A. Spencer, S. M. Billerman, B. L. Sullivan, M. Smith, and C. L. Wood. 2024. The eBird/Clements checklist of birds of the world: v2024. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved October 23, 2024HBW and BirdLife International (2024). Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 9. Available at: https://datazone.birdlife.org/about-our-science/taxonomy retrieved December 23, 2024
The paramo ground tyrant has three subspecies, the nominate M. a. alpinus (Jardine, 1849), M. a. columbianus (Chapman, 1912), and M. a. quesadae (Meyer de Schauensee, 1942).
Description
The paramo ground tyrant is {{convert|18|to|20|cm|in|abbr=on}} long. The sexes have the same plumage. Adults of the nominate subspecies have a sepia-brown crown, dusky lores, a wide white supercilium that extends past the eye, and a small white patch below the eye. Their nape and back are grayish brown. Their wings are a duskier grayish brown with some pale edges on the coverts. Their tail is blackish with thin white edges on the outermost feathers. Their underparts are grayish white that is darker on the breast than on the throat and belly. They have a dark brown iris, a shortish black bill, and black legs and feet. Juveniles have cinnamon tips on the wing coverts and thin cinnamon edges on the inner remiges. Their belly and vent have a buff tinge with whitish streaks. Subspecies M. a. columbianus is almost identical to the nominate but has slightly darker upperparts. M. a. quesadae has the same plumage as the nominate but is slightly smaller.Farnsworth, A., G. Langham, and E. de Juana (2020). Plain-capped Ground-Tyrant (Muscisaxicola alpinus), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.plcgrt1.01 retrieved April 20, 2025
Distribution and habitat
The paramo ground tyrant is a bird of the northern high Andes and has a disjunct distribution. Subspecies M. a. columbianus is found in Colombia's Central Andes between Caldas and Cauca departments. M. a. quesadae is found in Colombia's Eastern Andes of Boyacá, Santander, and Cundinamarca departments. The nominate subspecies is found from Nariño Department in extreme southwestern Colombia south on both slopes of the Andes through Ecuador to Azuay Province.{{cite book | last1 =McMullan | first1 =Miles | last2 =Donegan | first2 =Thomas M. | last3 =Quevedo | first3 =Alonso | title = Field Guide to the Birds of Colombia | publisher = Fundación ProAves| date =2010 | location =Bogotá | pages =162 |isbn =978-0-9827615-0-2 }}{{cite book | last =Ridgely | first =Robert S. | last2 =Greenfield | first2 =Paul J. | title =The Birds of Ecuador: Field Guide | publisher =Cornell University Press | volume = II| date =2001 | location =Ithaca | pages =512–513 | isbn =978-0-8014-8721-7 }} There are also sight records in northern Peru that might be of this species; the SACC calls it hypothetical in that country.{{cite book | last =Schulenberg | first =T.S. | last2 =Stotz | first2 =D.F. | last3 =Lane | first3 =D.F. | last4 =O'Neill | first4 =J.P. | last5 =Parker | first5 =T.A. III | title =Birds of Peru | publisher =Princeton University Press | edition =revised and updated |series=Princeton Field Guides | date =2010 | location =Princeton, NJ | pages =456 |isbn = 978-0691130231 }}
The paramo ground tyrant inhabits puna and páramo grasslands, often in rocky areas, and arid montane scrub. In elevation it ranges between {{convert|3200|and|4400|m|ft|abbr=on}} in Colombia and mostly between {{convert|3800|and|4600|m|ft|abbr=on}} in Ecuador. The sight records in Peru were between {{convert|2900|and|3350|m|ft|-2|abbr=on}}.
Behavior
=Movement=
=Feeding=
The paramo ground tyrant feeds on insects. It usually forages by itself in the breeding season; outside that it often is in small flocks that may include other species. It is almost wholly terrestrial; it will perch on rocks or a wall, though less often than some other ground tyrants. It runs and hops along the ground, stopping to stand erect before grabbing prey, or makes a short sally to the ground from a low perch.
=Breeding=
The paramo ground tyrant breeds between September and December in Colombia and August to October and possibly beyond in Ecuador. Males make a display flight in which they hover and then drop to the ground. The species' nest is a cup of grass and roots with minimal lining and typically is placed in a crevice or hole under a rock. The clutch is four eggs that are white with cinnamon flecks. Nothing else is known about the species' breeding biology.
{{birdsong|url=https://xeno-canto.org/species/Muscisaxicola-alpinus |species=the paramo ground tyrant}}
=Vocalization=
As of April 2025 xeno-canto had 10 recordings of ochre-naped ground tyrant vocalizations.{{cite web |url=https://xeno-canto.org/species/Muscisaxicola-alpinus |title=Paramo Ground Tyrant - Muscisaxicola alpinus |author= |date=April 20, 2025 |website=xeno-canto |access-date=April 20, 2025 }} The Cornell Lab of Ornithology's Macaulay Library had only three, all of which are also on xeno-canto.{{cite web |url=https://search.macaulaylibrary.org/catalog?taxonCode=plcgrt1&mediaType=audio&sort=rating_rank_desc |title=Plain-capped Ground-Tyrant Muscisaxicola alpinus |author= |date=April 20, 2025 |website=Birds of the World |publisher=Cornell Lab of Ornithology |access-date=April 20, 2025 }} The species is "[g]enerally quiet, though [it] occasionally gives a soft tik note".
Status
The IUCN has assessed the paramo ground tyrant as being of Least Concern. Its population size is not known and is believed to be stable. No immediate threats have been identified. It is considered "local but abundant where present" in Colombia and "fairly common" in Ecuador. It occurs in national parks and other protected areas in both countries.