2011 in arthropod paleontology

{{Short description|none}}

{{Year nav topic20|2011|arthropod paleontology|paleontology|science|reptile paleontology|paleobotany|paleoentomology |paleomalacology |archosaur paleontology |mammal paleontology |paleoichthyology}}

This list of fossil arthropods described in 2011 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2011. The list only includes taxa at the level of genus or species.

Arachnids

class="wikitable sortable" align="center" width="80%"
Name

! Novelty

! Status

! Authors

! Age

! Unit

! Location

! Notes

! Images

Acarus indicus{{cite journal |author1=Prabhat Kumar |author2=Neerja Jha |author3=D.D. Bhattacharya |author4=A.C. Pande |title=Acarid mites from Early Permian sediments of the Chamba valley, Himachal Pradesh, India |journal=Journal of the Palaeontological Society of India |year=2011 |url=http://palaeontologicalsociety.in/vol56_2/v6.pdf |volume=56 |issue=2 |pages=171–179 }}

|

Sp. nov

|

Disputed

|

Kumar et al.

|

Early Permian

|

Manjir Formation

|

{{Flag|India}}

|

A mite. Originally described as a species of Acarus; Sidorchuk (2018) considered it more likely to be a misidentified immature nothrine oribatid mite (probably a member of the family Nothridae), and more likely to be a contaminant from the extant soil fauna rather than a genuine Paleozoic fossil.{{cite journal |author=Ekaterina A. Sidorchuk |year=2018 |title=Mites as fossils: forever small? |journal=International Journal of Acarology |volume=44 |issue=8 |pages=349–359 |doi=10.1080/01647954.2018.1497085 |bibcode=2018IJAca..44..349S |s2cid=92357151 }}

|

Ameticos{{cite journal |author1=Russell J. Garwood |author2=Jason A. Dunlop |author3=Gonzalo Giribet |author4=Mark D. Sutton |title=Anatomically modern Carboniferous harvestmen demonstrate early cladogenesis and stasis in Opiliones |journal=Nature Communications |year=2011 |doi=10.1038/ncomms1458 |volume=2 |pages=Article number 444 |pmid=21863011 |bibcode=2011NatCo...2..444G |doi-access=free }}

|

Gen. et sp. nov

|

|

Garwood et al.

|

Carboniferous (late Stephanian)

|

|

{{Flag|France}}

|

A member of Opiliones. The type species is A. scolos.

|

Araneaovoius columbiae{{cite journal |last1=Dunlop |first1=J.A. |last2=Braddy |first2=S.J. |year=2011 |title=Cteniza bavincourti and the nomenclature of arachnid-related trace fossils |journal=The Journal of Arachnology |volume=39 |issue=2 |pages=250–257|doi=10.1636/CHa10-70.1 |s2cid=55206472 |url=https://www.biodiversitylibrary.org/part/229298 }}

|

Gen et comb nov

|

valid

|

Dunlop & Braddy

|

?Ypresian

|

Quesnel
?Fraser Formation

|

{{Flag|Canada}}
{{Flag|British Columbia}}

|

An orb-web spider egg sack ichnogenus.
New ichnogenus for Aranea columbiae (Scudder 1878).{{cite journal |last=Scudder |first=S. H |year=1878 |title=Additions to the Insect-fauna of the Tertiary beds at Quesnel, British Columbia |journal=Geological Survey of Canada, Report of Progress for |volume=1876-1877 |pages=457–464}}

|

File:Araneaovoius columbiae Scudder 1890 pl2 Fig1.png
(1890 illustration]]

Burmesiola{{cite book |author =Jörg Wunderlich |year=2011 |chapter=Some fossil spiders (Araneae) in Cretaceous ambers |editor=Jörg Wunderlich |title=Beiträge zur Araneologie, 6 |pages=539–557 }}

|

Gen. et sp. nov

|

Valid

|

Wunderlich

|

Late Cretaceous (Cenomanian)

|

Burmese amber

|

{{Flag|Myanmar}}

|

A tree trunk spider. Genus includes new species B. cretacea.

|

Chaerilobuthus{{cite journal |author1=Wilson R. Lourenço |author2=Alex Beigel |title=A new scorpion fossil from the Cretaceous amber of Myanmar (Burma). New phylogenetic implications |journal=Comptes Rendus Palevol |year=2011 |doi=10.1016/j.crpv.2011.08.001 |volume=10 |issue=8 |pages=635–639 |bibcode=2011CRPal..10..635L }}

|

Gen. et sp. nov

|

Valid

|

Lourenço & Beigel

|

Late Cretaceous (Cenomanian)

|

Burmese amber

|

{{Flag|Myanmar}}

|

A chaerilobuthid scorpion. The type species is C. complexus.

|

Eoscaphiella

|

Gen. et sp. nov

|

Valid

|

Wunderlich

|

Late Cretaceous (Cenomanian)

|

Burmese amber

|

{{Flag|Myanmar}}

|

A member of the family Tetrablemmidae. Genus includes new species E. ohlhoffi.

|

Episinus tibiaseta{{cite book |author =Jörg Wunderlich |year=2011 |chapter=Some fossil spiders in Dominican amber (Araneae: Hersiliidae, Theridiidae, Gnaphosidae) |editor=Jörg Wunderlich |title=Beiträge zur Araneologie, 6 |pages=461–471 }}

|

Sp. nov

|

Valid

|

Wunderlich

|

Miocene

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A member of the family Theridiidae, a species of Episinus.

|

Femurraptor

|

Gen. et sp. nov

|

Valid

|

Wunderlich

|

Miocene

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A member of the family Theridiidae. Genus includes new species F. dominicanus.

|

Fictotama maculosa

|

Sp. nov

|

Valid

|

Wunderlich

|

Miocene

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A tree trunk spider.

|

Glaesacarus{{cite journal |author1=Ekaterina A. Sidorchuk |author2=Pavel B. Klimov |title=Redescription of the mite Glaesacarus rhombeus (Koch & Berendt, 1854) from Baltic amber (Upper Eocene): evidence for female-controlled mating |journal=Journal of Systematic Palaeontology |year=2011 |doi=10.1080/14772019.2011.566585 |volume=9 |issue=2 |pages=183–196 |bibcode=2011JSPal...9..183S |s2cid=84860361 }}

|

Gen. et comb. nov

|

Valid

|

Sidorchuk & Klimov

|

Eocene

|

Baltic amber

|

Europe (Baltic Sea region)

|

A mite; a new genus for "Acarus" rhombeus Koch & Berendt (1854)

|

Jerseyuloborus

|

Gen. et sp. nov

|

Valid

|

Wunderlich

|

Late Cretaceous

|

New Jersey amber

|

{{Flag|United States}}

|

A member of the family Uloboridae. Genus includes new species J. longisoma.

|

Macrogyion

|

Gen. et sp. nov

|

|

Garwood et al.

|

Carboniferous (late Stephanian)

|

|

{{Flag|France}}

|

A member of Opiliones. The type species is M. cronus.

|

Nephila jurassica{{cite journal |last1=Selden |first1=P. A. |last2=Shih |first2=C.K. |last3=Ren |first3=D. |title=A golden orb-weaver spider (Araneae: Nephilidae: Nephila) from the Middle Jurassic of China |journal= Biology Letters |year=2011 |doi=10.1098/rsbl.2011.0228 |pmid=21508021 |pmc=3169061 |volume=7 |issue=5 |pages=775–8}}{{cite journal |last1=Selden |first1=P. A. |last2=Shih |first2=C.K. |last3=Ren |first3=D. |year= 2013 |title=A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group |journal= Naturwissenschaften |doi=10.1007/s00114-013-1121-7 |pmid=24317464 |volume=100 |issue=12 |pages=1171–1181|bibcode = 2013NW....100.1171S |pmc=3889289 }}

|

sp nov

|

Objective synonym

|

Selden, Shih & Ren

|

Middle Jurassic

|

Jiulongshan Formation

|

{{Flag|China}}

|

largest fossil spider described.
objective syn of Mongolarachne jurassica

|

File:Mongolarachne.jpg]]

Labidostomma paleoluteum{{cite journal |author1=Jason A. Dunlop |author2=Michel Bertrand |title=Fossil labidostomatid mites (Prostigmata: Labidostommatidae) from Baltic amber |journal=Acarologia |year=2011 |doi=10.1051/acarologia/20112006 |volume=51 |issue=2 |pages=191–198 |url=https://hal.archives-ouvertes.fr/hal-01600007/file/Acarologia-2011-51-191-198.pdf }}

|

Sp. nov

|

Valid

|

Dunlop & Bertrand

|

Eocene

|

Baltic amber

|

Europe (Baltic Sea region)

|

A mite, a species of Labidostomma.

|

Praeterleptoneta tibialis

|

Sp. nov

|

Valid

|

Wunderlich

|

Late Cretaceous (Cenomanian)

|

Burmese amber

|

{{Flag|Myanmar}}

|

A member of Araneomorphae belonging to the family Praeterleptonetidae. Originally described as a species of Praeterleptoneta, subsequently transferred to the separate genus Parvispina.{{cite book |author =Jörg Wunderlich |year=2015 |chapter=On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae) |editor=Jörg Wunderlich |title=Beiträge zur Araneologie, 9: Mesozoic spiders and other fossil arachnids |pages=21–408 }}

|

Scopoides dominicanus

|

Sp. nov

|

Valid

|

Wunderlich

|

Miocene

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A ground spider, a species of Scopoides.

|

Siro balticus{{cite journal |author1=Jason A. Dunlop |author2=Plamen G. Mitov |title=The first fossil cyphophthalmid harvestman from Baltic amber |journal=Arachnologische Mitteilungen |year=2011 |url=http://www.european-arachnology.org/wdp/wp-content/uploads/2015/08/47-54_Dunlop.pdf |volume=40 |pages=47–54 |doi=10.5431/aramit4006 |doi-access=free }}

|

Sp. nov

|

Valid

|

Dunlop & Mitov

|

Eocene

|

Baltic amber

|

Europe (Baltic Sea region)

|

A member of Opiliones, a species of Siro.

|

Spinocharinus{{cite journal |author1=Markus Poschmann |author2=Jason A. Dunlop |title=Trigonotarbid arachnids from the Lower Devonian (Siegenian) of Bürdenbach (Lahrbach Valley, Westerwald area, Rhenish Slate Mountains, Germany) |journal=Paläontologische Zeitschrift |year=2011 |doi=10.1007/s12542-011-0109-x |volume=85 |issue=4 |pages=433–447 |bibcode=2011PalZ...85..433P |s2cid=129040136 }}

|

Gen. et sp. nov

|

Valid

|

Poschmann & Dunlop

|

Early Devonian

|

|

{{Flag|Germany}}

|

A member of Trigonotarbida. The type species is S. steinmeyeri.

|

Crustaceans

class="wikitable sortable" align="center" width="100%"
Name

! Novelty

! Status

! Authors

! Age

! Unit

! Location

! Notes

! Images

Aenigmastacus{{cite journal |last1=Feldmann |first1=R. |last2=Schweitzer |first2=C. |last3=Leahy |first3=J. |year=2011 |title=New Eocene crayfish from the McAbee Beds in British Columbia: First record of Parastacoidea in the Northern Hemisphere |journal=Journal of Crustacean Biology |volume=31 |issue=2 |pages=320–331 |doi=10.1651/10-3399.1|doi-access=free }}

|

Gen et sp nov

|

valid

|

Feldmann, Schweitzer, & Leahy

|

Eocene
Ypresian

|

Okanagan Highlands
Tranquille Formation

|

{{Flag|Canada}}
{{Flag|British Columbia}}

|

A parastacid freshwater crayfish

|

Dynamenella miettoi{{cite journal |author1=Antonio De Angeli |author2=Alessandro Lovato |title=Dynamenella miettoi n. sp., nuovo isopode (Crustacea, Flabellifera, Sphaeromatidae) dell'Eocene del Veneto (Vicenza, Italia settentrionale) |journal=Lavori – Società Veneziana di Scienze Naturali |year=2011 |url=http://decapoda.nhm.org/pdfs/31742/31742.pdf |volume=36 |pages=111–118 }}

|

Sp. nov

|

Valid

|

De Angeli & Lovato

|

Eocene
Lutetian

|

|

{{Flag|Italy}}

|

An isopod belonging to the family Sphaeromatidae.

|

Insects

class="wikitable sortable" align="center" width="100%"
Name

! Novelty

! Status

! Authors

! Age

! Unit

! Location

! Notes

! Images

Agulla protomaculata{{Cite journal|last1=Engel |first1=Michael S. |year=2011 |title=A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae)|journal= Transactions of the Kansas Academy of Science |volume=114 |issue= 1–2 |pages = 77–87 |doi=10.1660/062.114.0107 |s2cid=85011364 }}

|

sp nov

|

jr synonym

|

Engel

|

Lutetian

|

Green River Formation

|

{{Flag|USA}}

|

A raphidiid snakefly.
Moved to Archiinocellia protomaculata in 2021{{Cite journal|last1=Archibald |first1=S. B. |last2=Makarkin |first2=V. N. |title=Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation |year=2021 |journal=Zootaxa |volume=4951 |issue=1 |pages=41–79 |doi=10.11646/zootaxa.4951.1.2 |pmid=33903413 |s2cid=233411745 }}

|

file:Archiinocellia protomaculata holotype NMNH-USNM PAL 569865 img1.tif]]

Alicodoxa{{cite journal |last1=Emeljanov |first1=A. F. |last2=Shcherbakov |first2=D. E. |year=2011 |title=A new genus and species of Dictyopharidae (Homoptera) from Rovno and Baltic amber based on nymphs |journal=ZooKeys |issue=130 |pages=175–184 |doi=10.3897/zookeys.130.1775|pmid=22259275 |pmc=3260758 |doi-access=free |bibcode=2011ZooK..130..175E }}

|

Gen et sp nov

|

Valid

|

Emeljanov & Shcherbakov

|

late Eocene

|

Rovno amber

|

{{Flag|Ukraine}}

|

A planthopper, type species A. rasnitsyni
also found in Baltic amber

|

File:Alicodoxa rasnitsyni holotype Fig 3.jpg]]

Anonychomyrma geinitzi{{cite journal |last1=Heterick |first1=B. E. |last2=Shattuck |first2=S. |year=2011 |title=Revision of the ant genus Iridomyrmex (Hymenoptera: Formicidae) |journal=Zootaxa |volume=2845 |pages=169|doi=10.11646/zootaxa.2845.1.1 |doi-access=free }}

|

Comb nov

|

valid

|

(Mayr)

|

Lutetian

|

Baltic amber

|

{{Flag|Europe}}

|

Fossil Dolichoderin ant

|

File:Yantaromyrmex geinitzi MBI2289 02.jpg]]

Aradus andancensis{{cite journal |last1=Marchal |first1=L. |last2=Guilbert |first2=E. |last3=Brisac |first3=P. |last4=Nel. |first4=A. |year=2011 |title=A new record and a new species of Aradidae fossils (Hemiptera: Heteroptera) |journal=Zootaxa |volume=2832 |pages=56–62|doi=10.11646/zootaxa.2832.1.4 }}

|

Sp nov

|

valid

|

Marchal, Guilbert,
Brisac, & Nel

|

Turolian

|

Montagne d'Andance

|

{{Flag|France}}

|

A flat bug

|

File:Aradus andancensis holotype MNHN.F.A33748 direct lighting.jpg]]

Archiaustroconops borkenti{{cite journal|last1=Pérez-de la Fuente |first1=R. |last2=Delclòs |first2=X. |last3=Peñalver |first3=E. |last4=Arillo |first4=A. |year=2011 |title=Biting midges (Diptera: Ceratopogonidae) from the Early Cretaceous El Soplao amber (N Spain) |journal=Cretaceous Research |volume=32 |issue=6 |pages=750–761 |doi=10.1016/j.cretres.2011.05.003 |bibcode=2011CrRes..32..750P }}

|

Sp. nov

|

Valid

|

Pérez-de la Fuente et al.

|

Early Cretaceous (Albian)

|

El Soplao amber deposit

|

{{Flag|Spain}}

|

A fly belonging to the family Ceratopogonidae.

|

Arostropsis{{Cite journal |last1=Yunakov |first1=N.N. |last2=Kirejtshuk |first2=A.G. |year=2011 |title=New genus and species of broad-nosed weevils from Baltic amber and notes on fossils of the subfamily Entiminae (Coleoptera, Curculionidae) |journal= ZooKeys |issue=160 |pages = 73–96 |doi= 10.3897/zookeys.160.2108 |pmid=22303121 |pmc=3253632|doi-access=free |bibcode=2011ZooK..160...73Y }}

|

Gen et sp nov

|

valid

|

Yunakov & Kirejtshuk

|

Eocene

|

Baltic amber

|

{{Flag|Europe}}

|

Monotypic broad-nosed weevil genus, one species A. groehni

|

File:ZooKeys-160-073-g001 Arostropsis groehni 02.jpg]]

Astreptolabis{{Cite journal |last1=Engel |first1=M.S. |year=2011 |title=New earwigs in mid-Cretaceous amber from Myanmar (Dermaptera, Neodermaptera) |journal= ZooKeys |issue=130 |pages = 137–152 |doi= 10.3897/zookeys.130.1293 |pmid=22259272 |pmc=3260755|doi-access=free |bibcode=2011ZooK..130..137E }}

|

Subfam, Gen et sp nov

|

valid

|

Engel

|

Albian

|

Burmese amber

|

{{Flag|Myanmar}}

|

Monotypic earwig subfamily, one species A. ethirosomatia

|

File:Astreptolabis ethirosomatia holotype ZooKeys-130-137-g001.jpg]]

Atriculicoides szadziewskii

|

Sp. nov

|

Valid

|

Pérez-de la Fuente et al.

|

Early Cretaceous (Albian)

|

El Soplao amber deposit

|

{{Flag|Spain}}

|

A fly belonging to the family Ceratopogonidae.

|

Baltimartyria rasnitsyni{{Cite journal |last1=Mey |first1=Wolfram |title=On the systematic position of Baltimartyria Skalski, 1995 and description of a new species from Baltic amber (Lepidoptera, Micropterigidae) |journal= ZooKeys |year=2011 |issue=130 |pages=331–342 |url=https://www.pensoft.net/journals/zookeys/article/1480/on-the-systematic-position-of-baltimartyria-skalski-1995-and-description-of-a-new-species-from-baltic-amber-lepidoptera-|doi=10.3897/zookeys.130.1480 |pmid=22259287 |pmc=3260770|doi-access=free |bibcode=2011ZooK..130..331M }}

|

sp nov

|

valid

|

Mey

|

Lutetian

|

Baltic amber

|

{{Flag|Europe}}

|

the second species in Baltimartyria

|

File:Baltimartyria rasnitsyni.JPG]]

Baryshnyala{{cite journal|last1=Ilger|first1=J.-M.|last2=Brauckmann|first2=C.|year=2011|title=The smallest Neoptera (Baryshnyalidae fam. n.) from Hagen-Vorhalle (early Late Carboniferous: Namurian B; Germany)|journal=ZooKeys|issue=130|pages=91–102|doi=10.3897/zookeys.130.1422|doi-access=free|pmid=22259268 |pmc=3260751 |bibcode=2011ZooK..130...91I }}

|

Gen. et sp. nov

|

Valid

|

Ilger & Brauckmann

|

Late Carboniferous (early Bashkirian)

|

Ziegelschiefer Formation

|

{{flag|Germany}}

|

A neopteran insect of uncertain systematic placement, assigned to the new family Baryshnyalidae. The type species is B. occulta.

|

File:Baryshnyala occulta.jpg]]

Ctenoplectrella phaeton{{cite journal |last1=Gonzalez |first1=V. H. |last2=Engel |first2=M. S. |year=2011 |title=A new species of the bee genus Ctenoplectrella in middle Eocene Baltic amber (Hymenoptera, Megachilidae) |journal=ZooKeys |issue=111 |pages=41–49 |doi=10.3897/zookeys.111.1593|pmid=21852938 |pmc=3142689|doi-access=free |bibcode=2011ZooK..111...41G }}

|

sp nov

|

Valid

|

Gonzalez & Engel

|

Lutetian

|

Baltic Amber

|

{{flag|Europe}}

|

A leaf cutter bee

|

file:Ctenoplectrella phaeton ZooKeys fig2.jpg]]

Dasyleptus triassicus{{cite journal|last1=Bechly|first1=G.|last2=Stockar|first2=R.|year=2011|title=The first Mesozoic record of the extinct apterygote insect genus Dasyleptus (Insecta: Archaeognatha: Monura: Dasyleptidae) from the Triassic of Monte San Giorgio (Switzerland)|journal=Palaeodiversity|volume=4|pages=23–37|url=https://bechly.lima-city.de/Palaeodiversity_4_BechlyStockar.pdf}}

|

Sp. nov

|

Valid

|

Bechly & Stockar

|

Middle Triassic (Ladinian)

|

Meride Limestone

|

{{Flag|Switzerland}}

|

A member of Archaeognatha, a species of Dasyleptus.

|

Deinodryinus velteni{{Cite journal |last1= Guglielmino |first1=A. |last2=Olmi |first2=M. |year=2011 |title=Revision of fossil species of Deinodryinus, with description of a new species (Hymenoptera, Dryinidae) |journal= ZooKeys |issue=130 |pages = 495–504 |doi=10.3897/zookeys.130.1326 |pmid=22259295 |pmc=3260777|doi-access=free |bibcode=2011ZooK..130..495G }}

|

sp nov

|

Valid

|

Guglielmino & Olmi

|

Lutetian

|

Baltic amber

|

{{Flag|Europe}}

|

A dryinid wasp.

|

File:ZooKeys-130-495-g003 Deinodryinus velteni.jpg]]

Diochus electrus{{Cite journal |last1=Chatzimanolis |first1=S. |last2=Engel |first2=M. S. |title=A new species of Diochus from Baltic amber (Coleoptera, Staphylinidae, Diochini) |journal=ZooKeys |year=2011 |issue=138 |pages=65–73 |doi=10.3897/zookeys.138.1896 |url=https://www.pensoft.net/J_FILES/1/articles/1896/1896-G-3-layout.pdf |pmid=22144854 |pmc=3208521|doi-access=free |bibcode=2011ZooK..138...65C }}

|

sp nov

|

valid

|

Chatzimanolis & Engel

|

Lutetian

|

Baltic amber

|

{{Flag|Europe}}

|

first valid Staphylininae species from the fossil record

|

File:Diochus electrus ZooKeys-138-065-g001a.jpg]]

Dryinus rasnitsyni{{Cite journal |last1=Olmi |first1=Massimo |last2=Guglielmino |first2=Adalgisa |title=Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae) |journal=ZooKeys |year=2011 |issue=130 |pages=505–514 |doi= 10.3897/zookeys.130.1335|pmid=22259296 | pmc=3260778|url=https://www.pensoft.net/J_FILES/1/articles/1335/1335-G-3-layout.pdf|doi-access=free |bibcode=2011ZooK..130..505O }}

|

sp nov

|

valid

|

Olmi & Guglielmino

|

Burdigalian

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

second fossil Dryinus lamellatus group species

|

File:Dryinus rasnitsyni ZooKeys-130-505-g004.jpg]]

Eldermyrmex{{cite journal |last1=Heterick |first1=B. E. |last2=Shattuck |first2=S. |year=2011 |title=Revision of the ant genus Iridomyrmex (Hymenoptera: Formicidae) |journal=Zootaxa |volume=2845 |pages=1–174|doi=10.11646/zootaxa.2845.1.1 |doi-access=free }}

|

Gen et comb nov

|

Valid

|

Heterick & Shattuck

|

Lutetian

|

Baltic amber

|

{{Flag|Russia}}

|

Fossil Dolichoderine ant,
Type species "Iridomyrmex" oblongiceps

|

File:Eldermyrmex oblongiceps GZG-BST05036 profile.tif]]

Iberoraphidia{{cite journal |last1=Jepson |first1=J.E. |last2=Ansorge |first2=J. |last3=Jarzembowski |first3=E.A. |year=2011 |title=New snakeflies (Insecta: Raphidioptera) from the Lower Cretaceous of the UK, Spain and Brazil |journal= Palaeontology |volume=54 |issue=2 |pages=385–395 |doi=10.1111/j.1475-4983.2011.01038.x|doi-access=free |bibcode=2011Palgy..54..385J }}

|

sp nov

|

valid

|

Jepsen, Ansorge, & Jarzembowski

|

Lower Barremian

|

Serra del Montsec

|

{{Flag|Spain}}

|

A mesoraphidiid snakefly, one species Iberoraphidia dividua

|

Lebanoculicoides excantabris

|

Sp. nov

|

Valid

|

Pérez-de la Fuente et al.

|

Early Cretaceous (Albian)

|

El Soplao amber deposit

|

{{Flag|Spain}}

|

A fly belonging to the family Ceratopogonidae.

|

Lebanoraphidia{{cite journal|last1=Bechly |first1=G. |last2=Wolf-Schwenninger |first2=K. |year=2011 |title=A new fossil genus and species of snakefly (Raphidioptera: Mesoraphidiidae) from Lower Cretaceous Lebanese amber, with a discussion of snakefly phylogeny and fossil history |journal=Insect Systematics and Evolution |volume=42 |issue=2 |pages=221–236 |doi=10.1163/187631211X568164 |url=http://www.bernstein.naturkundemuseum-bw.de/odonata/ISE-Raphidioptera.pdf |archive-url=https://web.archive.org/web/20140305182820/http://www.bernstein.naturkundemuseum-bw.de/odonata/ISE-Raphidioptera.pdf |url-status=dead |archive-date=2014-03-05 }}

|

sp nov

|

valid

|

Bechly & Wolf-Schwenninger

|

Upper Neocomian

|

Jezzine

|

{{Flag|Lebanon}}

|

A mesoraphidiid snakefly, one species Lebanoraphidia nana

|

Leptolingia calonervis{{cite journal|last1=Shi |first1=C. |last2=Yang |first2=Q. |last3=Ren |first3=D. |year=2011 |title=Two new fossil lacewing species from the Middle Jurassic of Inner Mongolia, China (Neuroptera: Grammolingiidae) |journal=Acta Geologica Sinica |volume=85 |issue=2 |pages=482–489 |url=http://202.204.209.200/upload/20110408010459.pdf |archive-url=https://web.archive.org/web/20120324183119/http://202.204.209.200/upload/20110408010459.pdf |url-status=dead |archive-date=2012-03-24 |doi=10.1111/j.1755-6724.2011.00416.x |bibcode=2011AcGlS..85..482S }}

|

sp.

|

Valid

|

Shi, Liu, & Ren

|

Middle Jurassic

|

Jiulongshan Formation

|

{{Flag|China}}

|

|

Leptolingia imminuta{{cite journal |last1=Liu |first1=Y. |last2=Shi |first2=C. |last3=Ren |first3=D. |year= 2011 |title= A new lacewing (Insecta: Neuroptera: Grammolingiidae) from the Middle Jurassic of Inner Mongolia, China |journal= Zootaxa |volume= 2897 |pages= 51–56 |url=http://www.mapress.com/zootaxa/2011/f/zt02897p056.pdf|doi=10.11646/zootaxa.2897.1.5 |doi-access=free }}

|

sp.

|

Valid

|

Liu, Shi, & Ren

|

Middle Jurassic

|

Daohugou Beds

|

{{Flag|China}}

|

The smallest species known in the family Grammolingiidae

|

Litholingia ptesa

|

sp.

|

Valid

|

Shi, Yang, & Ren

|

Middle Jurassic

|

Jiulongshan Formation

|

{{Flag|China}}

|

|

Mickoleitia longimanus{{cite journal|last1=Staniczek |first1=A.H. |last2=Bechly |first2=G. |last3=Godunko |first3=R.J. |year=2011 |title=Coxoplectoptera, a new fossil order of Palaeoptera (Arthropoda: Insecta), with comments on the phylogeny of the stem group of mayflies (Ephemeroptera) |journal=Insect Systematics & Evolution |volume=42 |issue=2 |pages=101–138 |doi=10.1163/187631211X578406 |url=http://www.bernstein.naturkundemuseum-bw.de/odonata/Coxoplectoptera.pdf |archive-url=https://web.archive.org/web/20111003171623/http://www.bernstein.naturkundemuseum-bw.de/odonata/Coxoplectoptera.pdf |url-status=dead |archive-date=2011-10-03 }}

|

Ord., fam., gen. et sp.

|

valid

|

Staniczek, Bechly, & Godunko

|

Lower Cretaceous

|

Crato Formation

|

{{Flag|Brazil}}

|

A new genus in the new family Mickoleitiidae and a new insect order Coxoplectoptera.

|

File:Mickoleitia longimanus.jpg]]

Myanmymar{{cite journal|last1=Poinar |first1=G. |last2=Huber |first2=J. T. |year=2011 |title=A new genus of fossil Mymaridae (Hymenoptera) from Cretaceous amber and key to Cretaceous mymarid genera |journal=ZooKeys |issue=130 |pages=461–472 |doi=10.3897/zookeys.130.1241 |url=https://www.pensoft.net/inc/journals/download.php?fileId=2891&fileTable=J_GALLEYS |pmid=22259293 |pmc=3260775|doi-access=free |bibcode=2011ZooK..130..461P }}

|

gen et sp nov

|

valid

|

Huber

|

Albian

|

Burmese amber

|

{{Flag|Myanmar}}

|

oldest mymarid described

|

File:Myanmymar aresconoides.jpg]]

Nanotermes{{Cite journal |last1=Engel |first1=M.S. |last2=Grimaldi |first2=D.A. |last3=Nascimbene |first3=P.C. |last4=Singh |first4=H. |year=2011 |title=The termites of Early Eocene Cambay amber, with the earliest record of the Termitidae (Isoptera) |journal= ZooKeys |issue=148 |pages = 105–123 |doi= 10.3897/zookeys.148.1797 |pmid=22287892 |pmc=3264413|doi-access=free |bibcode=2011ZooK..148..105E }}

|

Gen et sp. nov

|

valid

|

Engel & Grimaldi

|

Ypresian

|

Cambay amber

|

{{Flag|India}}

|

A Termitid termite

|

File:ZooKeys-148-105-g001 Nanotermes isaacae.jpg]]

Ordralfabetix{{Cite journal |last1=Szwedo |first1=J. |title=Ordralfabetix sirophatanis gen. et sp. n.—the first Lophopidae from the Lowermost Eocene Oise amber, Paris Basin, France (Hemiptera: Fulgoromorpha) |journal=Zootaxa |year=2011 |volume=2822 |pages=52–60 |url=http://www.mapress.com/zootaxa/2011/f/z02822p060f.pdf|doi=10.11646/zootaxa.2822.1.3 }}

|

gen et sp.

|

valid

|

Szwedo

|

Ypresian

|

Oise amber

|

{{Flag|France}}

|

Second fossil lophopid planthopper from Europe

|

File:Ordralfabetix sirophatanis holotype fig 5.jpg]]

Parastylotermes krishnai

|

Sp. nov

|

valid

|

Engel & Grimaldi

|

Ypresian

|

Cambay amber

|

{{Flag|India}}

|

A Stylotermitid termite

|

File:ZooKeys-148-105-g001 Parastylotermes krishnai.jpg]]

Platyperla marquati{{Cite journal |last1= Gallego |first1= O.F. |last2= Rébori |first2=L |last3=Zavattieri |first3=A.M. |last4= Sinitshenkova |first4= N. |last5=Lara |first5= M.B. |last6=Martins-Neto |first6=R.G. |year=2011 |title=The most ancient Platyperlidae (Insecta, Perlida= Plecoptera) from early Late Triassic deposits in southern South America |journal=Ameghiniana |volume=48 |issue=4 |pages=447–461 |url=http://www.ameghiniana.org.ar/index.php/ameghiniana/article/view/360 |doi=10.5710/amgh.v48i4(360)|s2cid= 131681078 |url-access=subscription }}

|

Species

|

Valid

|

Gallego et al.

|

Late Triassic

|

Potrerillos Formation

|

{{flag|Argentina}}

|

A platyperlid stonefly, a species of Platyperla.

|

Plumalexius{{Cite journal |last1=Brothers |first1=Denis J. |title=A new Late Cretaceous family of Hymenoptera, and phylogeny of the Plumariidae and Chrysidoidea (Aculeata) |year=2011 |issue=130 |pages=515–542 |doi=10.3897/zookeys.130.1591|url=https://www.pensoft.net/J_FILES/1/articles/1591/1591-G-3-layout.pdf |pmid=22259297 |pmc=3260779 |journal=ZooKeys|doi-access=free |bibcode=2011ZooK..130..515B }}

|

fam, gen et sp.

|

valid

|

Brothers

|

Turonian

|

New Jersey amber

|

{{Flag|USA}}

|

sister genus to Plumariidae

|

File:Plumalexius ZooKeys-130-515-g002c.jpg]]

Pristomyrmex rasnitsyni{{cite journal |last1=Dlussky |first1=G. M. |last2=Radchenko |first2=A. G. |year=2011 |title= Pristomyrmex rasnitsyni sp. n., the first known fossil species of the ant genus Pristomyrmex Mayr (Hymenoptera: Formicidae) from the Late Eocene Danish amber |journal=Russian Entomological Journal |volume=20 |issue=3 |pages=251–254|doi=10.15298/rusentj.20.3.05 |doi-access=free }}

|

sp nov

|

valid

|

Dlussky & Radchenko

|

Late Eocene

|

Scandinavian amber

|

{{Flag|Denmark}}

|

A fossil ant

|

Prostylotermes

|

Gen et sp. nov

|

valid

|

Engel & Grimaldi

|

Ypresian

|

Cambay amber

|

{{Flag|India}}

|

A Stylotermitid termite

|

File:ZooKeys-148-105-g002 Prostylotermes kamboja a.jpg]]

Renphasma{{cite journal |last1=Nel |first1=A. |last2=Delfosse |first2=E. |year=2011 |title=A new Chinese Mesozoic stick insect |journal=Acta Palaeontologica Polonica |volume=56 |issue=2 |pages=429–432 |doi=10.4202/app.2009.1108 |doi-access=free }}

|

Gen. et sp.

|

Valid

|

Nel & Delfosse

|

Early Cretaceous

|

Yixian Formation

|

{{Flag|China}}

|

A stick insect

|

File:Renphasma sinica.jpg]]

Termitaradus dominicanus

|

Sp nov

|

valid

|

Poinar

|

Burdigalian?

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A termitaphidid termite bug

|

Titanomyrma lubei{{cite journal |last1=Archibald| first1=S. Bruce |last2=Johnson |first2=Kirk R. |last3=Mathewes |first3=Rolf W. |last4=Greenwood |first4=David R. |year=2011 |title=Intercontinental dispersal of giant thermophilic ants across the Arctic during early Eocene hyperthermals |journal=Proceedings of the Royal Society B |volume=278 |issue=1725 |pages=3679–3686 |doi=10.1098/rspb.2011.0729 |pmid= 21543354|pmc=3203508}}

|

Gen, sp, et comb nov.

|

valid

|

Archibald, Johnson,
Mathewes, & Greenwood

|

Early Eocene

|

Messel shales

|

{{Flag|Germany}}

|

new genus for the largest known ants

|

File:Titanomyrma simillima SMFMEI01006 01.jpg]]

Tytthodiplatys

|

Gen et sp. nov

|

Valid

|

Engel

|

Albian

|

Burmese amber

|

{{Flag|Myanmar}}

|

Oldest member of the family Diplatyidae

|

File:ZooKeys-130-137-g002 Tytthodiplatys mecynocercus.jpg]]

Undulopsychopsis{{cite journal |last1=Peng |first1=Y. |last2=Makarkin |first2=V.N. |last3=Wang |first3=X. |last4=Ren |first4=D. |year=2011 |title=A new fossil silky lacewing genus (Neuroptera, Psychopsidae) from the Early Cretaceous Yixian Formation of China |journal=ZooKeys |issue=130 |pages=217–228 |doi=10.3897/zookeys.130.1576 |pmid=22259278 |pmc=3260761|doi-access=free |bibcode=2011ZooK..130..217P }}

|

Gen et sp. nov

|

Valid

|

Peng, Makarkin, Wang, & Ren

|

Lower Barremian to Upper Aptian

|

Yixian Formation

|

{{Flag|China}}

|

A psychopsid silky lacewing, only species U. alexi

|

File:ZooKeys-130-217-g001 Undulopsychopsis alexi.jpg]]

Vetuformosa{{cite journal |last1=Poinar |first1=G. |year=2011 |title=Vetuformosa buckleyi n. gen., n. sp. (Ephemeroptera: Baetidae; Vetuformosinae n. subfam.), a new subfamily of mayflies in Early Cretaceous Burmese amber |journal=Historical Biology |volume=23 |issue=4 |pages=369–374|doi=10.1080/08912963.2011.559084 |bibcode=2011HBio...23..369P |s2cid=83703856 }}

|

Sub. Fam., Gen., et Sp. Nov

|

valid

|

Poinar

|

Albian

|

Burmese amber

|

{{Flag|Myanmar}}

|

A baetid mayfly.
The type species is V. buckleyi,
type genus of subfamily Vetuformosinae

|

File:Vetuformosa buckleyi holotype.jpg]]

Zophotermes

|

Gen et sp. nov

|

valid

|

Engel & Singh

|

Ypresian

|

Cambay amber

|

{{Flag|India}}

|

A Rhinotermitid termite

|

File:ZooKeys-148-105-g001 Zophotermes ashoki.jpg]]

Trilobites

class="wikitable sortable" align="center" width="100%"
Name

! Novelty

! Status

! Authors

! Age

! Unit

! Location

! Notes

! Images

Chasbellus{{cite journal |author1=James D. Loch |author2=John F. Taylor |title=New symphysurinid trilobites from the Cambrian-Ordovician boundary interval in the western United States |journal=Memoirs of the Association of Australasian Palaeontologists |volume=42 |pages=417–436 |year=2011 |citeseerx=10.1.1.828.4661 }}

|

Gen. et 2 sp. nov

|

Valid

|

Loch & Taylor

|

Early Ordovician

|

Bliss Formation

House Limestone

Tanyard Formation

Wilberns Formation

|

{{Flag|United States}}

|

A member of Asaphida belonging to the family Symphysurinidae. The type species is C. milleri; genus also includes C. repetskii.

|

Gladiatoria crowei{{cite journal |author1=Jonathan M. Adrain |author2=Neo E.B. McAdams |author3=Stephen R. Westrop |title=Affinities of the Lower Ordovician (Tulean; lower Floian) trilobite Gladiatoria, with species from the Great Basin, western United States |journal=Memoirs of the Association of Australasian Palaeontologists |volume=42 |pages=321–367 |year=2011 |url=http://myweb.uiowa.edu/jadrain/resources/Publications/Adrainetal2011b.pdf }}

|

Sp. nov

|

Valid

|

Adrain, McAdams & Westrop

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Gladiatoria harrisi

|

Sp. nov

|

Valid

|

Adrain, McAdams & Westrop

|

Ordovician (Floian)

|

Fillmore Limestone

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Gladiatoria nielsenae

|

Sp. nov

|

Valid

|

Adrain, McAdams & Westrop

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Gladiatoria phoenixi

|

Sp. nov

|

Valid

|

Adrain, McAdams & Westrop

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Gladiatoria reedi

|

Sp. nov

|

Valid

|

Adrain, McAdams & Westrop

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Hebediscus williamsi{{cite journal |author1=Stephen R. Westrop |author2=Ed Landing |title=Lower Cambrian (Branchian) eodiscoid trilobites from the lower Brigus formation, Avalon Peninsula, Newfoundland, Canada |journal=Memoirs of the Association of Australasian Palaeontologists |volume=42 |pages=209–262 |year=2011 |url=https://www.researchgate.net/publication/285923830 }}

|

Sp. nov

|

Valid

|

Westrop & Landing

|

Early Cambrian

|

Brigus Formation

|

{{Flag|Canada}}

|

A member of the family Hebediscidae.

|

Hintzeia parafirmimarginis{{cite journal |author1=Neo E.B. McAdams |author2=Jonathan M. Adrain |title=Systematics of the Lower Ordovician pliomerid trilobite Hintzeia, with species from the Great Basin, western USA |journal=Zootaxa |volume=2910 |pages=1–45 |year=2011 |url=http://myweb.uiowa.edu/jadrain/resources/Publications/McAdamsAdrain2011a.pdf }}

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Luhops coquus{{cite journal |author=Adrian W.A. Rushton |title=The mid-Cambrian (Drumian) centropleurid trilobite Luhops and its relatives from the Abbey Shale formation near Nuneaton, central England |journal=Memoirs of the Association of Australasian Palaeontologists |volume=42 |pages=93–104 |year=2011 }}

|

Sp. nov

|

Valid

|

Rushton

|

Cambrian (Drumian)

|

Abbey Shale

|

{{Flag|United Kingdom}}

|

|

Panisaspis{{cite journal |author1=Neo E.B. McAdams |author2=Jonathan M. Adrain |title=Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA |journal=Zootaxa |volume=2969 |pages=1–68 |year=2011 |url=http://myweb.uiowa.edu/jadrain/resources/Publications/McAdamsAdrain2011b.pdf |doi=10.11646/zootaxa.2969.1.1 }}

|

Gen. et 6 sp. et comb. nov

|

Valid

|

McAdams & Adrain

|

Early Ordovician

|

Fillmore Limestone

Garden City Formation

Yellow Hill Limestone

|

{{Flag|United States}}

|

A member of the family Pliomeridae. The type species is P. millardensis; genus also includes new species P. deltaensis, P. loganensis, P. rancherensis, P. sevierensis and P. topscityensis, as well as "Protopliomerops" quattuor Hintze (1953).

|

Protopliomerella bowlesi{{cite journal |author1=Neo E.B. McAdams |author2=Jonathan M. Adrain |title=Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA |journal=Zootaxa |volume=3144 |pages=1–113 |year=2011 |url=http://myweb.uiowa.edu/jadrain/resources/Publications/McAdamsAdrain2011c.pdf |doi=10.11646/zootaxa.3144.1.1 }}

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Protopliomerella kerouaci

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

Yellow Hill Limestone

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Protopliomerella okeeffeae

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

Yellow Hill Limestone

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Protopliomerella seegeri

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Protopliomerella stegneri

|

Sp. nov

|

Valid

|

McAdams & Adrain

|

Ordovician (Floian)

|

Fillmore Limestone

|

{{Flag|United States}}

|

A member of the family Pliomeridae.

|

Psalikilopsis newmani{{cite journal |author1=Jonathan M. Adrain |author2=Neo E.B. McAdams |author3=Stephen R. Westrop |author4=Talia S. Karim |title=Systematics and affinity of the Lower Ordovician (Tulean; lower Floian) trilobite Psalikilopsis |journal=Memoirs of the Association of Australasian Palaeontologists |volume=42 |pages=369–416 |year=2011 |url=http://myweb.uiowa.edu/jadrain/resources/Publications/Adrainetal2011a.pdf }}

|

Sp. nov

|

Valid

|

Adrain et al.

|

Ordovician (Floian)

|

Fillmore Limestone

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Psalikilopsis paracuspidicauda

|

Sp. nov

|

Valid

|

Adrain et al.

|

Ordovician (Floian)

|

Fillmore Limestone

Yellow Hill Limestone

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Psalikilopsis redfordi

|

Sp. nov

|

Valid

|

Adrain et al.

|

Ordovician (Floian)

|

Fillmore Limestone

Garden City Formation

|

{{Flag|United States}}

|

A member of the family Bathyuridae.

|

Symphysurina ethingtoni

|

Sp. nov

|

Valid

|

Loch & Taylor

|

Late Cambrian and Early Ordovician

|

House Limestone

|

{{Flag|United States}}

|

A member of Asaphida belonging to the family Symphysurinidae.

|

Symphysurina straatmannae

|

Sp. nov

|

Valid

|

Loch & Taylor

|

Late Cambrian and Early Ordovician

|

Bliss Formation

House Limestone

Wilberns Formation

|

{{Flag|United States}}

|

A member of Asaphida belonging to the family Symphysurinidae.

|

References