Ashley Formation

{{Short description|Geologic formation in South Carolina, U.S.}}{{More categories|date=December 2024}}{{Infobox rockunit

| name = Ashley Formation

| image =

| caption =

| type = Geological formation

| age = Lower Oligocene (Rupelian)

| period = Paleogene

| prilithology =

| otherlithology =

| namedfor =

| namedby =

| region = South Carolina

| country = United States

| coordinates =

| unitof =

| subunits = Gettysville Member, Runnymeade Marl Member and Givhans Ferry Member.{{Cite journal |last1=Cicimurri |first1=David J. |last2=Knight |first2=James L. |last3=Ebersole |first3=Jun A. |date=2022 |title=Early Oligocene (Rupelian) fishes (Chondrichthyes, Osteichthyes) from the Ashley Formation (Cooper Group) of South Carolina, USA |url=https://escholarship.org/uc/item/0xv324c8 |journal=PaleoBios |language=en |volume=39 |issue=1 |doi=10.5070/P939056976 |s2cid=247912932 |issn=0031-0298|doi-access=free }}

| underlies =

| overlies =

| thickness =

| extent =

| area =

| map =

| map_caption =

}}

The Ashley Formation is a geologic formation in South Carolina. It preserves fossils dating back to the Paleogene period.

Vertebrate fauna

=Mammals=

class="wikitable" align="center"
colspan="6" align="center" | Carnivorans
GenusSpeciesPresence

!Material

NotesImages
Phocidae

|Gen. et. sp. indeterminate

|

|Proximal portion of a right femur (ChM PV5713).{{Cite book |last=J. |first=Ray, Clayton Edward. Emry, Robert |url=https://www.researchgate.net/publication/267408194 |title=Cenozoic mammals of land and sea : tributes to the career of Clayton E. Ray |date=2002 |publisher=Smithsonian Institution Press |pages=179–183 |oclc=1035595001}}

| "A taxon closely comparable to the most specialized phocid, the modern genus Cystophora".

|

class="wikitable" align="center"
colspan="6" align="center" | Cetaceans
GenusSpeciesPresence

!Material

NotesImages
Agorophius

|A. pygmaeus

| Underwater cliff face from the base of the formation.{{Cite journal |last1=Godfrey |first1=Stephen J. |last2=Uhen |first2=Mark D. |last3=Osborne |first3=Jason E. |last4=Edwards |first4=Lucy E. |date=January 2016 |title=A new specimen of Agorophius pygmaeus (Agorophiidae, Odontoceti, Cetacea) from the early Oligocene Ashley Formation of South Carolina, USA |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/new-specimen-of-agorophius-pygmaeus-agorophiidae-odontoceti-cetacea-from-the-early-oligocene-ashley-formation-of-south-carolina-usa/C7BF5C1676EB9FE7CF0A26033AD416BD |journal=Journal of Paleontology |language=en |volume=90 |issue=1 |pages=154–169 |doi=10.1017/jpa.2016.4 |bibcode=2016JPal...90..154G |s2cid=132593720 |issn=0022-3360|url-access=subscription }}

|A skull.

|

|File:Agorophius pygmaeus.jpg

Albertocetus

|A. meffordorum

|Givhans Ferry Member and Runnymede Marl Member.{{Cite journal |last1=Boessenecker |first1=Robert W. |last2=Ahmed |first2=Erum |last3=Geisler |first3=Jonathan H. |date=2017-11-08 |title=New records of the dolphin Albertocetus meffordorum (Odontoceti: Xenorophidae) from the lower Oligocene of South Carolina: Encephalization, sensory anatomy, postcranial morphology, and ontogeny of early odontocetes |journal=PLOS ONE |language=en |volume=12 |issue=11 |pages=e0186476 |doi=10.1371/journal.pone.0186476 |issn=1932-6203 |pmc=5695589 |pmid=29117197|bibcode=2017PLoSO..1286476B |doi-access=free }}

|A partial skeleton and an isolated braincase.

|A xenorophid.

|File:Albertocetus reconstruction outline.jpg

Ankylorhiza

|A. tiedemani

|

|AMNH 10445 (partial rostrum) and CCNHM 220 (partial skull and associated vertebrae).{{Cite journal |last1=Boessenecker |first1=Robert W. |last2=Churchill |first2=Morgan |last3=Buchholtz |first3=Emily A. |last4=Beatty |first4=Brian L. |last5=Geisler |first5=Jonathan H. |date=2020-08-17 |title=Convergent Evolution of Swimming Adaptations in Modern Whales Revealed by a Large Macrophagous Dolphin from the Oligocene of South Carolina |journal=Current Biology |language=en |volume=30 |issue=16 |pages=3267–3273.e2 |doi=10.1016/j.cub.2020.06.012 |pmid=32649912 |s2cid=220435400 |issn=0960-9822|doi-access=free |bibcode=2020CBio...30E3267B }}

|An odontocete.

|File:Ankylorhiza tiedemani life reconstruction by PaleoGeek.png

Ashleycetus

|A. planicapitis

|Near Charleston, South Carolina.{{Cite journal |last1=Sanders |first1=Albert E. |last2=Geisler |first2=Jonathan H. |date=2015-01-02 |title=A new basal odontocete from the upper Rupelian of South Carolina, U.S.A., with contributions to the systematics of Xenorophus and Mirocetus (Mammalia, Cetacea) |url=https://doi.org/10.1080/02724634.2014.890107 |journal=Journal of Vertebrate Paleontology |volume=35 |issue=1 |pages=e890107 |doi=10.1080/02724634.2014.890107 |bibcode=2015JVPal..35E0107S |s2cid=85040115 |issn=0272-4634|url-access=subscription }}

|A partial skull.

|

|

Coronodon

|C. havensteini

|"Wando River near Highway 41 Bridge, Berkeley County, South Carolina".{{Cite journal |last1=Geisler |first1=Jonathan H. |last2=Boessenecker |first2=Robert W. |last3=Brown |first3=Mace |last4=Beatty |first4=Brian L. |date=2017-07-10 |title=The Origin of Filter Feeding in Whales |journal=Current Biology |language=en |volume=27 |issue=13 |pages=2036–2042.e2 |doi=10.1016/j.cub.2017.06.003 |pmid=28669761 |s2cid=36765725 |issn=0960-9822|doi-access=free |bibcode=2017CBio...27E2036G }}

|

|A basal mysticete.

|File:Coronodon anterolateral at Mace Brown Museum of Natural History.jpg

Ediscetus

|E. osbornei

|

|

|"Slightly outside the odontocete crown group".{{Cite journal |last1=Albright |first1=L. Barry |last2=Sanders |first2=Albert E. |last3=Geisler |first3=Jonathan H. |date=April 2019 |title=An Unexpectedly Derived Odontocete from the Ashley Formation (Upper Rupelian) of South Carolina, U.S.A. |url=https://bioone.org/journals/journal-of-vertebrate-paleontology/volume-38/issue-4/02724634.2018.1482555/An-Unexpectedly-Derived-Odontocete-from-the-Ashley-Formation-Upper-Rupelian/10.1080/02724634.2018.1482555.full |journal=Journal of Vertebrate Paleontology |volume=38 |issue=4 |pages=(1)-(15) |doi=10.1080/02724634.2018.1482555 |s2cid=92830510 |issn=0272-4634|url-access=subscription }}

|

Micromysticetus

|M. rothauseni

|

|2 braincases without rostra.{{Cite journal |last1=Sanders Albert |first1=E. |last2=Barnes Lawrence |first2=G. |date=2002-11-21 |title=Paleontology of the late Oligocene Ashley and Chandler Bridge Formations of South Carolina; 2, Micromysticetus rothauseni, a primitive cetoteriid mysticete (Mammalia, Cetacea) |url=https://eurekamag.com/research/019/667/019667420.php |journal=Smithsonian Contributions to Paleobiology |language=en |volume=93 |pages=271–293}}

|

|

Saurocetus

|S. gibbesi

|L. Agassiz. 1848. [on the species Saurocetus gibbesii]. Proceedings of the Academy of Natural Sciences of Philadelphia 4:4-5

|A cheek tooth (MCZ 8760).{{Cite journal |last=Dooley |first=Alton C. Jr|date=2003 |title=A Review of the Eastern North American Squalodontidae (Mammalia: Cetacea) |journal=Jeffersoniana |issue=11 |pages=1–26}}

|

|

Xenorophus

|X. sloanii

|

|Incomplete skull.{{Cite journal |last=Kellogg |first=Remington |date=1923 |title=Description of an apparently new toothed cetacean from South Carolina |url=https://repository.si.edu/handle/10088/23638 |journal=Smithsonian Miscellaneous Collections |volume=76 |issue=7 |pages=1–7|hdl=10088/23638 }}

|A xenorophid.

|

class="wikitable" align="center"
colspan="6" align="center" | Sirenians
GenusSpeciesPresence

!Material

NotesImages
Crenatosiren

|C. olseni

|

|{{Cite journal |last=Domning |first=Daryl P. |date=1997-06-19 |title=Fossil Sirenia of the west Atlantic and Caribbean region. VI. Crenatosiren olseni (Reinhart, 1976) |url=https://doi.org/10.1080/02724634.1997.10010984 |journal=Journal of Vertebrate Paleontology |volume=17 |issue=2 |pages=397–412 |doi=10.1080/02724634.1997.10010984 |bibcode=1997JVPal..17..397D |issn=0272-4634|url-access=subscription }}

| A dugongid also found in the Chandler Bridge and Parachucla formations.

|

Stegosiren

|S. macei

|

|{{Cite journal |last1=Domning |first1=Daryl P. |last2=Beatty |first2=Brian L. |date=2019-05-04 |title=Fossil Sirenia of the West Atlantic and Caribbean region. XII. Stegosiren macei, gen. et sp. nov. |url=https://doi.org/10.1080/02724634.2019.1650369 |journal=Journal of Vertebrate Paleontology |volume=39 |issue=3 |pages=e1650369 |doi=10.1080/02724634.2019.1650369 |bibcode=2019JVPal..39E0369D |s2cid=203407242 |issn=0272-4634|url-access=subscription }}

|A dugongid also found in the Chandler Bridge Formation.

|File:Stegosiren skeleton.jpg

=Reptiles=

{{Paleobiota-key-compact}}

class="wikitable" align="center"
colspan="6" align="center" | Crocodilians
GenusSpeciesPresence

!Material

NotesImages
style="background:#f3e9f3;" |Gavialosuchus

|style="background:#fbdddb;" |G. carolinensis

|style="background:#fbdddb;" |Dorchester County, South Carolina.{{Cite book |last=R.. |first=Erickson, Bruce |url=https://www.smm.org/sites/default/files/public/attachments/estuarine-crocodile-gavialosuchus-carolinensis.pdf |title=The estuarine crocodile Gavialosuchus Carolinensis n.sp. (Crocodylia : Eusuchia) from the late Oligocene of South Carolina, North America |date=1996 |publisher=Science Museum of Minnesota |oclc=491733633}}

|style="background:#fbdddb;" |ChM PV 4282 (mandible).

|style="background:#fbdddb;" |Now moved to the genus Thecachampsa.

|style="background:#fbdddb;" |

Thecachampsa

|T. carolinensis

|Dorchester County, South Carolina.

|ChM PV 4282 (mandible).

|A gavialoid.

|File:Thecachampsa carolinensis.jpg

class="wikitable" align="center"
colspan="6" align="center" | Turtles
GenusSpeciesPresence

!Material

NotesImages
Ashleychelys

|A. palmeri

| East bank of Limehouse Branch, Berkeley County, South Carolina.{{Cite journal |last1=Weems |first1=Robert E. |last2=Sanders |first2=Albert E. |date=January 2014 |title=Oligocene pancheloniid sea turtles from the vicinity of Charleston, South Carolina, U.S.A. |url=https://www.researchgate.net/publication/263409112 |journal=Journal of Vertebrate Paleontology |volume=34 |issue=1 |pages=80–99 |doi=10.1080/02724634.2013.792826 |bibcode=2014JVPal..34...80W |s2cid=128543145 |issn=0272-4634}}

|

|A cheloniid also found in the Chandler Bridge Formation.

|

Carolinochelys

|C. wilsoni

|

|MCZ 1005-A (a nearly complete skull) and ChM PV8309.

|A cheloniid also found in the Chandler Bridge Formation.

|File:Carolinachelys wilsoni skull and shell.jpg

rowspan=2|Natemys

|N. sp. 1

|Givhan's Ferry Member.{{Cite web |title=Multispecies leatherback turtle assemblage from the Oligocene Chandler Bridge and Ashley formations of South Carolina, USA - Acta Palaeontologica Polonica |url=https://www.app.pan.pl/article/item/app007402020.html |access-date=2022-09-11 |website=www.app.pan.pl |language=en}}

|CCNHM 4288 (a non-ridged carapacial ossicle).

|A dermochelyid also found in the Chandler Bridge Formation.

|

N. sp. 2

|?Givhan's Ferry Member and ?Runnymede Marl Member.

|CCNHM 4287.1 and 4287.2 (a pair of associated non-ridged carapacial ossicles) and CCNHM

4910 (a non-ridged ossicle).

|A dermochelyid.

|

Procolpochelys

|P. charlestonensis

|

|A humerus (MCZ 1005-B).

|A cheloniid also found in the Chandler Bridge Formation.

|File:Procolpochelys charlestonensis skull and shell.jpg

cf. Psephophorus

|cf. P. sp.

|Givhan's Ferry Member.

|CCNHM 5460 (an isolated non-ridged carapacial ossicle).

|A dermochelyid also found in the Chandler Bridge Formation.

|File:Psephophorus sp. LACM.jpg

=Fish=

==Cartilaginous fish==

class="wikitable" align="center"
colspan="6" align="center" | Rays
GenusSpeciesPresence

!Material

NotesImages
Dasyatis

|"D." sp.

|Givhans Ferry Member.{{Cite journal |last1=Cicimurri |first1=David J. |last2=Knight |first2=James L. |last3=Ebersole |first3=Jun A. |date=2022 |title=Early Oligocene (Rupelian) fishes (Chondrichthyes, Osteichthyes) from the Ashley Formation (Cooper Group) of South Carolina, USA |url=https://escholarship.org/uc/item/0xv324c8 |journal=PaleoBios |language=en |volume=39 |issue=1 |doi=10.5070/P939056976 |s2cid=247912932 |issn=0031-0298|doi-access=free }}

|Multiple teeth.

|A stingray.

|File:Common stingray tynemouth.jpg

Mobula

|"M." sp.

|Givhans Ferry Member.

|

|A devil ray.

|File:Mobula breaching.jpg

Plinthicus

|P. sp.

|Givhans Ferry Member.

|A tooth (SC2007.36.48).

|

|

rowspan="2" |Raja

|R. mccollumi

| Givhans Ferry Member.

|Abundant teeth.

| A skate also found in the Chandler Bridge Formation.

|

"R." sp.

|Givhans Ferry Member.

|Multiple teeth.

|A skate, appears to be conspecific with R. sp. from the Chandler Bridge Formation.

|

Rhinoptera

|"R." sp.

|Givhans Ferry Member.

|A broken symphyseal tooth (SC2015.29.30).

|A cownose ray.

|File:Rhinoptera steindachneri.jpg

Rhynchobatus

|R. sp.

|Givhans Ferry Member.

|Teeth (SC2007.36.39, SC2015.29.13, SC2015.29.32).

|A wedgefish.

|File:Rhynchobatus djiddensis durban.jpg

Taeniurops

|"T." cavernosus

|Givhans Ferry Member.

|Multiple teeth.

|A stingray.

|

class="wikitable" align="center"
colspan="6" align="center" | Sharks
GenusSpeciesPresence

!Material

NotesImages
Araloselachus

|A. sp.

|May be derived from the Givhans Ferry Member.

|Teeth.

|

|

Carcharhinus

|C. gibbesi

|Givhans Ferry Member.

|A large number of teeth.

|The most common shark in the Ashley Formation sample.

|File:Carcharhinus gibbesi - Chandler Bridge Formation - USA.jpg

Galeocerdo

|G. aduncus

|Givhans Ferry Member.

|Teeth.

|A ground shark.

|File:Galeocerdo aduncus 323.jpg

Ginglymostomatidae

|Gen. et. sp. indeterminate

| Givhans Ferry Member.

|A partial tooth (SC2007.36.208).

| A nurse shark.

|

Hemipristis

|H. cf. H. serra

|Givhans Ferry Member.

|4 teeth (SC2007.36.7, SC2007.36.8, SC2007.36.9).

|A weasel shark.

|File:Hemipristis serra teeth.jpg

Otodus

|O. angustidens

|Ashley River, in the Givhans Ferry Member.

|Ablated specimens.

|A megatoothed shark.

|File:4-25 Angustidens.jpg

Pachyscyllium

|P. sp.

|Givhans Ferry Member.

|A tooth (SC2007.36.5).

|A catshark.

|

rowspan=2|Physogaleus

|P. cf. P. contortus

|Givhans Ferry Member.

|SC2007.36.19, SC2007.36.20 (four teeth), SC2015.29.27.

|A ground shark.

|

P. sp.

|Givhans Ferry Member.

|SC2007.36.21 and SC2007.36.22.

|A ground shark.

|

Pristiophorus

|P. sp.

|Givhans Ferry Member.

|An incomplete tooth crown (SC2015.29.20).

|A sawshark.

|File:Pristiophorus delicatus csiro-nfc.jpg

Scyliorhinus

|S. weemsi

|Givhans Ferry Member.

|Multiple teeth.

|A catshark.

|

Sphyrnidae

|Gen. et. sp. indeterminate

|Givhans Ferry Member.

|SC2007.36.23, SC2007.36.24, SC2007.36.25 (11 teeth), SC2007.36.26, SC2007.36.27, SC2007.36.28 (five teeth).

|A hammerhead shark.

|

Squalus

|S. sp.

|Givhans Ferry Member.

|Teeth.

|A spurdog.

|File:Squalus acanthias2.jpg

Squatina

|S. sp.

|Givhans Ferry Member.

|Teeth (SC2007.36.4, SC2007.36.126, SC2007.36.149, SC2007.36.227, SC2015.29.18, SC2015.29.19, SC2015.33.2).

|An angelshark.

|File:Squatina dumeril nefsc.jpg

Trigonotodus

|T. alteri

|

|Teeth.

|A thresher shark.

|File:Trigonotodus alteri lateral tooth (Oligocene, Chandler Bridge FM, SC).jpg

==Bony fish==

class="wikitable" align="center"
colspan="6" align="center" | Bony fish
GenusSpeciesPresence

!Material

NotesImages
Albula

|A. sp.

| Givhans Ferry Member.

|SC2007.36.165, SC2007.36.228 (five teeth), SC2015.29.200 (66 teeth), SC2015.29.201, SC2015.29.204.

| A bonefish.

| File:Bonefish Albula vulpes.jpg

Archosargus

|A. sp.

|Givhans Ferry Member.

|Multiple teeth.

|A sea bream.

|File:Sheepshead.jpg

?Calamus

|?C. sp.

|Givhans Ferry Member.

|Teeth.

|A sea bream.

|File:Calamus bajonado.jpg

Diplodus

|D. sp.

|Givhans Ferry Member.

|SC2015.29.195 (13 teeth), SC2015.29.222, SC2015.29.223, SC2015.29.224.

|A sea bream.

|File:Sargo real (Diplodus cervinus), franja marina Teno-Rasca, Tenerife, España, 2022-01-09, DD 49.jpg

?Labridae

|Gen. et. sp. indet.

|Givhans Ferry Member.

|Multiple teeth.

|A wrasse.

|

Osteoglossidae

|Gen. et. sp. indeterminate

|Givhans Ferry Member.

|Teeth.

|

|

Palaeocybium

|P. sp.

|Givhans Ferry Member.

|A tooth (SC2007.36.122), a neurocranium and a left dentary.

|A scombrid.

|

Paralichthyidae

|Gen. et. sp. indeterminate

|Givhans Ferry Member.

|A tooth (SC2015.36.259).

|A flatfish.

|

?Pogonias

|?P. sp.

|Givhans Ferry Member.

|Teeth.

|A croaker.

|File:Pogonias cromis (S0897) (12639311174).jpg

Scomberomorus

|S. sp.

|Givhans Ferry Member.

|Teeth.

|A scombrid.

|File:Scomberomorus cavalla.png

?Sciaenops

|?S. sp.

|Givhans Ferry Member.

|Many teeth.

|A croaker.

|File:Sciaenops ocellatus (S0230) (12527956414).jpg

Sphyraena

|S. sp.

|Givhans Ferry Member.

|Many cheek and laniary teeth.

|A barracuda.

|File:Barracuda with prey.jpg

Trichiurides

|T. cf. T. sagittidens

|Givhans Ferry Member.

|2 ablated laniary teeth.

|A cutlassfish.

|

See also

References

{{Reflist}}

Further reading

  • {{cite web|title= Fossilworks: Gateway to the Paleobiology Database|author= ((Various Contributors to the Paleobiology Database))|url= https://www.fossilworks.org|access-date= 17 December 2021}}

Category:Paleogene geology of South Carolina