Eucharideae
{{Short description|Tribe of flowering plants}}
{{Automatic taxobox
| image = Eucharis candida (14277378282).jpg
| image_caption = Urceolina candida
| taxon = Eucharideae
| authority = Hutch.Fam. Fl. Pl. 2: 130. 1934
| subdivision_ranks = Genera
| subdivision = See text
| type_genus = Eucharis (now a synonym of Urceolina)
| type_genus_authority = Planch. & Linden
| synonyms=
- Euchareae Traub
}}
Eucharideae is a tribe of plants within the family Amaryllidaceae. It was augmented in 2000 by Meerow et al. following a molecular phylogenetic study that revealed that many elements of the tribe Stenomesseae segregated with it, rather than separately, and were subsequently submerged in it (although there was an initial proposal to rename this clade Stenomesseae).{{sfn|Meerow et al.|2000b}} Further revisions were made in 2020, when three genera were merged.{{sfn|Meerow|Gardner|Nakamura|2020}} It forms one of the tribes of the Andean subclade of the American clade of the subfamily.{{sfn|Vigneron|2008}}
Taxonomy
In Traub's 1963 monograph on the Amaryllidaceae, he conceived of a tribe Euchareae based on the type genus, Eucharis and constituted from six genera in total.{{sfn|Traub|1963}} In 1996, the Müller-Doblies deemed this to be polyphyletic, and redistributed the genera over three separate tribes, retaining Eucharideae as one of those tribes, but divided into two subtribes, Eucharidinae and Hymenocallidinae with a total of seven genera.{{sfn|Müller-Doblies|Müller-Doblies|1996}} In 1998, Meerow and Snijman considered these separate tribes, retaining only four genera in their Eucharideae.{{sfn|Meerow|Snijman|1998}} In 2000, the tribe was then considerably reconstituted following a deconstruction of tribe Stenomesseae based on molecular phylogenetics, resulting in seven genera (Plagiolirion was not included). However, relationships were not fully resolved.{{sfn|Meerow et al.|2000b}}
= Phylogeny =
The placement of Eucharideae within subfamily Amaryllidoideae is shown in the following cladogram:{{sfn|Meerow et al.|2000b}}
{{cladogram
| title= {{anchor|Clad3}}Cladogram: Tribes of subfamily Amaryllidoideae
| align=left
| cladogram={{clade|style=font-size:92%;line-height:100%;width:700px;
| label1 = Subfamily Amaryllidoideae
|1= {{Clade
| label1= Africa
| 1= Tribe Amaryllideae
| 2= {{Clade
| 1= {{clade
| label1=Africa
| 1= Tribe Cyrtantheae
| 2= {{clade
| label1=Africa
| 1= Tribe Haemantheae
| label2= Australasia
| 2= Tribe Calostemmateae
}}
}}
| 2= {{clade
| label1= Eurasian clade
| label2= American clade
| 1= {{clade
| label1= Asia
| 1= Tribe Lycorideae
| label2= Mediterranean
| 2= {{clade
| 1= Tribe Galantheae
| 2= {{clade
| 1= Tribe Pancratieae
| 2= Tribe Narcisseae
}}
}}
}}
| 2= {{clade
| label1= Hippeastroid clade
| label2= Andean clade
| 1= {{clade
| 1= Tribe Griffineae
| 2= Tribe Hippeastreae
}}
| 2= {{clade
| 1= {{clade
| 1= Tribe Eustephieae
}}
| 2= {{clade
| 1= Tribe Eucharideae/Stenomesseae
| 2= {{clade
| 1= {{clade
| 1= Tribe Clinantheae
| 2= Tribe Hymenocallideae
}}
}}
}}
}}
}}
}}
}}
}}
}}
}}
{{clr}}
A 2020 molecular phylogenetic study showed that genera accepted in 2000 required further revision. A summary cladogram from the study combined Eucharis, Caliphruria and Urceolina in a single clade, as well as showing that two species were wrongly placed.{{sfn|Meerow|Gardner|Nakamura|2020}}
{{clade
|label1=Eucharideae
|1={{clade
|1=Rauhia
|2={{clade
|1=Plagiolirion (Plagiolirion horsmannii)
|2={{clade
|1={{clade
|1=Phaedranassa
|2=Eucrosia
}}
|2={{clade
|1=Stenomesson (including Caliphruria korsakoffii)
|2=Urceolina (including Eucharis, Caliphruria, Eucrosia dodsonii)
}}
}}
}}
}}
}}
= Genera =
Genera accepted in the 2020 study are:{{sfn|Meerow|Gardner|Nakamura|2020}}
- Eucrosia Ker Gawl.
- Phaedranassa Herb.
- Plagiolirion Baker
- Rauhia Traub
- Stenomesson Herb.
- Urceolina Rchb. (including Eucharis and Caliphruria)
References
{{Reflist}}
Bibliography
{{Refbegin|30em}}
- {{cite book|last1=Traub|first1=H.P.|author-link=Hamilton Traub|title=Genera of the Amaryllidaceae|url=http://babel.hathitrust.org/cgi/pt?id=mdp.39015006929270;view=1up;seq=3|archive-url=https://web.archive.org/web/20181001175229/https://babel.hathitrust.org/cgi/pt?id=mdp.39015006929270;view=1up;seq=3|url-status=dead|archive-date=2018-10-01|date=1963|publisher=American Plant Life Society|location=La Jolla, California}}
- {{cite book|last1=Meerow|first1=Alan|author-link=Alan Meerow|title=Towards a phylogeny of the Amaryllidaceae|pages=169–179}} In {{harvtxt|Rudall|Cribb|Cutler|Humphries|1995}}
- {{cite book |editor1-last=Rudall |editor1-first=P.J. |editor2-last=Cribb |editor2-first=P.J. |editor3-last=Cutler |editor3-first=D.F. |editor4-last=Humphries |editor4-first=C.J. |editor-link1=Paula Rudall |year=1995 |title=Monocotyledons: systematics and evolution (Proceedings of the International Symposium on Monocotyledons: Systematics and Evolution, Kew 1993) |publisher=Royal Botanic Gardens |location=Kew |isbn=978-0-947643-85-0 |url=http://www.press.uchicago.edu/ucp/books/book/distributed/M/bo9856357.html |access-date=14 January 2014 }}
- {{cite journal|last1=Müller-Doblies|first1=U.|last2=Müller-Doblies|first2=D.|author-link2=Dietrich Müller-Doblies|author-link1=Ute Müller-Doblies|title=Tribes and subtribes and some species combinations in Amaryllidaceae J St Hil R Dahlgren & al. 1985|journal=Feddes Repertorium|date=1996|volume=107|issue=5–6|pages=S.c.1–S.c.9}}
- {{cite book|last1=Meerow|first1=AW|title=Flowering Plants · Monocotyledons|last2=Snijman|first2=DA|author-link1=Alan Meerow|pages=83–110|doi=10.1007/978-3-662-03533-7_11|year=1998|isbn=978-3-642-08377-8|s2cid=240114483 }}, in {{harvtxt|Kubitzki|1998}}. [https://books.google.com/books?id=FyPVYzL76sMC&pg=PA85 (additional excerpts)]
- {{cite book|editor-last=Kubitzki|editor-first=K.|editor-link=Klaus Kubitzki|title=The families and genera of vascular plants. Vol.3|year=1998|publisher=Springer-Verlag|location=Berlin, Germany|isbn=978-3-540-64060-8|url=https://books.google.com/books?id=FyPVYzL76sMC|access-date=14 January 2014}}
- {{cite journal | last1 = Meerow | first1 = A.W. | last2 = Fay | first2 = M.F. | last3 = Guy | first3 = C.L. | last4 = Li | first4 = Q.-B. | last5 = Zaman | first5 = F.Q. | last6 = Chase | first6 = M.W. | author-link1=Alan Meerow|author-link2=Michael F Fay|author-link6=Mark W Chase|title = Systematics of Amaryllidaceae based on cladistic analysis of plastid rbcL and trnL-F sequence data | journal = Am. J. Bot. |year = 1999 | volume = 86 | issue = 9 | pages = 1325–1345 | doi = 10.2307/2656780|ref={{harvid|Meerow et al.|1999}} | pmid=10487820 | jstor=2656780}}
- {{cite journal | last1 = Meerow | first1 = A.W. | last2 = Guy | first2 = C.L. | last3 = Li | first3 = Q.-B. | last4 = Yang | first4 = S.-L. | author-link1 = Alan Meerow | title = Phylogeny of the American Amaryllidaceae Based on nrDNA ITS Sequences | journal = Systematic Botany | year = 2000 | volume = 25 | issue = 4 | pages = 708–726 | doi = 10.2307/2666729 | url = http://www.bulbsociety.org/meerow/Meerow%20et%20al-American.pdf | access-date = 25 January 2015 | ref = {{harvid|Meerow et al.|2000b}} | archive-url = https://web.archive.org/web/20150923195144/http://www.bulbsociety.org/meerow/Meerow%20et%20al-American.pdf | archive-date = 2015-09-23 | url-status = dead | jstor = 2666729 | s2cid = 20392462 }}
- {{cite journal|last1=Meerow|first1=Alan W.|last2=Snijman|first2=Deirdre A.|author-link1=Alan Meerow|title=The never-ending story: multigene approaches to the phylogeny of Amaryllidaceae|journal=Aliso|date=2006|volume=22|pages=355–366|doi=10.5642/aliso.20062201.29 |url=http://naldc-legacy.nal.usda.gov/naldc/download.xhtml?id=2425&content=PDF|access-date=25 January 2015|doi-access=free}}
- {{cite web|last1=Vigneron|first1=Pascal|title=Amaryllidaceae|url=http://www.amaryllidaceae.org//index.htm|website=Amaryllidaceae.org|date=2008|access-date=23 October 2014|language=fr|archive-url=https://web.archive.org/web/20150104114211/http://www.amaryllidaceae.org/index.htm|archive-date=4 January 2015|url-status=dead}}
- {{citation |last=Stevens |first=P.F. |date=2001{{ndash}}2012 |title=Angiosperm Phylogeny Website: Amaryllidoideae |url=http://www.mobot.org/mobot/research/apweb/orders/asparagalesweb.htm#Amaryllidaceae }}
- {{cite web|title=Amaryllidaceae: A taxonomic tool for the Amaryllidaceae of the world|url=http://amaryllidaceae.e-monocot.org/|publisher=eMonocot|ref={{harvid|eMonocot|2016}}|access-date=2016-02-25|archive-date=2016-03-04|archive-url=https://web.archive.org/web/20160304161517/http://amaryllidaceae.e-monocot.org/|url-status=dead}}
- {{Cite journal |last1=Meerow |first1=Alan W. |last2=Gardner |first2=Elliot M. |last3=Nakamura |first3=Kyoko |date=2020 |title=Phylogenomics of the Andean Tetraploid Clade of the American Amaryllidaceae (Subfamily Amaryllidoideae): Unlocking a Polyploid Generic Radiation Abetted by Continental Geodynamics |journal=Frontiers in Plant Science |volume=11 |page=582422 |doi=10.3389/fpls.2020.582422 |pmid=33250911 |pmc=7674842 |issn=1664-462X |doi-access=free }}
{{Refend}}
{{Commons}}
{{Wikispecies}}
{{Amaryllidaceae}}
{{Taxonbar|from=Q10489715}}