Grimmen Formation
{{short description|Jurassic geologic formation in Germany}}
{{Infobox rockunit
| name = Grimmen Formation
| image = Jurassic_deposits_near_Grimmen.png
| caption = Clay pit outcrop near Grimmen
| type = Geological formation
| age = Lower Toarcian
~{{fossil range|182|179}}Tenuicostatum-Exaratum
| period = Toarcian
| prilithology = Claystone, Siltstone
| otherlithology = Mudstone, fine-grained Sandstone, Siderite and carbonate concretions
| namedfor = Grimmen, Germany
| region = Mecklenburg-Vorpommern
| country = Germany
| unitof = Kamienna Group
| underlies = Glashütte Formation, Dörnten Member (Posidonia Shale Formation)
| overlies = Posidonia Shale Formation, Wolgast Formation
| thickness = Up to {{convert|100|m|ft|abbr=on}}
| extent = North German Basin
}}
The Grimmen Formation is a Lower Jurassic (Toarcian) geological formation in northeastern Germany, primarily exposed in the Grimmen and Klein Lehmhagen clay pits and documented in wells such as Reinberg 1E. Formally established in 2025, it was previously part of the informal “Green Series” of the Ciechocinek Formation.{{Cite journal |last1=Ansorge |first1=J. |last2=Franz |first2=M. |last3=Götz |first3=A.E. |title=Stratigraphy and palaeoecology of the Toarcian in NE Germany: organo-detrital and detrital sedimentation in response to the productivity of the planktic ecosystem |journal=PalZ |date=2025 |doi=10.1007/s12542-025-00718-z|doi-access=free |bibcode=2025PalZ..tmp...18A }}{{Cite journal |last1=Ruebsam |first1=Wolfgang |last2=Franz |first2=Matthias |last3=Ansorge |first3=Jörg |last4=Obst |first4=Karsten |last5=Schwark |first5=Lorenz |date=2024-05-20 |title=Late Triassic to Early Jurassic carbon isotope chemostratigraphy and organo-facies evolution in a distal to proximal transect of the North German Basin |journal=International Journal of Earth Sciences |volume=113 |issue=8 |pages=1977–1998 |doi=10.1007/s00531-024-02418-6 |doi-access=free |bibcode=2024IJEaS.113.1977R }} It represents a prodelta to brackish-marine depositional system in the eastern North German Basin (NGB), shaped by sea-level changes and the Toarcian anoxic event.{{cite journal |last1=Ernst |first1=W. |title=Die Liastongrube Grimmen. Sediment, Makrofauna und Stratigraphie |journal=Ein Überblick Geologie |date=1967 |volume=16 |issue=1 |pages=550–569}}{{cite journal |last1=Prauss |first1=M. |title=The Lower Toarcian Posidonia Shale of Grimmen, Northeast Germany |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |date=1996 |volume=434 |issue=1 |pages=107–132 |doi=10.1127/njgpa/200/1996/107 |url=https://www.researchgate.net/publication/285862272 |access-date=30 July 2021}}{{clear}}
History
Studies of Toarcian strata near Grimmen began in 1874 with the discovery of fossiliferous clays in a railway cutting near Schönenwalde, 5 km north of Grimmen, initially misidentified as Middle Jurassic due to ammonite finds.{{cite journal |last1=Stolley |first1=E. |title=Über den oberen Lias und den unteren Dogger Norddeutschlands |journal=Neues Jahrbuch für Mineralogie, Geologie und Paläontologie |date=1909 |volume=28 |issue=1 |pages=286–334}} In 1909, the succession was reclassified as Lower Toarcian (Lias epsilon), distinct from the Posidonia Shale Formation. From 1959 to 1995, the Grimmen clay pit was excavated, revealing a glacially dislocated raft of Liassic clays and sands deformed by Pleistocene ice advances.{{cite journal |last1=Ansorge |first1=J. |title=Lower Jurassic clay pit of Klein Lehmhagen near Grimmen. The Central European Basin System–from the Bottom to the Top |journal=Geo-Pomerania, Szczechin |date=2007 |volume=23 |issue=1 |pages=37–41 |url=https://www.researchgate.net/publication/285166668 |access-date=8 September 2021}}{{cite journal |last1=Ernst |first1=W. |title=Der Lias der Scholle von Dobbertin (Mecklenburg) |journal=Fundgrube |date=1992 |volume=28 |issue=1 |pages=56–70}} Exploration wells (e.g., Reinberg 1E, Kb Barth 10, Kb Grambow 5) since the 1950s provided extensive core data, with Reinberg 1E designated as the reference section in 2025. The clay pit, abandoned and water-filled since 1995, was established as the type section in 2025, with 2016 and 2020 excavations refining an 18 m composite log spanning the Pliensbachian-Toarcian boundary to the elegans Subzone.{{cite journal |last1=Stumpf |first1=Sebastian |last2=Ansorge |first2=Jörg |last3=Grimmberger |first3=Gunther |title=Grätensandsteine und andere Geschiebe des oberen Lias (Toarcium) aus Norddeutschland |journal=Geschiebekunde Aktuell |date=2016 |volume=32 |issue=4 |pages=121–141 |url=https://www.researchgate.net/publication/312212775 |access-date=30 July 2021}}
Sedimentology/Lithology
File:Grimmen Clay Pit interpretation.png
The Grimmen Formation comprises greenish to bluish claystones, siltstones, and fine-grained sandstones, with siderite and carbonate concretions, reaching up to 100 m thick in the eastern North German Basin. The type section in the Grimmen clay pit forms an 18 m composite log, transitioning from Pliensbachian sandy deposits of the Wolgast Formation to Toarcian organo-detrital clays and heteroliths of the Lehmhagen Member (Posidonia Shale Formation), followed by the Reinberg Member of the Grimmen Formation. The Reinberg Member (~10 m thick) starts with bluish clays, grading to greenish, pellet-laminated clays with exaratum and lower elegans marker beds (marly limestones with fecal pellets).{{cite journal |last1=Leonowicz |first1=P. |title=The Ciechocinek Formation (Lower Jurassic) of SW Poland: petrology of green clastic rocks |journal=Geological Quarterly |date=2005 |volume=49 |issue=3 |pages=317–330 |url=https://www.researchgate.net/publication/228645104 |access-date=21 December 2021}} Pellet laminae, up to 5 mm thick, decrease upsection, while silty/sandy intercalations and siderite concretions increase, forming a coarsening-upward trend with pyrite-filled burrows. The upper Grimmen Formation features greenish clays with thin silt/sand laminae in 8–15 m symmetric cycles, capped by the Glashütte Formation’s deltaic sandstones. Heavy minerals (zircon, rutile, tourmaline) and smectite-group clay minerals indicate volcanogenic input, while kaolinite and chlorite are common.{{cite journal |last1=Brański |first1=P. |title=Kaolinite peaks in early Toarcian profiles from the Polish Basin–an inferred record of global warming |journal=Geological Quarterly |date=2010 |volume=54 |issue=1 |pages=15–24 |url=https://www.researchgate.net/publication/296794079 |access-date=21 December 2021}} Glacially dislocated Eocene greensand and clay are locally intercalated. Organic-rich layers contain charcoal fragments, suggesting wildfires.
Toarcian material found in glacial Erratics in Ahrensburg and the Hagen Forest have been in controversy due to its dubious origin, being linked with the Rya Formation and Sorthat Formation, as well this unit. They were originally considered or local or Baltic in derivation, but that changued with the recovery of erratic concretions in the Baltic sea cliffs near Lübeck, being found as part of the Weichselian Glacial Maximum. Liassic–Cretaceous sediments in the assemblage are most probably associated with the tectonic Sorgenfrei-Tornquist Zone. The origin of this erratics from southwestern Baltic, Poland or Danish archipelago is unlikely, as those zones are dominated by Late Cretaceous–Paleocene strata, suggesting that this Toarcian assamblages should come from south/SW between STZ–TTZ and the German Baltic coast. The most clear hint link this deposits with the Grimmen Fm, as they're identical in fauna and facies composition of Grimmen and Dobbertin, also affected by subglacial erosion and thrusting, suggesting a close stratigraphic and palaeogeographical origin.
Paleoenvironment
File:Jurassic NGB plankton.png
File:Grimmen Formation distribution.png
The Grimmen Formation was deposited in a prodelta to brackish-marine environment in the eastern North German Basin, influenced by sea-level changes and the Toarcian anoxic event. The Reinberg Member reflects a transition from organo-detrital Posidonia Shale Formation to brackish-marine clays, with high total organic carbon (TOC, ~3 wt%) at the base decreasing to <1 wt% upsection, indicating a shift to dysoxic conditions. Basal bluish clays (exaratum Subzone) represent a restricted marine basin, followed by greenish clays with pellet laminae, suggesting increased fluvial input and suspension-load plumes during wetter phases. The coarsening-upward trend and onset of bioturbation (pyrite-filled burrows) reflect prodelta progradation from Fennoscandia, culminating in the Glashütte Formation’s deltaic systems. High kaolinite and smectite content indicate a warm, humid climate with biochemical weathering and volcanogenic influence, akin to modern tropical settings. The absence of major ripples suggests minimal wave action, with sedimentation driven by fluvial input and storm events. The formation extends from western Poland to the Ringkøbing-Fyn-Møn-Arkona High, interfingering with the Ciechocinek Formation in the east. The Grimmen Formation’s biostratigraphy is primarily based on ammonite zones, with the falciferum Zone (elegantulum, exaratum, elegans, falciferum subzones) for the lower Toarcian. The type section spans the Pliensbachian-Toarcian boundary (tenuicostatum Subzone) to the elegans Subzone, with key fossils preserved in concretions and clays.File:Grimmen fossil invertebrates.pngAmmonites define subzones, with Dactylioceras semicelatum and possible D. tenuicostatum at the boundary, followed by Lobolytoceras siemensi and Tiltoniceras antiquum in the Lehmhagen Member (semicelatum Subzone). Elegantulum concretions yield abundant Eleganticeras elegantulum and rare Hildaites murleyi, while exaratum concretions in the Reinberg Member contain Cleviceras exaratum, Phylloceras heterophyllum, and Lytoceras crenatum. Belemnites (Passaloteuthis bisulcata) are common in the semicelatum Subzone but absent from the elegantulum Subzone due to anoxia. Teuthoids and coleoids occur in elegantulum and exaratum concretions.
Holoplanktonic gastropod Coelodiscus minutus and larval ‘Inoceramus’ dubius are abundant, with adults in elegantulum and exaratum concretions, often in regurgitalites. Ostracods and foraminifera are absent in black shales but occur above the Dörnten Member. Calcareous nannoplankton (e.g., Rhombolithion) are diverse in the elegantulum Subzone, correlating with NJ5b–NJ6 zones. Palynofacies show a dinoflagellate cyst blackout in the elegantulum-exaratum Subzones, with amorphous organic matter (AOM) dominance and micro-charcoal. Megaspores (e.g., Paxillitriletes phyllicus) from Fennoscandia indicate a humid climate.
= Reinberg Member Fossil Record =
The Reinberg Member, notably at Grimmen and Dobbertin, is a Konservatlagerstätte with exceptional marine and terrestrial fossils in exaratum concretions. Ammonites (e.g., Cleviceras exaratum), Coelodiscus minutus, and ‘Inoceramus’ dubius are preserved in calcite, with predation traces. Decapod crustaceans (e.g., Palaeoastacus-like), coleoid gladii, and fish (leptolepids, Grimmenodon aureum) occur in calcium phosphate. Coprolites, likely from fish, sharks, and marine reptiles (e.g., ichthyosaurs, plesiosaurs), are common. Terrestrial insects, especially in exaratum concretions types 1 and 2, include well-preserved wings and rare 3D specimens, comparable to Dobbertin’s fauna. These fossils, formed in anoxic, fine-grained limestone below the storm wave base, reflect a nutrient-rich environment fertilized by insect carcasses.
Biota
{{Paleobiota-key-compact}}
= Foraminifera =
class="wikitable sortable"
!Genus !Species !Location !Material !Notes !Images |
style="background:#D1FFCF;" |Berthelinella{{cite journal |last1=Pietrzeniuk |first1=E. |date=1961 |title=Zur Mikrofauna einiger Liasvorkommen der Deutschen Demokratischen Republik |url=https://digital.slub-dresden.de/data/kitodo/PietZur_167933834X/PietZur_167933834X_tif/jpegs/PietZur_167933834X.pdf |journal=Freiberger Forschungshefte |volume=21 |issue=5 |pages=113–129 |access-date=4 November 2021}}
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Plectofrondiculariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Bolivina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Bolivinidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Cornuspira
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Cornuspirinae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Dentalina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Eoguttulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Polymorphininae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Falsopalmula
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Robuloididae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Frondicularia
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Glomospira
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Usbekistaniinae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Lagena
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Lenticulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Vaginulinidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Lingulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Lingulininae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Marginulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Marginulininae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Nodosaria
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Ophtalmidium
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Ophthalmidiidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Rectoglandulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Stilostomella
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Stilostomellidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Trochamminoides
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Trochamminoidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Turrispirillina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the family Spirillinidae | style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Vaginulina
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |
| style="background:#D1FFCF;" |Tests | style="background:#D1FFCF;" |Foraminiferan of the subfamily Vaginulininae | style="background:#D1FFCF;" | |
= Ichnofossils =
class = "wikitable" |
Genus
! Species ! Type ! Location ! Origin ! Images |
---|
style="background:#FEF6E4;" |
|style="background:#FEF6E4;" |
|style="background:#FEF6E4;" | Fodinichnia |style="background:#FEF6E4;" |
|style="background:#FEF6E4;" |
|style="background:#FEF6E4;" | |
style="background:#FEF6E4;" |
| style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | Pascichnia | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | |
style="background:#FEF6E4;" |
Planolites{{cite journal |last1=Leonowicz |first1=P. |title=Trace fossils from the Lower Jurassic Ciechocinek Formation, SW Poland |journal=Volumina Jurassica |date=2008 |volume=6 |issue=6 |pages=89–98 |url=https://www.academia.edu/14260648 |access-date=21 December 2021}} | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | Pascichnia | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | |
style="background:#FEF6E4;" |
| style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | Domichnia | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | |
style="background:#FEF6E4;" |
Taenidium{{cite journal |last1=Geinitz |first1=F. E. |title=Der Jura von Dobbertin in Mecklenburg und seine Versteinerungen |journal=Zeitschrift der Deutschen Geologischen Gesellschaft |date=1880 |volume=32 |issue=1–2 |pages=510–535 |url=https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142446/Geinitz-F.-E.-1880-Der-Jura-von-Dobbertin-in-Mecklenburg-und-seine-Versteinerungen.pdf |access-date=14 November 2021 |archive-date=21 December 2021 |archive-url=https://web.archive.org/web/20211221174522/https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142446/Geinitz-F.-E.-1880-Der-Jura-von-Dobbertin-in-Mecklenburg-und-seine-Versteinerungen.pdf |url-status=dead }}{{cite journal |last1=Suhr |first1=P.|title= Lebensspuren aus dem Lias von Dobbertin |journal=Fundgrube|date=1988 |volume=24 |issue=1 |pages=22–26}} | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | Fodinichnia | style="background:#FEF6E4;" |
| style="background:#FEF6E4;" | | style="background:#FEF6E4;" | |
=Brachiopoda=
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Discina{{cite book |last1=Denckmann|first1=A.|title=Ueber die geognostischen Verhältnisse der Umgegend von Dörnten nördlich Goslar: mit besonderer Berücksichtigung der Fauna des oberen Lias (Vol. 8, No. 2) |date=1887 |publisher=Schropp |location=Berlin |url=https://digital.ub.uni-potsdam.de/content/titleinfo/385711 |access-date=22 October 2021}}
|
|
|Shells | A saltwater brachiopod, member of Discinidae inside Discinida. | |
=Bivalvia=
=Gastropoda=
=Cephalopoda=
=Crustacea=
Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.
=Arachnida=
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Seppo{{cite journal |last1=Selden|first1=Paul A.|last2=Dunlop|first2=Jason A.|title=The first fossil spider (Araneae: Palpimanoidea) from the Lower Jurassic (Grimmen, Germany)|journal=Zootaxa |date=2014 |volume=3894 |issue=1 |pages=161–168 |doi=10.11646/zootaxa.3894.1.13|pmid=25544628|url=https://www.researchgate.net/publication/270292420|access-date=30 July 2021}}
|
|
| Single incomplete specimen | A spider, possible member of the superfamily Palpimanoidea. It is the first confirmed spider from the lower Jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action. Probably a ground-dwelling predator that hunted the abundant insect fauna present on the layers. | |
=Insecta=
Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation.{{cite journal |last1=Ansorge |first1=J.|title=Insects from the Lower Toarcian of Middle Europe and England |journal=Proceedings of the Second Palaeoentomological Congress, Acta Zoologica Cracoviensia|date=2003 |volume=46 |issue=1 |pages=291–310 |url=https://www.researchgate.net/publication/271444424 |access-date=30 July 2021}} In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of Eleganticeras elegantulum macrochonchs and in fish coprolites which are the most frequent fossils at all. In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex. On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.
==Notoptera==
==Odonatoptera==
===Odonata===
==Paraneoptera==
==Eoblattida==
==Thysanoptera==
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Liadoptilia{{cite journal |last1=Handlirsch |first1=A. |title=Neue Untersuchungen u ̈ber die fossilen Insektenmit Erga ̈nzungen und Nachtra ̈gen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemeine biologische Probleme. II Teil |journal=Annalen des Naturhistorischen Museums in Wien |date=1939 |volume=49 |issue=1 |pages=1–240}}
|
|
|rowspan="3" | Specimens |rowspan="3" | Thysanopterans, members of the family Lophioneurida. Aeroplankton is extraordinarily well preserved in Grimmen, with the most abundant representatives of the aeroplankton (around 3 mm) being Lophioneurids, specially Undacypha europaea. |rowspan="3" | File:Thrips5.jpg, extinct genera were probably similar]] |
Trichorthophlebia
|
|
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Undacypha
|
|
|
==Plecoptera==
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Dicronemoura
|
|
| Specimens | A stonefly, member of the family Perlariopseidae. | |
Dobbertiniopteryx{{cite journal |last1=Ansorge |first1=J. |title=Dobbertiniopteryx capniomimus gen. et sp. nov. die erste Steinfliege (Insecta: Plecoptera) aus dem europäischen Jura |journal=Paläontologische Zeitschrift |date=1993 |volume=67 |issue=3–4 |pages=287–292 |doi=10.1007/BF02990281 |s2cid=130530628 |url=https://www.researchgate.net/publication/225503210 |access-date=23 October 2021}}
|
|
| Specimens | A winter stonefly, member of the family Capniidae. | |
==Orthoptera==
==Phasmatodea==
==Blattodea==
==Hemiptera==
==Hymenoptera==
==Megaloptera==
==Neuroptera==
==Coleoptera==
==Trichoptera==
==Lepidoptera==
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Eolepidopterigidae{{cite journal |last1=Zhang|first1=Q. Q. |last2=Mey |first2=W. |last3=Ansorge |first3=J. |last4=Starkey |first4=T. A. |last5=McDonald |first5=L. T. |last6=McNamara |first6=M. E. |last7=Jarzembowski |first7=E. A. |last8=Wichard |first8=W. |last9=Kelly |first9=R. |last10=Ren |first10=X. Y. |last11=Chen |first11=J. |last12=Zhang |first12=H. C. |last13=Wang |first13=B. |title=Fossil scales illuminate the early evolution of lepidopterans and structural colors |journal=Science Advances |date=2018 |volume=4 |issue=1 |pages=e1700988 |doi=10.1126/sciadv.1700988 |pmid=29651455 |pmc=5895446 |bibcode=2018SciA....4..988Z }}
|
|
|
| A moth, member of the family Eolepidopterigidae. | |
==Mecoptera==
==Diptera==
=Echinodermata=
In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%. In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.{{cite thesis |last1= Stumpf |first1=Sebastian |title=A synoptic review of the vertebrate fauna from the "Green Series" (Toarcian) of northeastern Germany with descriptions of new taxa: A contribution to the knowledge of Early Jurassic vertebrate palaeobiodiversity patterns|type=PhD Thesis |publisher=Greifswald University |date=2017 |pages=1–47|url=https://epub.ub.uni-greifswald.de/frontdoor/deliver/index/docId/1941/file/STUMPF2017DISS.pdf|access-date=9 September 2021}}
class = "wikitable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
Crinoidea
| Indeterminate |
|Columnar ossicles |Incertae sedis | |
Holothuriida
| Indeterminate |
|Holothurian wheels |Incertae sedis | |
Sinosura
|
|
|Isolated and semiarticulated specimens | A brittle star, member of Ophioleucidae inside Ophioleucida. Identified as Ophiura longivertebralis. | |
=Vertebrates=
==Fishes==
==Amniotes==
= Flora =
class="wikitable sortable" |
Genus
! Species ! Stratigraphic position ! Material ! Notes ! Images |
---|
style="background:#D1FFCF;" | Botryococcus
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Freshwater algae |style="background:#D1FFCF;" | Indicates freshwater influx in palynofacies assemblage IV, associated with coastal or deltaic settings. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Equisetites{{Cite journal |last=Mantei |first=S. |date=2021 |title=Ein dolomitisierter Schachtelhalm (Equisetites sp.) in einem Liasgeschiebe (Toarcium) Vorpommerns |url=https://www.researchgate.net/publication/369088426 |journal=Geschiebekunde Aktuell |volume=37 |issue=4 |pages=106–113}}
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" |Isolated stems, rhizomes |style="background:#D1FFCF;" |Equisetalean stems of the family Equisetidae. Dominant in mono-to oligotypic stands near water bodies, indicative of high terrestrial input in palynofacies assemblages I and IV. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" | Erlansonisporites
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Transported from northern sources (e.g., Fennoscandia), indicative of humid climates. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" | Hughesisporites
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Likely transported via suspension load plumes, reflecting freshwater influx. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" | Minerisporites
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Associated with warm, humid conditions and high atmospheric CO2. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" |Pagiophyllum{{Cite journal |last=Salfeld |first=H. |date=1909 |title=Beiträge zur Kenntnis jurassischer Pflanzenreste aus Norddeutschland |url=https://www.schweizerbart.de/papers/palae/detail/56/66129/Beitrage_zur_Kenntnis_jurassischer_Pflanzenreste_aus_Norddeutschland_Pompeckj_J_F_Beitrage_zur_Palaontologie_und_Stratigraphie_des_nordwestdeutschen_Jura_I |journal=PALAEONTOGRAPHICA Beiträge zur Naturgeschichte der Vorzeit |volume=56 |issue=1 |pages=1–36}}{{Cite journal |last=Neuwald |first=HK. |date=2008 |title=Jura-Hölzer aus Vorpommern |journal=Fossilien |issue=1/2008 |pages=59–60}}
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" |Grimmen clay pit |style="background:#D1FFCF;" |Leaves, driftwood |style="background:#D1FFCF;" |Conifer macrofossils of the family Cheirolepidiaceae or Araucariaceae. High phytoclasts and cuticles in palynofacies assemblages I, II, and IV support conifer presence. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" | Paxillitriletes
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with the Isoetales. Transported from Fennoscandia, indicates humid climate, biostratigraphic marker (Paxillitriletes phyllicus Zone, also seen in the Ciechocinek Formation) |style="background:#D1FFCF;" |File:Isoetes_echinospora_NRCS-1.png, typical example of Isoetales. Paxillitriletes probably come from a similar or a related Plant]] |
style="background:#D1FFCF;" | Striatriletes
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Reflects freshwater influx from northern sources. |style="background:#D1FFCF;" | |
style="background:#D1FFCF;" | Trileites
|style="background:#D1FFCF;" |
|style="background:#D1FFCF;" | Grimmen clay pit |style="background:#D1FFCF;" | Megaspores |style="background:#D1FFCF;" | Affinities with lycophytes or ferns. Indicates diverse megaspore flora transported from northern sources. |style="background:#D1FFCF;" | |