Grimmen Formation

{{short description|Jurassic geologic formation in Germany}}

{{Infobox rockunit

| name = Grimmen Formation

| image = Jurassic_deposits_near_Grimmen.png

| caption = Clay pit outcrop near Grimmen

| type = Geological formation

| age = Lower Toarcian
~{{fossil range|182|179}}Tenuicostatum-Exaratum

| period = Toarcian

| prilithology = Claystone, Siltstone

| otherlithology = Mudstone, fine-grained Sandstone, Siderite and carbonate concretions

| namedfor = Grimmen, Germany

| region = Mecklenburg-Vorpommern

| country = Germany

| unitof = Kamienna Group

| underlies = Glashütte Formation, Dörnten Member (Posidonia Shale Formation)

| overlies = Posidonia Shale Formation, Wolgast Formation

| thickness = Up to {{convert|100|m|ft|abbr=on}}

| extent = North German Basin

}}

The Grimmen Formation is a Lower Jurassic (Toarcian) geological formation in northeastern Germany, primarily exposed in the Grimmen and Klein Lehmhagen clay pits and documented in wells such as Reinberg 1E. Formally established in 2025, it was previously part of the informal “Green Series” of the Ciechocinek Formation.{{Cite journal |last1=Ansorge |first1=J. |last2=Franz |first2=M. |last3=Götz |first3=A.E. |title=Stratigraphy and palaeoecology of the Toarcian in NE Germany: organo-detrital and detrital sedimentation in response to the productivity of the planktic ecosystem |journal=PalZ |date=2025 |doi=10.1007/s12542-025-00718-z|doi-access=free |bibcode=2025PalZ..tmp...18A }}{{Cite journal |last1=Ruebsam |first1=Wolfgang |last2=Franz |first2=Matthias |last3=Ansorge |first3=Jörg |last4=Obst |first4=Karsten |last5=Schwark |first5=Lorenz |date=2024-05-20 |title=Late Triassic to Early Jurassic carbon isotope chemostratigraphy and organo-facies evolution in a distal to proximal transect of the North German Basin |journal=International Journal of Earth Sciences |volume=113 |issue=8 |pages=1977–1998 |doi=10.1007/s00531-024-02418-6 |doi-access=free |bibcode=2024IJEaS.113.1977R }} It represents a prodelta to brackish-marine depositional system in the eastern North German Basin (NGB), shaped by sea-level changes and the Toarcian anoxic event.{{cite journal |last1=Ernst |first1=W. |title=Die Liastongrube Grimmen. Sediment, Makrofauna und Stratigraphie |journal=Ein Überblick Geologie |date=1967 |volume=16 |issue=1 |pages=550–569}}{{cite journal |last1=Prauss |first1=M. |title=The Lower Toarcian Posidonia Shale of Grimmen, Northeast Germany |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |date=1996 |volume=434 |issue=1 |pages=107–132 |doi=10.1127/njgpa/200/1996/107 |url=https://www.researchgate.net/publication/285862272 |access-date=30 July 2021}}{{clear}}

History

Studies of Toarcian strata near Grimmen began in 1874 with the discovery of fossiliferous clays in a railway cutting near Schönenwalde, 5 km north of Grimmen, initially misidentified as Middle Jurassic due to ammonite finds.{{cite journal |last1=Stolley |first1=E. |title=Über den oberen Lias und den unteren Dogger Norddeutschlands |journal=Neues Jahrbuch für Mineralogie, Geologie und Paläontologie |date=1909 |volume=28 |issue=1 |pages=286–334}} In 1909, the succession was reclassified as Lower Toarcian (Lias epsilon), distinct from the Posidonia Shale Formation. From 1959 to 1995, the Grimmen clay pit was excavated, revealing a glacially dislocated raft of Liassic clays and sands deformed by Pleistocene ice advances.{{cite journal |last1=Ansorge |first1=J. |title=Lower Jurassic clay pit of Klein Lehmhagen near Grimmen. The Central European Basin System–from the Bottom to the Top |journal=Geo-Pomerania, Szczechin |date=2007 |volume=23 |issue=1 |pages=37–41 |url=https://www.researchgate.net/publication/285166668 |access-date=8 September 2021}}{{cite journal |last1=Ernst |first1=W. |title=Der Lias der Scholle von Dobbertin (Mecklenburg) |journal=Fundgrube |date=1992 |volume=28 |issue=1 |pages=56–70}} Exploration wells (e.g., Reinberg 1E, Kb Barth 10, Kb Grambow 5) since the 1950s provided extensive core data, with Reinberg 1E designated as the reference section in 2025. The clay pit, abandoned and water-filled since 1995, was established as the type section in 2025, with 2016 and 2020 excavations refining an 18 m composite log spanning the Pliensbachian-Toarcian boundary to the elegans Subzone.{{cite journal |last1=Stumpf |first1=Sebastian |last2=Ansorge |first2=Jörg |last3=Grimmberger |first3=Gunther |title=Grätensandsteine und andere Geschiebe des oberen Lias (Toarcium) aus Norddeutschland |journal=Geschiebekunde Aktuell |date=2016 |volume=32 |issue=4 |pages=121–141 |url=https://www.researchgate.net/publication/312212775 |access-date=30 July 2021}}

Sedimentology/Lithology

File:Grimmen Clay Pit interpretation.png

The Grimmen Formation comprises greenish to bluish claystones, siltstones, and fine-grained sandstones, with siderite and carbonate concretions, reaching up to 100 m thick in the eastern North German Basin. The type section in the Grimmen clay pit forms an 18 m composite log, transitioning from Pliensbachian sandy deposits of the Wolgast Formation to Toarcian organo-detrital clays and heteroliths of the Lehmhagen Member (Posidonia Shale Formation), followed by the Reinberg Member of the Grimmen Formation. The Reinberg Member (~10 m thick) starts with bluish clays, grading to greenish, pellet-laminated clays with exaratum and lower elegans marker beds (marly limestones with fecal pellets).{{cite journal |last1=Leonowicz |first1=P. |title=The Ciechocinek Formation (Lower Jurassic) of SW Poland: petrology of green clastic rocks |journal=Geological Quarterly |date=2005 |volume=49 |issue=3 |pages=317–330 |url=https://www.researchgate.net/publication/228645104 |access-date=21 December 2021}} Pellet laminae, up to 5 mm thick, decrease upsection, while silty/sandy intercalations and siderite concretions increase, forming a coarsening-upward trend with pyrite-filled burrows. The upper Grimmen Formation features greenish clays with thin silt/sand laminae in 8–15 m symmetric cycles, capped by the Glashütte Formation’s deltaic sandstones. Heavy minerals (zircon, rutile, tourmaline) and smectite-group clay minerals indicate volcanogenic input, while kaolinite and chlorite are common.{{cite journal |last1=Brański |first1=P. |title=Kaolinite peaks in early Toarcian profiles from the Polish Basin–an inferred record of global warming |journal=Geological Quarterly |date=2010 |volume=54 |issue=1 |pages=15–24 |url=https://www.researchgate.net/publication/296794079 |access-date=21 December 2021}} Glacially dislocated Eocene greensand and clay are locally intercalated. Organic-rich layers contain charcoal fragments, suggesting wildfires.

Toarcian material found in glacial Erratics in Ahrensburg and the Hagen Forest have been in controversy due to its dubious origin, being linked with the Rya Formation and Sorthat Formation, as well this unit. They were originally considered or local or Baltic in derivation, but that changued with the recovery of erratic concretions in the Baltic sea cliffs near Lübeck, being found as part of the Weichselian Glacial Maximum. Liassic–Cretaceous sediments in the assemblage are most probably associated with the tectonic Sorgenfrei-Tornquist Zone. The origin of this erratics from southwestern Baltic, Poland or Danish archipelago is unlikely, as those zones are dominated by Late Cretaceous–Paleocene strata, suggesting that this Toarcian assamblages should come from south/SW between STZ–TTZ and the German Baltic coast. The most clear hint link this deposits with the Grimmen Fm, as they're identical in fauna and facies composition of Grimmen and Dobbertin, also affected by subglacial erosion and thrusting, suggesting a close stratigraphic and palaeogeographical origin.

Paleoenvironment

File:Jurassic NGB plankton.png

File:Grimmen Formation distribution.png

The Grimmen Formation was deposited in a prodelta to brackish-marine environment in the eastern North German Basin, influenced by sea-level changes and the Toarcian anoxic event. The Reinberg Member reflects a transition from organo-detrital Posidonia Shale Formation to brackish-marine clays, with high total organic carbon (TOC, ~3 wt%) at the base decreasing to <1 wt% upsection, indicating a shift to dysoxic conditions. Basal bluish clays (exaratum Subzone) represent a restricted marine basin, followed by greenish clays with pellet laminae, suggesting increased fluvial input and suspension-load plumes during wetter phases. The coarsening-upward trend and onset of bioturbation (pyrite-filled burrows) reflect prodelta progradation from Fennoscandia, culminating in the Glashütte Formation’s deltaic systems. High kaolinite and smectite content indicate a warm, humid climate with biochemical weathering and volcanogenic influence, akin to modern tropical settings. The absence of major ripples suggests minimal wave action, with sedimentation driven by fluvial input and storm events. The formation extends from western Poland to the Ringkøbing-Fyn-Møn-Arkona High, interfingering with the Ciechocinek Formation in the east. The Grimmen Formation’s biostratigraphy is primarily based on ammonite zones, with the falciferum Zone (elegantulum, exaratum, elegans, falciferum subzones) for the lower Toarcian. The type section spans the Pliensbachian-Toarcian boundary (tenuicostatum Subzone) to the elegans Subzone, with key fossils preserved in concretions and clays.File:Grimmen fossil invertebrates.pngAmmonites define subzones, with Dactylioceras semicelatum and possible D. tenuicostatum at the boundary, followed by Lobolytoceras siemensi and Tiltoniceras antiquum in the Lehmhagen Member (semicelatum Subzone). Elegantulum concretions yield abundant Eleganticeras elegantulum and rare Hildaites murleyi, while exaratum concretions in the Reinberg Member contain Cleviceras exaratum, Phylloceras heterophyllum, and Lytoceras crenatum. Belemnites (Passaloteuthis bisulcata) are common in the semicelatum Subzone but absent from the elegantulum Subzone due to anoxia. Teuthoids and coleoids occur in elegantulum and exaratum concretions.

Holoplanktonic gastropod Coelodiscus minutus and larval Inoceramus’ dubius are abundant, with adults in elegantulum and exaratum concretions, often in regurgitalites. Ostracods and foraminifera are absent in black shales but occur above the Dörnten Member. Calcareous nannoplankton (e.g., Rhombolithion) are diverse in the elegantulum Subzone, correlating with NJ5b–NJ6 zones. Palynofacies show a dinoflagellate cyst blackout in the elegantulum-exaratum Subzones, with amorphous organic matter (AOM) dominance and micro-charcoal. Megaspores (e.g., Paxillitriletes phyllicus) from Fennoscandia indicate a humid climate.

= Reinberg Member Fossil Record =

The Reinberg Member, notably at Grimmen and Dobbertin, is a Konservatlagerstätte with exceptional marine and terrestrial fossils in exaratum concretions. Ammonites (e.g., Cleviceras exaratum), Coelodiscus minutus, and ‘Inoceramus’ dubius are preserved in calcite, with predation traces. Decapod crustaceans (e.g., Palaeoastacus-like), coleoid gladii, and fish (leptolepids, Grimmenodon aureum) occur in calcium phosphate. Coprolites, likely from fish, sharks, and marine reptiles (e.g., ichthyosaurs, plesiosaurs), are common. Terrestrial insects, especially in exaratum concretions types 1 and 2, include well-preserved wings and rare 3D specimens, comparable to Dobbertin’s fauna. These fossils, formed in anoxic, fine-grained limestone below the storm wave base, reflect a nutrient-rich environment fertilized by insect carcasses.

Biota

{{Paleobiota-key-compact}}

= Foraminifera =

class="wikitable sortable"

!Genus

!Species

!Location

!Material

!Notes

!Images

style="background:#D1FFCF;" |Berthelinella{{cite journal |last1=Pietrzeniuk |first1=E. |date=1961 |title=Zur Mikrofauna einiger Liasvorkommen der Deutschen Demokratischen Republik |url=https://digital.slub-dresden.de/data/kitodo/PietZur_167933834X/PietZur_167933834X_tif/jpegs/PietZur_167933834X.pdf |journal=Freiberger Forschungshefte |volume=21 |issue=5 |pages=113–129 |access-date=4 November 2021}}

| style="background:#D1FFCF;" |

  • B. paradoxa

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Plectofrondiculariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Bolivina

| style="background:#D1FFCF;" |

  • B. liasica

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Bolivinidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Cornuspira

| style="background:#D1FFCF;" |

  • C. orbicula

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Cornuspirinae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Dentalina

| style="background:#D1FFCF;" |

  • D. deslongchampsi
  • D. primaeva
  • D. pseudocommunis
  • D. submucronata
  • D. subtenuicollis
  • D. tenuistriata
  • D. torta
  • D. vetusta

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Eoguttulina

| style="background:#D1FFCF;" |

  • E. bilocularis
  • E. liassica
  • E. cf.simplex

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Polymorphininae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Falsopalmula

| style="background:#D1FFCF;" |

  • F. insignis

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Robuloididae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Frondicularia

| style="background:#D1FFCF;" |

  • F. bicostata
  • F. sulcata
  • F. terquemi
  • F. sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Glomospira

| style="background:#D1FFCF;" |

  • G. gordialis
  • G. sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Usbekistaniinae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Lagena

| style="background:#D1FFCF;" |

  • L. globosa
  • L. oxystoma
  • L. globosa
  • L. tenuicostata
  • L. sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Lenticulina

| style="background:#D1FFCF;" |

  • L. acutiangulata
  • L. müensteri
  • L. varians
  • L. sublaevis

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Vaginulinidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Lingulina

| style="background:#D1FFCF;" |

  • L. tenera

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Lingulininae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Marginulina

| style="background:#D1FFCF;" |

  • M. prima
  • M. simplex
  • M. oolithica

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Marginulininae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Nodosaria

| style="background:#D1FFCF;" |

  • N. dispar
  • N. agglutinans
  • N. elongata
  • N. germanica
  • N. hirsuta
  • N. dispar
  • N. mitis
  • N. regularis
  • N. simplex

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Ophtalmidium

| style="background:#D1FFCF;" |

  • O. orbiculare
  • O. carinatum
  • O. sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Ophthalmidiidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Rectoglandulina

| style="background:#D1FFCF;" |

  • R. oviformis
  • R. sexcostata

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Nodosariidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Stilostomella

| style="background:#D1FFCF;" |

  • S.? sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Stilostomellidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Trochamminoides

| style="background:#D1FFCF;" |

  • T. sp.

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Trochamminoidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Turrispirillina

| style="background:#D1FFCF;" |

  • T. conoidea

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the family Spirillinidae

| style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Vaginulina

| style="background:#D1FFCF;" |

  • V. anomala

| style="background:#D1FFCF;" |

  • Dobbertin

| style="background:#D1FFCF;" |Tests

| style="background:#D1FFCF;" |Foraminiferan of the subfamily Vaginulininae

| style="background:#D1FFCF;" |

= Ichnofossils =

class = "wikitable"
Genus

! Species

! Type

! Location

! Origin

! Images

style="background:#FEF6E4;" |

Chondrites

|style="background:#FEF6E4;" |

  • C. bollensis

|style="background:#FEF6E4;" |

Fodinichnia

|style="background:#FEF6E4;" |

  • Dobbertin Clay Pit

|style="background:#FEF6E4;" |

  • Sipunculans
  • Polychaetes
  • Enteropneustans
  • Echiurans

|style="background:#FEF6E4;" |

File:Chondrites.JPG

style="background:#FEF6E4;" |

Megagrapton

| style="background:#FEF6E4;" |

  • M. isp.

| style="background:#FEF6E4;" |

Pascichnia

| style="background:#FEF6E4;" |

  • Dobbertin Clay Pit

| style="background:#FEF6E4;" |

  • Polychaetes
  • Sipunculans
  • Enteropneustans
  • Echiurans

| style="background:#FEF6E4;" |

style="background:#FEF6E4;" |

Planolites{{cite journal |last1=Leonowicz |first1=P. |title=Trace fossils from the Lower Jurassic Ciechocinek Formation, SW Poland |journal=Volumina Jurassica |date=2008 |volume=6 |issue=6 |pages=89–98 |url=https://www.academia.edu/14260648 |access-date=21 December 2021}}

| style="background:#FEF6E4;" |

  • P. montanus
  • P. beverleyensis

| style="background:#FEF6E4;" |

Pascichnia

| style="background:#FEF6E4;" |

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

| style="background:#FEF6E4;" |

  • Polychaetes
  • Insects

| style="background:#FEF6E4;" |

File:Planolites.jpg

style="background:#FEF6E4;" |

Skolithos

| style="background:#FEF6E4;" |

  • S. isp.

| style="background:#FEF6E4;" |

Domichnia

| style="background:#FEF6E4;" |

  • Müssentin gravel pit

| style="background:#FEF6E4;" |

  • Polychaetes
  • Phoronidans

| style="background:#FEF6E4;" |

style="background:#FEF6E4;" |

Taenidium{{cite journal |last1=Geinitz |first1=F. E. |title=Der Jura von Dobbertin in Mecklenburg und seine Versteinerungen |journal=Zeitschrift der Deutschen Geologischen Gesellschaft |date=1880 |volume=32 |issue=1–2 |pages=510–535 |url=https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142446/Geinitz-F.-E.-1880-Der-Jura-von-Dobbertin-in-Mecklenburg-und-seine-Versteinerungen.pdf |access-date=14 November 2021 |archive-date=21 December 2021 |archive-url=https://web.archive.org/web/20211221174522/https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142446/Geinitz-F.-E.-1880-Der-Jura-von-Dobbertin-in-Mecklenburg-und-seine-Versteinerungen.pdf |url-status=dead }}{{cite journal |last1=Suhr |first1=P.|title= Lebensspuren aus dem Lias von Dobbertin |journal=Fundgrube|date=1988 |volume=24 |issue=1 |pages=22–26}}

| style="background:#FEF6E4;" |

  • T. serpentinum

| style="background:#FEF6E4;" |

Fodinichnia

| style="background:#FEF6E4;" |

  • Dobbertin Clay Pit

| style="background:#FEF6E4;" |

| style="background:#FEF6E4;" |

=Brachiopoda=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Discina{{cite book |last1=Denckmann|first1=A.|title=Ueber die geognostischen Verhältnisse der Umgegend von Dörnten nördlich Goslar: mit besonderer Berücksichtigung der Fauna des oberen Lias (Vol. 8, No. 2) |date=1887 |publisher=Schropp |location=Berlin |url=https://digital.ub.uni-potsdam.de/content/titleinfo/385711 |access-date=22 October 2021}}

|

  • D. cornucopiae
  • D. papyracea
  • D. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|Shells

|

A saltwater brachiopod, member of Discinidae inside Discinida.

|

=Bivalvia=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Bositra

|

  • B. buchii
  • B. radiata

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Ahrensburg

|

Shells

|

A saltwater oyster, member of Posidoniidae.

|

File:Bositra buchi 45.jpg

Goniomya

|

  • G. rhombifera

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

Shells

|

A saltwater clam, member of Pholadomyidae.

|

Gresslya

|

  • G. sp.

|

  • Ahrensburg

|

Shells

|

A saltwater clam, member of Ceratomyidae.

|

Lucina

|

  • L. plana

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

Shells

|

A saltwater clam, member of Lucinidae.

|File:Lucina aurantia 01.jpg]]

Meleagrinella{{cite journal |last1=Hesselbo |first1=S.P. |last2=Pieńkowski |first2=G. |title=Stepwise atmospheric carbon-isotope excursion during the Toarcian oceanic anoxic event (Early Jurassic, Polish Basin) |journal=Earth and Planetary Science Letters |date=2011 |volume=301 |issue=1–2 |pages=365–372 |doi=10.1016/j.epsl.2010.11.021 |bibcode=2011E&PSL.301..365H |url=https://www.sciencedirect.com/science/article/pii/S0012821X10007168 |access-date=21 December 2021|url-access=subscription }}

|

  • M. substriata

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin clay pit

|

Shells

|

A saltwater scallop, member of Oxytomidae.

|

File:Meleagrinella substriata 65.jpg]]

Nucula

|

  • N. caecilia

|

  • Dobbertin

|

Shells

|

A saltwater clam, member of Nuculidae.

|

File:Nucula nucleus.jpg]]

Parainoceramya{{cite journal |last1=Geinitz |first1=F. E. |title=Ueber die Fauna des Dobbertiner Lias |journal=Zeitschrift der Deutschen Geologischen Gesellschaft Band |date=1884 |volume=36 |issue=3 |pages=566–583 |url=https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142516/Geinitz-F.-E.-1884-%C3%9Cber-die-Fauna-des-Dobbertiner-Lias.pdf |access-date=8 September 2021 |archive-date=21 December 2021 |archive-url=https://web.archive.org/web/20211221180009/https://cdn.canr.udel.edu/wp-content/uploads/sites/2/2017/03/17142516/Geinitz-F.-E.-1884-%C3%9Cber-die-Fauna-des-Dobbertiner-Lias.pdf |url-status=dead }}

|

  • P. dubia
  • P. amygdaloides
  • P. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade

|

Shells

|

A saltwater/brackish clam, member of Inoceramidae.

|

File:Inoceramus dubius - Naturhistorisches Museum, Braunschweig, Germany - DSC05246.JPG]]

Parallelodon

|

  • P. cypriniformis

|

  • Ahrensburg

|

Shells

|

A saltwater clam, member of Parallelodontidae.

|

Stalagmina{{cite journal |last1=Zessin |first1=W.|title=The Dobbertine Jura (Lias ε, Mecklenburg) and its importance for paleoentomology |journal=Virgo, Newsletter of the Entomological Association Mecklenburg |date=2010 |volume=13|issue=1 |pages=4–9|url=https://www.entomologie-mv.de/download/virgo-13/virg%2013104%20%20Lias%20von%20Dobbertin.pdf |access-date=8 September 2021}}

|

  • S. koeneni

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

Shells

|

A saltwater clam, member of Inoceramidae.

|

Steinmannia

|

  • S. bronni
  • S. radiata

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

Shells

|

A saltwater oyster, member of Posidoniidae.

|

File:Steinmannia bronni.jpg

Tancredia

|

  • T. elegans

|

  • Ahrensburg

|

Shells

|

A saltwater clam, member of Tancrediidae.

|

=Gastropoda=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Cerithium

|

  • C. armatum
  • C. roeceri

|

  • Klein Lehmhagen pit, Grimmen

|

Shells

|

A sea snail, member of Cerithiinae.

|

File:Cerithium 4.JPG]]

Coelodiscus

|

  • C. minutus

|

  • Clay pit of Dobbertin
  • Klein Lehmhagen pit, Grimmen
  • Müssentin gravel pit
  • Ahrensburg

|

Shells

|

A holoplanktonic snail, type member of the family Coelodiscidae

|

Turritella

|

  • T. sp.

|

  • Dobbertin

|

Shells

|

A sea snail, member of Turritellidae.

|

=Cephalopoda=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Acrocoelites

|

  • A. pyramidalis

|

  • Klein Lehmhagen pit, Grimmen
  • Müssentin gravel pit

|

Multiple specimens.

|

A Belemnoidea member of the family Megateuthididae

|

File:Acrocoelites pyramidalis.JPG]]

Beloteuthis

|

  • B. schubleri

|

  • Klein Lehmhagen pit, Grimmen

|

Multiple specimens

|

A Mesoteuthoidea, member of the family Beloteuthidae.

|

File:Beloteuthis (7992625722).jpg]]

Dactylioceras{{cite journal |last1=Hoffmann |first1=K.|last2=Martin |first2=G. P. |title= Die Zone desDactylioceras tenuicostatum (Toarcien, Lias) in NW-und SW-Deutschland|journal=Paläontologische Zeitschrift|date=1960 |volume=34|issue=2 |pages=103–149|doi=10.1007/BF02987046|bibcode=1960PalZ...34..103H |s2cid=129436561}}

|

  • D. athleticum
  • D. temperatum
  • D. attenuatum
  • D. tenuicostatum
  • D. semicelatum
  • D. cf. aequistriatum
  • D. eickenbergi
  • D. ernsti

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

Shells

|

An ammonite, type member of the subfamily Dactylioceratinae.

|

File:Dactylioceras NT.jpg]]

Eleganticeras{{cite journal |last1=Sachs |first1=S. |last2=Hornung |first2=J. J. |last3=Lierl |first3=H. J |last4=Kear |first4=B. P. |title=Plesiosaurian fossils from Baltic glacial erratics: evidence of Early Jurassic marine amniotes from the southwestern margin of Fennoscandia |journal=Geological Society, London, Special Publications |date=2016 |volume=434 |issue=1 |pages=149–163 |doi=10.1144/SP434.14|bibcode=2016GSLSP.434..149S |s2cid=130195351}}{{cite journal |last1=Lehmann|first1=U. |title=Stratigraphie und Ammonitenführung der Ahrensburger Glazial-Geschiebe aus dem Lias epsilon (Unt. Toarcium) |journal=Mitteilungen aus dem Geologischen Staatsinstitut in Hamburg |date=1968 |volume=37 |issue=2 |pages=41–68 |url=http://mmtk.ginras.ru/pdf/lehmann1968.pdf |access-date=29 December 2021}}

|

  • E. elegantulum
  • E. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

Shells

|

An ammonite, member of the family Hildoceratidae.

|

File:Harpoceras elegans.jpg]]

Hildaites

|

  • H. subserpentinus

|

  • Ahrensburg

|

Shells

|

An ammonite, member of the family Hildoceratidae.

|

Jeletzkyteuthis

|

  • J. coriaceus

|

  • Klein Lehmhagen pit, Grimmen

|

Two specimens

|

A Loligosepiidae Loligosepiida (Vampyromorpha).

|

Lobolytoceras

|

  • L. siemensi
  • L. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Thüringen
  • Brandenburg
  • Braunschweig
  • Boulder from Ahrensburg

|

Shells

|

An ammonite, member of the family Lytoceratidae.

|

File:Lobolytoceras siemensi low.jpg]]

Nodicoeloceras

|

  • N. crassoides

|

  • Ahrensburg

|

Shells

|

An ammonite, type member of the subfamily Dactylioceratinae.

|

Peronoceras

|

  • P. fibulatum

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

Shells

|

An ammonite, member of the subfamily Dactylioceratinae.

|

Pseudolioceras

|

  • P. lythense

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

Shells

|

An ammonite, member of the subfamily Harpoceratinae.

|

File:Harpoceras lythense.jpg]]

Phylloceras

|

  • P. heterophyllum

|

  • Ahrensburg

|

Shells

|

An ammonite, type member of the subfamily Phylloceratinae.

|

File:Phylloceras NT.jpg]]

Teudopsis{{cite journal |last1=Schlögl|first1=J.|last2=Košt'ák|first2=M.|last3=Hyžný|first3=M.|title=First record of a gladius-bearing coleoid Teudopsis bollensis Voltz (Cephalopoda, Coleoidea) in the Toarcian of the Western Carpathians (Slovakia) |journal=Paläontologische Zeitschrift |date=2015 |volume=86 |issue=4 |pages=367–375 |doi=10.1007/s12542-012-0139-z|s2cid=129370764|url=https://www.researchgate.net/publication/280831512|access-date=30 July 2021}}

|

  • T. bollensis

|

  • Klein Lehmhagen pit, Grimmen

|

Few specimens

|

A vampire squid, member of Teudopsidae.

|

File:Kalmar Teudopsis bollensis, Schulp.jpg]]

Tiltoniceras

|

  • T. antiquum
  • T. costatum
  • T. acutum
  • T. schroederi
  • T. costatum
  • T. capillatum
  • T. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

Shells

|

An ammonite, member of the family Hildoceratidae.

|

File:Tiltoniceras antiquum - Naturhistorisches Museum, Braunschweig, Germany - DSC05134.JPG]]

Whitbyiceras

|

  • W. pingue

|

  • Ahrensburg

|

Shells

|

An ammonite, member of the family Phymatoceratidae.

|

=Crustacea=

Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Bairdia

|

  • B. hilda

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the family Bairdiidae.

|

Bythocypris

|

  • B. elongata

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the family Bythocyprididae.

|

Cytherelloidea

|

Indeterminate

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the superfamily Cytherelloidea.

|

Euestheria

|

  • E. opalina
  • E. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

Valves

|

A freshwater clam shrimp (Phyllopodan) of the family Lioestheriidae. Originally identified as a bivalve of the genus Posidonia and latter a member of the genus Lioestheria.

|

Glyphea

|

  • G. liasina

|

  • Dobbertin

|

  • Isolated molts
  • Isolated chelae

|

A marine decapod, type member of the family Glypheidae.

|

File:Glyphea.png]]

Hungarella

|

  • H. adenticulata
  • H. sp.

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the family Healdiidae

|

Paracypris

|

  • P. semidisca

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the subfamily Paracypridinae.

|

Polycope

|

  • P. cincinnata
  • P. sp.

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the family Polycopidae.

|

Procytheridea

|

  • P. sp.

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of the family Protocytheridae.

|

Proeryon

|

  • P. hartmanni
  • cf.P. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

  • Isolated molts
  • Single specimen with preserved upper thorax and chelae

|

A marine decapod, member of the subfamily Proeryoninae.

|

File:Proeryon hartmanni.jpg]]

Trachycythere

|

  • T. tubulosa

|

  • Dobbertin

|

Valves

|

A marine/brackish ostracod of incertae sedis affinity

|

=Arachnida=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Seppo{{cite journal |last1=Selden|first1=Paul A.|last2=Dunlop|first2=Jason A.|title=The first fossil spider (Araneae: Palpimanoidea) from the Lower Jurassic (Grimmen, Germany)|journal=Zootaxa |date=2014 |volume=3894 |issue=1 |pages=161–168 |doi=10.11646/zootaxa.3894.1.13|pmid=25544628|url=https://www.researchgate.net/publication/270292420|access-date=30 July 2021}}

|

  • S. koponeni

|

  • Klein Lehmhagen pit, Grimmen

|

Single incomplete specimen

|

A spider, possible member of the superfamily Palpimanoidea. It is the first confirmed spider from the lower Jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action. Probably a ground-dwelling predator that hunted the abundant insect fauna present on the layers.

|

=Insecta=

Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation.{{cite journal |last1=Ansorge |first1=J.|title=Insects from the Lower Toarcian of Middle Europe and England |journal=Proceedings of the Second Palaeoentomological Congress, Acta Zoologica Cracoviensia|date=2003 |volume=46 |issue=1 |pages=291–310 |url=https://www.researchgate.net/publication/271444424 |access-date=30 July 2021}} In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of Eleganticeras elegantulum macrochonchs and in fish coprolites which are the most frequent fossils at all. In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex. On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.

==Notoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Geinitzia

|

  • G. schlieffeni
  • G. debilis
  • G. minor

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A notopteran, member of the family Geinitziidae.

|

Nele

|

  • N. jurassica

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A notopteran, member of the family Bajanzhargalanidae.

|

File:Adult ice-crawler, Grylloblatta sp. “Trout Lake Caves” (18760843636).jpg, extant ice crawler, Nele was probably similar]]

Prosepididontus{{cite journal |last1=Handlirsch |first1=A.|title=Palaeontologie. I |journal=N C. Schröder (Ed.), Handbuch der Entomologie |date=1920 |volume=3 |issue=1 |pages=117–208}}{{cite journal|last1=Ansorge |first1=J. |title=Identity of Prosepididontus calopteryx HANDLIRSCH 1920 (Insecta: Grylloblattida: Geinitziidae) |journal=Acta Geológica Hispánica |date=2000 |volume=35 |issue=1–2 |pages=19–23 |url=https://www.academia.edu/1552855 |access-date=23 October 2021}}

|

  • P. calopteryx

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A notopteran, member of Geinitziidae.

==Odonatoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Obotritagrion{{cite journal |last1=Zessin |first1=W. |title=Die Phylogenie der Protomyrmeleontidae unter einbeziehung neuer Oberliassischer funde (Odonata: Archizygoptera sens. nov.) |journal=Odonatologica |date=1991 |volume=20 |issue=1 |pages=97–126 |url=https://www.researchgate.net/publication/342314333 |access-date=23 October 2021}}

|

  • O. tenuiformum
  • O. petersi

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)

|

Specimens

|rowspan="2" |

An odonatopteran, member of the family Protomyrmeleontidae.

|rowspan="2" |

Protomyrmeleon

|

  • P. brunonis
  • P. quadriordinum

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

===Odonata===

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Anisozygopteron

|

  • A. geinitzianum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A dragonfly, member of the family Myopophlebiidae.

|

Anomothemis

|

  • A. brevistigma

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="2" |

incertae sedis

|rowspan="2" |

Archithemis

|

  • A. brodiei

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Dialothemis

|

  • D. dubia

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A dragonfly, member of the family Selenothemistidae.

|

Eosagrion

|

  • E. risi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A dragonfly, member of the family Eosagrionidae.

|

Grimmenopteron

|

  • G. elegantulum

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A dragonfly, member of the family Liassophlebiidae.

|

Heterophlebia

|

  • H. buckmani
  • H. megapolitana

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)

|

Specimens

|

A dragonfly, member of the family Heterophlebiidae.

|

Heterothemis{{cite journal |last1=Handlirsch |first1=A. |title=Palaeontologie |journal=In C. Schröder (Ed.), Handbuch der Entomologie |date=1921 |volume=3 |issue=1 |pages=209–304}}{{cite journal |last1=Ansorge |first1=J. |title=Insekten aus Liasgeoden der Ahrensburger Geschiebesippe |trans-title=Insects from the Lias nodules of the Ahrensburg geschiebe assemblage |journal=Archiv für Geschiebekunde |date=2004 |volume=3 |issue=1 |page=8 |url=https://www.researchgate.net/publication/271442546 |access-date=16 July 2021 |language=de |via=ResearchGate}}

|

  • H. brodiei
  • H. germanica

|

  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg erratics

|

Specimens

|

A dragonfly, member of the family Liassogomphidae.

|

Liadothemis

|

  • L. hydrodictyon
  • L. major
  • L. geinitzi
  • L. insignis

|

  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="6" |

Incertae sedis. Magnasupplephlebia represents a very large dragonfly, with a wingspan of 13 cm. Other odonatan remains are unable to being referred to a concrete group due to their incomplete status.

|rowspan="6" |

Magnasupplephlebia{{cite journal |last1=Zessin |first1=W. |title=Durchsicht einiger liassischer Odonatopteroida unter Berücksichtigung neuer Funde von Dobbertin in Mecklenburg |journal=Deutsche Entomologische Zeitschrift |date=1982 |volume=29 |issue=1 |pages=101–106|doi=10.1002/mmnd.19820290117 }}{{cite journal |last1=Zessin |first1=W. |last2=Ansorge |first2=J. |title=Magnasupplephlebia intercalaria n. sp.-eine neue Anisozygopterenart aus dem oberen Lias von Mitteleuropa (Insecta, Odonata) |journal=Deutsche Entomologische Zeitschrift |date=1987 |volume=34 |issue=4–5 |pages=383–386 |doi=10.1002/mmnd.19870340416 |url=https://www.researchgate.net/publication/312502315 |access-date=23 October 2021}}

|

  • M. kallweita
  • M. intercalaria

|

  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

|Specimens

Parelthothemis

|

  • P. dobbertinensis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Petrothemis

|

  • P. singularis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Pycnothemis

|

  • P. densa

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Rhabdothemis

|

  • R. strigivena

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Selenothemis

|

  • S. liadis

|

  • Dobbertin, Mecklenburg (NHMW collection)

|

Specimens

|

A dragonfly, type member of the family Selenothemistidae.

|

Temnostigma

|

  • T. singulare

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Incertae sedis.

|

Trigonophlebia

|

  • T. zessini

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A dragonfly, member of the family Heterophlebiidae.

|

Turanopteron

|

  • T. pommerana

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A dragonfly, member of the family Asiopteridae. Sphenophlebia pommerana is a junior synonym.

|

==Paraneoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Archipsylla{{cite journal |last1=Handlirsch |first1=A. |title=Die Fossilen Insekten und die Phylogenie der Rezenten Formen, parts I-IV |journal=Ein Handbuch für Paläontologen und Zoologen |date=1906 |volume=12 |issue=1 |pages=1–640}}

|

  • A. primitiva

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A paraneopteran, member of the family Archipsyllidae.

|

Liassopsocus{{cite journal |last1=Ansorge |first1=J. |title=Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland) |journal=Neue Paläontologische Abhandlungen |date=1996 |volume=2 |issue=1 |pages=1–132 |url=https://www.researchgate.net/publication/284027049 |access-date=23 October 2021}}

|

  • L. lanceolatus

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A paraneopteran, member of the family Psocidiidae.

|

==Eoblattida==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Griphopteron

|

  • G. molle

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

An Eoblattidan, member of the family Blattogryllidae.

|

==Thysanoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Liadoptilia{{cite journal |last1=Handlirsch |first1=A. |title=Neue Untersuchungen u ̈ber die fossilen Insektenmit Erga ̈nzungen und Nachtra ̈gen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemeine biologische Probleme. II Teil |journal=Annalen des Naturhistorischen Museums in Wien |date=1939 |volume=49 |issue=1 |pages=1–240}}

|

  • L. misera

|

  • Klein Lehmhagen pit, Grimmen

|rowspan="3" |

Specimens

|rowspan="3" |

Thysanopterans, members of the family Lophioneurida. Aeroplankton is extraordinarily well preserved in Grimmen, with the most abundant representatives of the aeroplankton (around 3 mm) being Lophioneurids, specially Undacypha europaea.

|rowspan="3" |

File:Thrips5.jpg, extinct genera were probably similar]]

Trichorthophlebia

|

  • T. pilifera

|

  • Dobbertin, Mecklenburg (Greifswald collection)
Undacypha

|

  • U. europaea

|

  • Dobbertin, Mecklenburg (Greifswald collection)

==Plecoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Dicronemoura

|

  • D. furcata

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A stonefly, member of the family Perlariopseidae.

|

Dobbertiniopteryx{{cite journal |last1=Ansorge |first1=J. |title=Dobbertiniopteryx capniomimus gen. et sp. nov. die erste Steinfliege (Insecta: Plecoptera) aus dem europäischen Jura |journal=Paläontologische Zeitschrift |date=1993 |volume=67 |issue=3–4 |pages=287–292 |doi=10.1007/BF02990281 |s2cid=130530628 |url=https://www.researchgate.net/publication/225503210 |access-date=23 October 2021}}

|

  • D. capniomimus

|

  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A winter stonefly, member of the family Capniidae.

|

File:Small Winter Stonefly (Capniidae) (16475370865).jpg

==Orthoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Locustopsis{{cite journal |last1=Zessin |first1=W. |title=Revision der mesozoischen Familie Locustopsidae unter Berücksichtigung neuer Funde (Orthopteroida, Caelifera) |journal=Deutsche Entomologische Zeitschrift |date=1983 |volume=30 |issue=2 |pages=173–237|doi=10.1002/mmnd.19830300115 }}{{cite journal |last1=Zessin |first1=W. |title=Locustopsis kruegeri n. sp. (Orthopteroida, Caelifera) aus dem oberen Lias von Schandelah bei Braunschweig (BRD). |journal=Zeitschrift für Geologische Wissenschaften |date=1983 |volume=11 |issue=1–4 |pages=905–910}}

|

  • L. gyra
  • L. bernstorffi
  • L. sippeli
  • L. dubia
  • L. bernstorffi
  • L. nana
  • L. reducta
  • L. elongata
  • L. elegans
  • L. pulchella
  • L. procera
  • L. mecklenburgica
  • L. maculosa
  • L. sp 1
  • L. sp 2
  • L. sp 3
  • L. sp 4

|

  • Grimmen (Krempien collection)
  • Lehmhagen, near Grimmen (Zessin collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Grasshoppers of the family Locustopsidae.

|

Locustophanes

|

  • L. rhipidophorus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A locust, type member of the family Locustidae.

|

File:Garden locust (Acanthacris ruficornis).jpg, extant Locust, Locustophanes was probably similar]]

Panorpidium{{cite journal |last1=Zessin |first1=W. |title=Variabilität, Merkmalswandel und Phylogenie der Elcanidae im Jungpaläozoikum und Mesozoikum und die Phylogenie der Ensifera |journal=Deutsche Entomologische Zeitschrift |date=1987 |volume=34 |issue=1–3 |pages=1–76|doi=10.1002/mmnd.19870340102 }}

|

  • P. geinitzi
  • P. magna
  • P. minima
  • P. media
  • P. reticulata
  • P. mesostena

|

  • Grimmen (Krempien collection)
  • Lehmhagen, near Grimmen (Zessin collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="2" |

A grasshopper, member of the family Elcanidae.

|rowspan="2" |

Parelcana

|

  • P. tenuis

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Plesioschwinzia

|

  • P. thalassophila
  • P. lacera

|

  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Grasshoppers of the family Locustopsidae.

|

Protochaeta

|

  • P. masculina
  • P. lanceolata

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A grasshopper, member of the family Regiatidae.

|

Protogryllus{{cite book |last1=Zeuner |first1=F. E. |title=Fossil Orthoptera Ensifera |date=1939 |publisher=BM(NH) |location=London |pages=1–321}}

|

  • P. dobbertinensis
  • P. germanicus
  • P. sp.
  • P. vicinus
  • P. liadis
  • P. major
  • P. stenobasis
  • P. femina
  • P. acutipennis
  • P. minor
  • P. irregularis

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (BMNH collection)
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg Erratics

|

Specimens

|

A cricket-like grasshopper, type member of the family Protogryllidae.

|

Schwinzia

|

  • S. sola

|

  • Dobbertin, Mecklenburg (Zessin collection)

|

Specimens

|

Grasshoppers of the family Locustopsidae.

|

Synelcana{{cite journal |last1=Zessin |first1=W. |title=Neue Saltatoria (Insecta) aus dem Oberlias Mitteleuropas |journal=Freiberger Forschungshefte |date=1988 |volume=419 |issue=1 |pages=107–121}}

|

  • S. muelleri

|

  • Dobbertin, Mecklenburg (Zessin collection)

|

Specimens

|

A grasshopper, member of the family Elcanidae.

|

Zalmonites

|

  • Z. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A grasshopper, incertae sedis.

|

==Phasmatodea==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Durnovaria

|

  • D. cf. parallela

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A stick insect, member of the family Aerophasmidae.

|

File:Phasmatodea (16001849335).jpgn, Durnovaria was probably similar]]

==Blattodea==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Blattula

|

  • B. langfeldti
  • B. dubia

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg Erratics

|

Specimens

|

A cockroach, member of the family Blattulidae.

|

Caloblattina

|

  • C. mathildae

|

  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A cockroach, type member of the family Caloblattinidae.

|

Dipluroblattina

|

  • D. scudderi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="2" |

A cockroach, member of the family Blattulidae.

|rowspan="2" |

Eublattula

|

  • E. crassivena

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Liadoblattina

|

  • L. blakei

|

  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A carnivorous cockroach, type member of the family Raphidiomimidae.

|

Mesoblattina{{cite journal |last1=Vršanský |first1=P. |last2=Ansorge |first2=J. |title=Lower Jurassic cockroaches (Insecta: Blattaria ) from Germany and England |journal=African Invertebrates |date=2007 |volume=48 |issue=1 |pages=103–126 |url=https://www.researchgate.net/publication/242748378 |access-date=23 October 2021}}

|

  • M. protypa

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A cockroach, type member of the family Mesoblattinidae.

|

==Hemiptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Acromocoris{{cite journal |last1=Popov |first1=Y. A. |title=Jurassic bugs (Hemiptera: Heteroptera) from the Museum of Natural History in Vienna |journal=Annalen des Naturhistorischen Museums in Wien |date=1992 |volume=94 |issue=2–5 |pages=7–14}}

|

  • A. similis

|

  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A water boatman, member of the family Corixidae.

|

File:Corixidae sp - Flickr - S. Rae.jpg

Anosmus

|

  • A. spilopterus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Shore bugs, member of the family Archegocimicidae.

|

Aphlebocoris

|

  • A. punctata
  • A. nana

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|rowspan="2" |

Specimens

|rowspan="2" |

A creeping water bug, member of the family Naucoridae.

|rowspan="2" |

File:Ilyocoris cimicoides natur.jpg

Apopnus

|

  • A. magniclavus

|

  • Dobbertin, Mecklenburg (Greifswald collection)
Apsicoria

|

  • Apsicoria semideleta

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

True bugs, member of the family Pachymeridiidae.

|

Ardela

|

  • A. grimmenensis

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

Leafhoppers of the family Archijassidae.

|

Archegocimex

|

  • A. geinitzi
  • A. primitiva
  • A. liadis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Shore bugs, member of the family Archegocimicidae.

|

File:Shore Bug - Flickr - treegrow.jpg, Archegocimicidae genera where probably similar]]

Archicercopis

|

  • A. falcatus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Moss bugs of the family Progonocimicidae.

|

Archijassus

|

  • Archijassus heeri

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Leafhoppers of the family Archijassidae.

|

File:Membracidae (15402527004).jpg, Archijassidae genera where probably similar]]

Aphidulum

|

  • A. ciliatum

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A Sternorrhynchan, member of the family Protopsyllidiidae.

|

Archiconiopteryx

|

  • A. liasina

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A Stemorrhynchan, type member of the family Archiconiopterygidae.

|

Cathalus

|

  • Cathalus alutaceus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

True bugs, member of the family Pachymeridiidae.

|

Cixiites

|

  • C. liasinus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Planthoppers of the family Fulgoridiidae.

|

Cuneocoris

|

  • C. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A true bug, member of the family Cuneocoridae.

|

Diatillus

|

  • Diatillus debilis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Shore bugs, member of the family Archegocimicidae.

|

Eocercopis

|

  • E. ancyloptera
  • E. similis

|

  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Moss bugs of the family Progonocimicidae.

|

Fulgoridium

|

  • F. egens
  • F. lapideum
  • F. dubium
  • F. latum
  • F. venosum
  • F. pallidum
  • F. geinitzi
  • F. spilographum
  • F. oligoneurum
  • F. regulare
  • F. graphipterum
  • F. inconspicuum
  • F. quadrisignatum
  • F. breviradiatum
  • F. modestum
  • F. anale
  • F. marginepunctatum
  • F. picturatum
  • F. anomalum
  • F. stigmaticum
  • F. punctatum
  • F. mortuum
  • F. alatum
  • F. nodosum
  • F. megapolitanum
  • F. pulchrum
  • F. remotum
  • F. multivenosum
  • F. debile
  • F. trifurcatum
  • F. litorale
  • F. parvispilum
  • F. retractum
  • F. polyneurum
  • F. oligospilum
  • F. nubeculum
  • F. obtusum
  • F. grave
  • F. fenestratum
  • F. breve
  • F. dilutum
  • F. multipunctatum
  • F. ampliatum
  • F. clavatum
  • F. nebulosum
  • F. vicinum
  • F. curvipenne
  • F. elegantulum
  • F. defunctum
  • F. plicatum
  • F. exhumatum
  • F. angulosum
  • F. acutum
  • F. rotundatum
  • F. ancylla
  • F. bifurcatum
  • F. inaequale
  • F. brachyptilum
  • F. intercalatum
  • F. balticum

|

  • Grimmen (EMAU collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Planthoppers of the family Fulgoridiidae.

|

File:Fulgoridae at Mechode Padur.jpg, Fulgoridiidae genera where probably similar]]

Grimmenaphis

|

  • G. magnifica

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

An aphid, member of the family Oviparosiphidae.

|

File:Uroleucon nigrotuberculatum.jpg, Oviparosiphidae genera where probably similar]]

Hadrocoris

|

  • H. scutellaris

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A true bug, member of the family Hadrocoridae.

|

Hypocimex

|

  • Hypocimex membranaceus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

True bugs, member of the family Pachymeridiidae.

|

Liadopsylla

|

  • L. geinitzi
  • L. obtusa

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A jumping plant louse, member of the family Liadopsyllidae.

|

Liassocercopis

|

  • L. schnicki

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A cicadomorphan, member of the family Hylicellidae.

|

Liasocoris{{cite journal |last1=Wendt |first1=A. |title=Liasocoris hainmülleri n. sp., eine fossile Wanze aus Mecklenburg |journal=Archiv des Vereins der Freunde der Naturgeschichte in Mecklenburg |series=Neue Folge |date=1940 |volume=15 |issue=1 |pages=18–20}}

|

  • L. hainmuelleri

|

  • Schwinzer brickyard, near Goldberg

|

Specimens

|

A true bug, incertae sedis inside Hemiptera.

|

Margaroptilon

|

  • M. germanicum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Planthoppers of the family Fulgoridiidae.

|

Mesoledra

|

  • M. pachyneura

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Leafhoppers of the family Archijassidae.

|

Metafulgoridium

|

  • M. spilotum
  • M. ampliatum
  • M. graptum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Planthoppers of the family Fulgoridiidae.

|

Pachymeridium

|

  • P. dubium

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

True bugs, member of the family Pachymeridiidae.

|

Probascanion

|

  • P. megacephalum

|

  • Dobbertin, Mecklenburg (NHMW collection)

|

Specimens

|

A Coleorrhynchan, member of the family Probascanionidae.

|

Procercopis

|

  • P. alutacea
  • P. coriacea
  • P. liasina
  • P. similis
  • P. jurassica

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="2" |

Froghoppers of the family Procercopidae.

|rowspan="2" |

Procercopina

|

  • P. frenzeli

|

  • Klein Lehmhagen pit, Grimmen

|Specimens

Progonocimex{{cite journal |last1=Popov |first1=Y. A. |last2=Wootton |first2=R. J. |title=The upper Liassic Heteroptera of Mecklenburg and Saxony |journal=Systematic Entomology |date=1977 |volume=2 |issue=3 |pages=333–351|doi=10.1111/j.1365-3113.1977.tb00382.x |bibcode=1977SysEn...2..333P |s2cid=84814736 }}

|

  • P. jurassicus
  • P. liasinus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Moss bugs of the family Progonocimicidae.

|

File:HEMI Peloridiidae Oiophysa distincta 1.png, Progonocimicidae genera where probably similar]]

Progonocoris

|

  • P. pictus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Shore bugs, member of the family Archegocimicidae.

|

Sisyrocoris

|

  • S. rudis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

True bugs, member of the family Pachymeridiidae.

|

Toarcopsyllidium

|

  • T. ernsti

|

  • Klein Lehmhagen pit, Grimmen

|

Isolated Wings

|

A Sternorrhynchan, member of the family Protopsyllidiidae.

|

==Hymenoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Brigittepterus

|

  • B. brauckmanni

|

  • Dobbertin, Mecklenburg (Zessin collection)

|

Isolated Wings

|

Wasps, member of the family Ephialtitidae.

|

Grimmaratavites{{cite journal |last1=Rasnitsyn |first1=A. P. |last2=Ansorge |first2=J. |last3=Zhang |first3=H. C. |title=Ancestry of the orussoid wasps, with description of three new genera and species of Karatavitidae (Hymenoptera = Vespida: Karatavitoidea stat. nov.) |journal=Insect Systematics & Evolution |date=2006 |volume=37 |issue=3 |pages=179–190 |doi=10.1163/187631206788831137 |url=https://www.researchgate.net/publication/233599380 |access-date=23 October 2021}}

|

  • G. mirabilis

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A Wood Wasp, member of the family Karatavitidae.

|

Liadobracona{{cite journal |last1=Rasnitsyn |first1=A. P. |last2=Ansorge |first2=J. |last3=Zessin |first3=W. |title=New hymenopterous insects (Insecta: Hymenoptera) from the lower Toarcian (Lower Jurassic) of Germany |journal=Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen |date=2003 |volume=227 |issue=1–3 |pages=321–342|doi=10.1127/njgpa/227/2003/321 |bibcode=2003NJGPA.227..321R }}

|

  • L. raduhna

|

  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

|

Isolated Wings

|

Wasps, member of the family Ephialtitidae.

|

Xyelula

|

  • X. benderi

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A pseudo-Wasp, member of the family Sepulcidae.

|

==Megaloptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Dobbertinia

|

  • D. reticulata

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

An alderfly, member of the family Sialidae.

|

File:Sialidae fg01.jpg, Dobbertinia was probably similar]]

==Neuroptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Actinophlebia

|

  • A. megapolitana
  • A. aenea
  • A. parallela

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Lacewings of the family Prohemerobiidae

|

Apeirophlebia

|

  • A. grandis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A silky lacewing, member of the family Psychopsidae.

|

File:Silky Lacewing (6769953805).jpg

Epigambria{{cite journal |last1=Makarkin|first1=Vladimir|last2=Ansorge |first2=Jörg|last3=V Khramov |first3=Alexander|title=Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s.l. (Neuroptera) |journal=Palaeoentomology |date=2021 |volume=4 |issue=6 |pages=87–93 |doi=10.11646/palaeoentomology.4.6.1|bibcode=2021Plegy...4..6.1M |s2cid=245181387|url=https://www.mapress.com/pe/article/view/palaeoentomology.4.6.1 |access-date=16 December 2021|url-access=subscription }}

|

  • E. longipennis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A lacewing, member of the family Epigambriidae.

|

File:Rapisma tamilanum India.jpg, Epigambriidae members where probably similar]]

Liassochrysa{{cite journal |last1=Ansorge |first1=J. |last2=Schlüter |first2=T. |title=The earliest chrysopid: Liassochrysa stigmatica ng, n. sp. from the Lower Jurassic of Dobbertin, Germany |journal=Neuroptera International |date=1990 |volume=6 |issue=2 |pages=87–93 |url=https://www.researchgate.net/publication/271444550 |access-date=23 October 2021}}{{cite journal |last1=Jepson |first1=J. E. |title=A review of the current state of knowledge of fossil Mantispidae (Insecta: Neuroptera) |journal=Zootaxa |date=2015 |volume=3964 |issue=4 |pages=419–432 |doi=10.11646/zootaxa.3964.4.2 |pmid=26249453 |url=https://www.researchgate.net/publication/277668308 |access-date=23 October 2021}}

|

  • L. stigmatica

|

  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A Neuropteran of the family Mantispidae.

|

File:Mantispidae (5184942314).jpg

Loxophleps

|

  • L. costalis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

| rowspan="4" |

A lacewing, incertae sedis inside Neuroptera.

| rowspan="4" |

Melaneimon

|

  • M. dubium

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Melamnous

|

  • M. indistinctus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Mesoleon

|

  • M. dobbertinianus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Mesosmylina

|

  • M. falcifera

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A lance lacewing, member of the family Osmylidae.

|

Polyosmylus

|

  • P. excelsus

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A lacewing, member of the family Epigambriidae.

|

Prohemerobius

|

  • P. prodromus
  • P. ovatus
  • P. anomalus
  • P. costalis
  • P. persimilis
  • P. geinitzi
  • P. liasinus
  • P. major
  • P. parvulus
  • P. geinitzianus
  • P. dilaroides
  • P. chryseus
  • P. latus

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Lacewings of the family Prohemerobiidae.

|

Solenoptilon{{cite journal |last1=Geinitz |first1=F. E. |title=Beitrag zur Geologie Mecklenburgs |journal=Archiv des Vereins der Freunde der Naturgeschichte in Mecklenburg |date=1888 |volume=41 |issue=4–6 |pages=143–216}}

|

  • S. kochi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A lacewing, type member of the family Solenoptilidae.

|

Toarciconiopteryx{{cite journal |last1=Makarkin |first1=Vladimir N. |last2=Ansorge |first2=Jörg |title=The oldest dustywing (Neuroptera: Coniopterygidae) from the Lower Jurassic of Germany |journal=PalZ |date=2023 |volume=98 |pages=12–23 |doi=10.1007/s12542-023-00662-w |doi-access=free}}

|

  • T. dipterosimilis

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A dustywing, member of the subfamily Toarciconiopteryginae.

|

File:Coniopterygidae P1640716a.jpg, Toarciconiopteryx was probably similar]]

==Coleoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Allognosis

|

  • A. nitens

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="8" |

Beetles, incertae sedis inside Coleoptera.

|rowspan="8" |

Anancaeon

|

  • A. microcephalum

|

  • Dobbertin, Mecklenburg (NHMW collection)

|Specimens

Anomerus

|

  • A. punctifer

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Anypostatus

|

  • A. taurus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Apioderes

|

  • A. punctatus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Apsychus

|

  • A. alutaceus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Bareus

|

  • B. strigipennis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Bathygerus

|

  • B. bellus
  • B. divergens

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Carabites

|

  • C. dubius
  • C. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A ground beetle, member of the family Carabidae.

|

File:Platynus sp. - Kitchener, Ontario 2018-04-22 (01).jpg, Carabites was probably similar]]

Clinomerus

|

  • C. laticollis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Coptogyrinus

|

  • C. scutellatus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A whirligig beetle, member of the family Gyrinidae.

|

File:Escarabajo acuatico - Rhantus (Gyrinidae) (14805555906).jpg, Coptogyrinus was probably similar]]

Ecthlimma

|

  • E. forficuloides

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Enamma

|

  • E. striatum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Eurynucha

|

  • E. pseudobuprestis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A jewel beetle, member of the family Buprestidae.

|

File:Jewel Beetle . Chalcophora mariana Buprestidae - Flickr - gailhampshire.jpg, Eurynucha was probably similar]]

Gyrinulopsis

|

  • G. nanu

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Hydrobiites

|

  • H. dobbertinensis
  • H. punctatostriatus

|

  • Dobbertin, Mecklenburg (NHMW collection)

|

Specimens

|

A beetle, member of the family Permosynidae.

|

Keleusticus

|

  • K. zirkeli

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Liassodites{{cite journal |last1=Kirejtshuk |first1=A. G. |last2=Ansorge |first2=J. |title=An extraordinarily preserved new genus and species of Trachypachidae (Coleoptera, Adephaga) from the Early Jurassic of Germany and a review of fossil trachypachid genera |journal=Historical Biology |date=2022 |volume=34 |issue=6 |pages=958–975 |doi=10.1080/08912963.2022.2071709 |s2cid=248869983 |url=https://www.tandfonline.com/doi/abs/10.1080/08912963.2022.2071709 |access-date=15 June 2022|url-access=subscription }}

|

  • L. obsti

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A False Ground Beetle of the family Trachypachidae.

|

File:Systolosoma breve.jpg, Liassodites was probably similar]]

Masselytron

|

  • M. quinquestriatum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Nannoodes

|

  • N. pseudocistela

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Nebrioides

|

  • N. dobbertinensis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Onkedodimus

|

  • O. discicollis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Oxytoroptera

|

  • O. mediocris

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Paracurculium

|

  • P. punctatum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Parnidium

|

  • P. frechi
  • P. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Periboloptera

|

  • P. rotunda

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Plastobuprestites

|

  • P. elegans

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Plastonebria

|

  • P. scudderi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Polypamon{{cite journal |last1=Geinitz |first1=F. E.|title= Die Käferreste des Dobbertiner Lias|journal=Archiv des Vereins der Freunde der Naturgeschichte in Mecklenburg|date=1894 |volume=48 |issue=1 |pages=71–78}}

|

  • P. byrrhoides

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Pseudocyphon

|

  • P. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Pseudoprionites

|

  • P. liasinus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Beetles, incertae sedis inside Coleoptera.

|

Thoracotes

|

  • T. dubius

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A bark gnawing beetle, member of the family Trogossitidae.

|

File:Bark-gnawing beetle (Trogossitidae).jpg, Thoracotes was probably similar]]

==Trichoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Archiptilia

|

  • A. ovata

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="2" |

Caddisflies, member of the family Necrotauliidae.

|rowspan="2" |

Epididontus

|

  • E. geinitzianus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Liadotaulius

|

  • L. maior

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A finger-net caddisfly, member of the family Philopotamidae.

|

File:Philopotamus montanus.jpg, Liadotaulius was probably similar]]

Mesotrichopteridium{{cite journal |last1=Ansorge |first1=J. |title=Revision of the "Trichoptera" described by Geinitz and Handlirsch from the Lower Toarcian of Dobbertin (Germany) based on new material |journal=Proceedings of the 10th International Symposium on Trichoptera - Nova Supplementa Entomologica |date=2002 |volume=15 |issue=4–7 |pages=55–74 |url=https://www.researchgate.net/publication/296798102}}

|

  • M. intermedium
  • M. pusillum

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|rowspan="9" |

Caddisflies, member of the family Necrotauliidae.

|rowspan="9" |

Metarchitaulius

|

  • M. longus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Nannotrichopteron

|

  • N. gracile

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Necrotaulius

|

  • N. parvulus

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Palaeotaulius

|

  • P. vicinus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Parataulius

|

  • P. jurassicus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Paratrichopteridium

|

  • P. efossum
  • P. costale

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Pararchitaulius

|

  • P. ovalis

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Pseudorthophlebia

|

  • P. platyptera

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

==Lepidoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Eolepidopterigidae{{cite journal |last1=Zhang|first1=Q. Q. |last2=Mey |first2=W. |last3=Ansorge |first3=J. |last4=Starkey |first4=T. A. |last5=McDonald |first5=L. T. |last6=McNamara |first6=M. E. |last7=Jarzembowski |first7=E. A. |last8=Wichard |first8=W. |last9=Kelly |first9=R. |last10=Ren |first10=X. Y. |last11=Chen |first11=J. |last12=Zhang |first12=H. C. |last13=Wang |first13=B. |title=Fossil scales illuminate the early evolution of lepidopterans and structural colors |journal=Science Advances |date=2018 |volume=4 |issue=1 |pages=e1700988 |doi=10.1126/sciadv.1700988 |pmid=29651455 |pmc=5895446 |bibcode=2018SciA....4..988Z }}

|

  • Eolepidopterigidae indet. A
  • Eolepidopterigidae indet. B
  • Eolepidopterigidae indet. C

|

  • Klein Lehmhagen pit, Grimmen

|

  • Isolated wings
  • Isolated wing scales

|

A moth, member of the family Eolepidopterigidae.

|

==Mecoptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Mesobittacus

|

  • M. minutus

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Hangingflies, member of the family Bittacidae.

|

Mesochorista

|

  • M. sinuata

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A Scorpionfly, member of the family Permochoristidae.

|

Mesopanorpa

|

  • M. maculata

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Hangingflies, member of the family Orthophlebiidae.

|

Neorthophlebia

|

  • N. maculipennis
  • N. megapolitana
  • N. pallida
  • N. debilis
  • N. simillima
  • N. medialis
  • N. stigmatica
  • N. acutipennis

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Hangingflies, member of the family Bittacidae.

|

File:Bittacidae fg1.jpg

Orthophlebia

|

  • O. germanica
  • O. elongata
  • O. fuscipennis
  • O. limnophila
  • O. cf. limnophila
  • O. radialis
  • O. liadis
  • O. maculata
  • O. vicina
  • O. brunsvicensis

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Hangingflies, member of the family Orthophlebiidae.

|

Parabittacus{{cite journal |last1=Ansorge |first1=J. |title=Parabittacus analis Handlirsch 1939 und Parabittacus lingula (Bode 1953), Neorthophlebiiden (Insecta: Mecoptera) aus dem Oberen Lias von Deutschland |journal=Paläontologische Zeitschrift |date=1993 |volume=67 |issue=3–4 |pages=293–298 |doi=10.1007/BF02990282 |s2cid=130240591 |url=https://www.researchgate.net/publication/225408961 |access-date=23 October 2021}}

|

  • P. analis
  • P. lingula

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Hangingflies, member of the family Bittacidae.

|

Pseudopolycentropus

|

  • P. triangularis
  • P. perlaeformis
  • P. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A scorpionfly, type member of the family Pseudopolycentropodidae.

|

==Diptera==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Aenne{{cite journal |last1=Ansorge |first1=J. |title=Aenne liasina gen. et sp. n. - the most primitive non-biting midge (Diptera: Chironomidae: Aenninae subfam. n.)- from the Lower Jurassic of Germany. |journal=Polskie Pismo Entomologiczne |date=1999 |volume=68 |issue=1 |pages=431–443 |url=https://www.academia.edu/1572541 |access-date=23 October 2021}}

|

  • A. liasina

|

  • Schönenwald railway cutting, Grimmen
  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

The oldest known non biting midgess, members of the Chironomidae

|

File:Chironomidae Non-biting Midge.jpg

Antefungivora

|

  • A. germanica

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|rowspan="2" |

Flies, member of the family Antefungivoridae.

|rowspan="2" |

Archibio

|

  • A. mycetophilinus

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Archipleciomima{{cite journal |last1=Kopeć|first1=K. |last2=Soszyńska-Maj |first2=A. |last3=Gehler |first3=A. |last4=Ansorge |first4=J. |last5=Krzemiński |first5=W. |title=Mecoptera and Diptera from the early Toarcian (Early Jurassic) deposits of Wolfsburg – Große Kley (Lower Saxony, Germany) |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |date=2018 |volume=107 |issue=1 |pages=163–171 |doi=10.1017/S1755691017000226 |s2cid=134568600 |url=https://www.cambridge.org/core/journals/earth-and-environmental-science-transactions-of-royal-society-of-edinburgh/article/abs/mecoptera-and-diptera-from-the-early-toarcian-early-jurassic-deposits-of-wolfsburg-grosse-kley-lower-saxony-germany/0BE2318BEAB7F0AAC5F34D0BBDF9E02C |access-date=23 October 2021|url-access=subscription }}

|

  • A. germanica
  • A. sp.

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A fly, incertae sedis inside Diptera

|

Archirhyphus

|

  • A. geinitzi

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Flies, member of the family Protorhyphidae.

|

Architipula

|

  • A. stigmatica
  • A. latipennis
  • A. elegans
  • A. seebachi
  • A. nana
  • A. pusilla
  • A. pulla
  • A. debilis
  • A. simplex
  • A. intermedia
  • A. obliqua
  • A. areolata
  • A. maior
  • A. conspicua
  • A. clara
  • A. dubia

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A crane fly of the family Limoniidae.

|

File:Epiphragma ocellare . Limoniidae - Flickr - gailhampshire (1).jpg

Crenoptychoptera{{cite journal |last1=Lukashevich |first1=E. |last2=Ansorge |first2=J. |last3=Krzemiński |first3=W. |last4=Krzemińska |first4=E. |title=Revision of Eoptychopterinae (Diptera: Eoptychopteridae) |journal=Polskie Pismo Entomologiczne |date=1998 |volume=67 |issue=1 |pages=311–343 |url=https://www.researchgate.net/publication/285749465 |access-date=23 October 2021}}

|

  • C. dobbertinensis

|

  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A Phantom crane fly, member of the family Ptychopteridae.

|

File:Ptychoptera contaminata (Ptychopteridae) - (male imago), Arnhem, the Netherlands.jpg

Eoditomyia

|

  • E. primitiva

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A fly, type member of the family Eoditomyidae.

|

Eolimnobia

|

  • E. geinitzi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

| rowspan="2" |

A phantom crane fly, member of the family Ptychopteridae.

| rowspan="2" |

Eoptychoptera

|

  • E. simplex
  • E. eximia

|

  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|Specimens

Eotipula

|

  • E. defuncta
  • E. mortua
  • E. coarctata

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A crane fly of the family Limoniidae.

|

Grimmenia

|

  • G. prima
  • G. secunda

|

  • Lehmhagen, near Grimmen (Zessin collection)

|

Specimens

|

A crane fly of the family Limoniidae.

|

Grimmyia

|

  • G. baltica

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A snipe fly, member of the family Rhagionidae.

|

File:Rhagionidae 01.jpg

Heterorhyphus

|

  • H. triangularis

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A fly, type member of the family Heterorhyphidae.

|

Liassobrachyceron{{cite journal |last1=Krzeminski |first1=W. |last2=Ansorge |first2=J. |title=A new rhagionid fly from the Lower. Jurassic |journal=Polskie Pismo Entomologiczne |date=2005 |volume=74 |issue=3 |pages=369–372 |url=https://www.researchgate.net/publication/268508600 |access-date=23 October 2021}}

|

  • L. kotejai

|

  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A snipe fly, member of the family Rhagionidae.

|

Liassopsychodina

|

  • L. pommerana

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A Moth fly, member of the family Psychodidae.

|

File:Psychodidae 01.jpg

Mailotrichocera{{cite journal |last1=Krzemińska |first1=E. |last2=Krzemiński |first2=W. |last3=Dahl |first3=C. |title=Monograph of Fossil Trichoceridae (Diptera): Over 180 Million Years of Evolution |journal=Monograph of Fossil Trichoceridae |date=2009 |volume=1 |issue=1 |pages=1–171 |url=https://www.researchgate.net/publication/274719749 |access-date=23 October 2021}}

|

  • M. mikereichi
  • M. variabilis
  • M. zessini

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A Winter Crane Fly, member of the family Trichoceridae.

|

File:Trichocera spec. (Trichoceridae) - (imago), Elst (Gld), the Netherlands - 3.jpg

Mesorhyphus

|

  • M. nanus

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A wood gnat, member of the family Anisopodidae.

|

File:Sylvicola sp 01.jpg

Mesotipula{{cite journal |last1=Krzemiński |first1=W. |last2=Zessin |first2=W. |title=The Lower Jurassic Limoniidae from Grimmen (GDR) (Dipt. Nematocera) |journal=Deutsche Entomologische Zeitschrift |date=1990 |volume=37 |issue=3 |pages=39–43 |doi=10.1002/mmnd.19900370113 |url=https://onlinelibrary.wiley.com/doi/abs/10.1002/mmnd.19900370113 |access-date=23 October 2021|url-access=subscription }}{{cite journal |last1=Kopeć |first1=K. |last2=Ansorge |first2=J. |last3=Soszyńska-Maj |first3=A. |last4=Krzemiński |first4=W. |title=Revision of the genus Mesotipula Handlirsch, 1920 (Diptera, Limoniidae, Architipulinae) from the Lower Jurassic of Northeast Germany |journal=Historical Biology |year=2020 |volume=32 |issue=4 |pages=1–8 |doi=10.1080/08912963.2018.1503257 |bibcode=2020HBio...32..500K |s2cid=92596808 |url=https://www.tandfonline.com/doi/abs/10.1080/08912963.2018.1503257?journalCode=ghbi20 |access-date=23 October 2021|url-access=subscription }}

|

  • M. siggiae
  • M. vicina
  • M. cf. vicina
  • M. mirabilis
  • M. curvata
  • M. geinitzi
  • M. minuta
  • M. lapidaria
  • M. parva

|

  • Lehmhagen, near Grimmen (Zessin collection)
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A Crane Fly of the family Limoniidae.

|

Metatrichopteridium{{cite journal |last1=Ansorge |first1=J. |title=Lower Jurassic Hennigmatidae (Diptera) from Germany |journal=Studia dipterologica |date=2001 |volume=8 |issue=1 |pages=97–102 |url=https://www.researchgate.net/publication/271444320 |access-date=23 October 2021}}

|

  • M. confusum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A fly, member of the family Hennigmatidae.

|

Nannotanyderus{{cite journal |last1=Ansorge |first1=J. |title=Tanyderidae and Psychodidae (Insecta: Diptera) from the Lower Jurassic of northeastern Germany |journal=Paläontologische Zeitschrift |date=1994 |volume=68 |issue=1 |pages=199–210 |doi=10.1007/BF02989440 |bibcode=1994PalZ...68..199A |s2cid=128958459 |url=https://www.researchgate.net/publication/227010157 |access-date=23 October 2021}}

|

  • N. grimmenensis
  • N. krzeminskii

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A primitive Crane Fly, member of the family Tanyderidae.

|

Oryctochlus

|

  • O. toarciensis

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

The oldest known non biting midges, members of the Chironomidae.

|

Rhaetomyia

|

  • R. herrigi

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A Phantom midge of the family Chaoboridae.

|

File:Chaoboridae 2 Richard Bartz.jpg

Palaeobrachyceron

|

  • P. willmanni

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A Snipe fly, member of the family Rhagionidae.

|

Phryganidium

|

  • P. minimum

|

  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A Crane Fly of the family Limoniidae.

|

Pleciofungivora

|

  • P. liassica

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A fly, member of the family Pleciofungivoridae.

|

Podonomius

|

  • P. tumidus

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

The oldest known non biting midges, members of the Chironomidae.

|

Praemacrochile{{cite journal |last1=Ansorge |first1=J. |last2=Krzemiński |first2=W. |title=Lower Jurassic tanyderids (Diptera: Tanyderidae) from Germany |journal=Studia Dipterologica |date=2002 |volume=9 |issue=1 |pages=21–29 |url=https://www.researchgate.net/publication/285901797 |access-date=23 October 2021}}

|

  • P. decipiens
  • P. dobbertinensis

|

  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

|

Specimens

|

A primitive Crane Fly, member of the family Tanyderidae.

|

Protobrachyceron

|

  • P. liasinum

|

  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A fly, member of the family Protobrachyceridae.

|

Protoplecia

|

  • P. klafacki
  • P. liasina

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

A fly, type member of the family Protopleciidae.

|

Protorhyphus

|

  • P. simplex

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

|

Specimens

|

Flies, member of the family Protorhyphidae.

|

Tanypsycha

|

  • T. connexa

|

  • Klein Lehmhagen pit, Grimmen

|

Specimens

|

A Moth fly, member of the family Psychodidae.

|

=Echinodermata=

In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%. In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.{{cite thesis |last1= Stumpf |first1=Sebastian |title=A synoptic review of the vertebrate fauna from the "Green Series" (Toarcian) of northeastern Germany with descriptions of new taxa: A contribution to the knowledge of Early Jurassic vertebrate palaeobiodiversity patterns|type=PhD Thesis |publisher=Greifswald University |date=2017 |pages=1–47|url=https://epub.ub.uni-greifswald.de/frontdoor/deliver/index/docId/1941/file/STUMPF2017DISS.pdf|access-date=9 September 2021}}

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Crinoidea

|

Indeterminate

|

  • Dobbertin

|Columnar ossicles

|Incertae sedis

|

Holothuriida

|

Indeterminate

|

  • Dobbertin

|Holothurian wheels

|Incertae sedis

|

Sinosura

|

  • S. brodiei
  • S. spp.

|

  • Dobbertin

|Isolated and semiarticulated specimens

|

A brittle star, member of Ophioleucidae inside Ophioleucida. Identified as Ophiura longivertebralis.

|

=Vertebrates=

==Fishes==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Dapedium{{cite journal |last1=Thies |first1=D. |title=Dapedium pholidotum (AGASSIZ, 1832) - (Pisces, Actinopterygii) from Unter-Toarcium NW-Germany. |journal=Geologica et Palaeontologica |date=1988 |volume=22 |issue=6 |pages=89–121 |url=https://www.researchgate.net/publication/284228912 |access-date=24 October 2021}}

|

  • D. pholidotum

|

  • Dobbertin

|

Incomplete skull

|

Member of the Dapediidae.

|File:Dapedium pholidotumJW.png

Grimmenichthys{{cite journal |last1=Konwert |first1=M. |last2=Hörnig |first2=M. |title=Grimmenichthys ansorgei, gen. et sp. nov. (Teleostei, "Pholidophoriformes"), and other "pholidophoriform" fishes from the early Toarcian of Grimmen (Mecklenburg-Western Pomerania, Germany) |journal=Journal of Vertebrate Paleontology |date=2018 |volume=38 |issue=3 |pages=1–16 |doi=10.1080/02724634.2018.1451872 |s2cid=90344418 |url=https://www.tandfonline.com/doi/abs/10.1080/02724634.2018.1451871 |access-date=24 October 2021|hdl=11336/84457 |hdl-access=free }}

|

  • G. ansorgei
  • Cf. G. ansorgei

|

  • Klein Lehmhagen pit, Grimmen

|

Articulated, but incompletely preserved specimens

|

Member of the group Pholidophoriformes. This genus was a small 'pholidophoriform' teleost.

|

Grimmenodon{{cite journal |last1=Stumpf |first1=S. |last2=Ansorge |first2=J. |last3=Pfaff |first3=C |last4=Kriwet |first4=J. |title=Early Jurassic diversification of pycnodontiform fishes (Actinopterygii, Neopterygii) after the end-Triassic extinction event: evidence from a new genus and species, Grimmenodon aureum |journal=Journal of Vertebrate Paleontology |date=2017 |volume=37 |issue=4 |pages=e1344679 |doi=10.1080/02724634.2017.1344679 |pmid=29170576 |pmc=5646184 |bibcode=2017JVPal..37E4679S }}

|

  • G. aureum

|

  • Klein Lehmhagen pit, Grimmen

|

Almost complete left prearticular with dentition

|

Member of the order Pycnodontiformes. Is of comparatively small size, suggesting a hypothetical small standard length of the fish of about 7–10 cm at the time of death.

|

Gyrosteus{{cite journal |last1=Hornung |first1=J. J. |last2=Sachs |first2=S. |title=First record of Gyrosteus mirabilis (Actinopterygii, Chondrosteidae) from the Toarcian (Lower Jurassic) of the Baltic region |journal=PeerJ |date=2020 |volume=1 |issue=1 |pages=1–10 |url=https://peerj.com/articles/8400/ |access-date=16 July 2021}}

|

  • G. mirabilis

|

  • Ahrensburg Erratic Assemblage

|

GPIH 4864, Hyomandibula

|

Member of the Chondrosteidae inside Acipenseriformes.

|

Hybodus

|

  • H. hauffianus
  • H. spp.

|

  • Klein Lehmhagen pit, Grimmen
  • Ahrensburg erratic assemblage

|

  • Fragmentary Fin spine
  • Supposed coprolithes
  • Teeth

|Member of the Hybodontiformes.

|File:Hybodus hauffianus.png

Lepidotes{{cite journal |last1=Thies |first1=D. |title=Sinneslinien bei dem Knochenfisch Lepidotes elvensis (Blainville 1818) (Actinopterygii, Semionotiformes) aus dem Oberlias (Unter-Toarcium) von Grimmen in der DDR |journal=Neues Jahrbuch für Geologie und Paläontologie, Monatshefte |date=1989 |volume=11 |issue=1 |pages=692–704 |doi=10.1127/njgpm/1989/1989/692 |url=https://www.researchgate.net/publication/284040703 |access-date=24 October 2021}}

|

  • L. buelowianus
  • L. elvensis
  • L. spp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Ahrensburg Erratic Assemblage

|

Various specimens

|

Member of the family Lepidotidae inside Lepisosteiformes.

|File:Lepidotes elvensis.JPG

Lepisosteidae{{cite journal |last1=Kovalchuk |first1=О. М. |last2=Anfimova |first2=G. V. |title=Lepisosteiform fish (Holostei) ganoid scales from the middle jurassic deposits of Ukraine |journal=Zoodiversity |date=2020 |volume=54 |issue=1 |pages=35–42 |doi=10.15407/zoo2020.01.035 |s2cid=216317837 |url=https://www.researchgate.net/publication/339628108 |access-date=24 October 2021|doi-access=free }}

|

Indeterminate

|

  • Grimmen

|

GG 439/4-7 scales

|

The oldest confirmed true Gar, member of the Lepisosteidae.

|

File:Lepisosteus oculatus 1.jpg extant example of Gar]]

Leptolepidae

|

Indeterminate

|

  • Former clay pit of Dobbertin

|

  • Isolated head with missing postcranial bones
  • Assigned Teeth

|

Member of the family Leptolepidae.

|

Leptolepis{{cite journal |last1=Jaekel |first1=O. |title=Lepidotus und Leptolepis aus dem oberen Lias von Dobbertin, Mecklenburg |journal=Mitteilungen aus der Mecklenburgischen Geologischen Landesanstalt |date=1929 |volume=38 |issue=3 |pages=13–25}}{{cite journal |last1=Agassiz |first1=L. |title=Untersuchungen über die fossilen Fische der Lias-Formation—Aus einem Briefe des Vfsan-Professor Bronn |journal=Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde |date=1832 |volume=3 |issue=1 |pages=139–149}}{{cite journal |last1=Konwert |first1=M. |last2=Stumpf |first2=S. |title=Exceptionally preserved Leptolepidae (Actinopterygii, Teleostei) from the late Early Jurassic Fossil Lagerstatten of Grimmen and Dobbertin (Mecklenburg-Western Pomerania, Germany) |journal=Zootaxa |date=2017 |volume=42 |issue=1 |pages=249–296 |doi=10.11646/zootaxa.4243.2.2 |pmid=28610149 |url=https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4243.2.2 |access-date=24 October 2021|url-access=subscription }}

|

  • L. bronni
  • L. coryphaenoides
  • L. normandica
  • L. jaegeri
  • L. spp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin clay pit
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

|

  • Slightly disarticulated, incomplete head; Subadult, almost complete; fragmentary head; isolated head with pectoral girdle; incomplete head
  • Isolated remains in form of "Grätensandsteine"

|

Type member of the family Leptolepidae. The most abundant vertebrate recovered on the formation, including 3D preserved specimens, as well, is the main component of the Fishbone sandstones from the upper layers.

|File:Leptolepis coryphaenoides.jpg

Lycopteroidarum

|

  • L. sp. A
  • L. sp. B

|

  • Ahrensburg Erratic Assemblage

|

  • Otoliths

|

Otoliths of the family Lycopteroidea inside Lycopteriformes.

|

Mengius{{Cite journal |last1=Thies |first1=Detlev |last2=Stevens |first2=Kevin |last3=Ansorge |first3=Jörg |date=2024 |title=A new lepisosteiform neopterygian (Actinopterygii) from the lower Toarcian Grimmen Formation (Lower Jurassic) of Mecklenburg and Western Pomerania, Germany |journal=PalZ |volume=98 |issue=2 |pages=21–42 |bibcode=2024PalZ..tmp...23T |doi=10.1007/s12542-023-00683-5 |doi-access=free}}

|

  • M. acutidens

|

  • Klein Lehmhagen pit, Grimmen
  • Schwinzer Ziegelei ca 3 km E of Dobbertin

|

  • Holotype: Articulated, incomplete fish
  • Referred articulated, semiarticulated and fragmentary specimens

|Member of the family Lepidotidae inside Lepisosteiformes. Stomach content is also preserved on a specimen from Dobbertin, and is composed by arthropod cuticles.{{cite journal |last1=Thies |first1=D. |last2=Stevens |first2=K. |last3=Stumpf |first3=S. |year=2021 |title=Stomach contents of the Early Jurassic fish† Lepidotes Agassiz, 1832 (Actinopterygii, Lepisosteiformes) and their palaeoecological implications |url=https://www.tandfonline.com/doi/abs/10.1080/08912963.2019.1665040?journalCode=ghbi20 |journal=Historical Biology |volume=33 |issue=6 |pages=868–879 |doi=10.1080/08912963.2019.1665040 |bibcode=2021HBio...33..868T |s2cid=203892069 |access-date=24 October 2021|url-access=subscription }}

|

Pachycormiformes

|

Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|

Unknown, only cited.

|

Member of the family Pachycormiformes.

|

Pholidophorus{{cite journal |last1=Lehmann |first1=U. |date=1971 |title=Faziesanalyse der Ahrensburger Liasknollen auf Grund ihrer Wirbeltierreste |url=https://repo.uni-hannover.de/collections/1e5352c0-7d27-4218-a6e9-58226e3075e3 |journal=Mitteilungen aus dem Geologischen Institut der Technischen Universität Hannover |volume=10 |issue=1 |page=21-42}}

|

  • P. sp.

|

  • Ahrensburg Erratic Assemblage

|

  • Assigned Fragmentary remains

|

Member of the family Pholidophoriformes.

|

Pholidophoriformes

|

Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|

  • Isolated skull roof with articulated ethmoid region, parasphenoid, and partial braincase

|

Member of the family Pholidophoriformes. Resembles Pholidophoroides crenulata and P. limbata.

|

Proleptolepis

|

P. sp.

|

  • Klein Lehmhagen pit, Grimmen

|

  • MV 202612, fragmentary head .
  • Assigned Teeth

|

Member of the family Leptolepidae. Specimens attributed to Proleptolepis have previously been reported only from the Sinemurian of western Europe.

|

Saurorhynchus{{cite journal |last1=Maxwell |first1=E. E. |last2=Stumpf |first2=S. |title=Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany |journal=European Journal of Taxonomy |date=2017 |issue=321 |pages=1–29 |doi=10.5852/ejt.2017.321 |url=https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/446/961 |access-date=24 October 2021|doi-access=free }}

|

  • S. hauffi
  • S. sp.

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

|

  • Diganostic Incomplete skull
  • Incomplete skull

|

The youngest representative of the family Saurichthyidae. Distinctive for its large jaws, similar to modern Belonidae.

|

File:Saurorhynchus brevirostris.JPG

Tetragonolepis

|

  • T. semicincta

|

  • Klein Lehmhagen pit, Grimmen

|

  • Incomplete specimen
  • Isolated Jaw

|

Member of the family Dapediidae.

|

File:Tetragonolepis semicinctus.JPG

==Amniotes==

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

Ichthyosauria{{cite journal |last1=Oertel |first1=W. |date=1925 |title=Der Lias in Schleswig-Holstein |journal=Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Beilage-Band |volume=42 |issue=1 |pages=175–213}}{{cite journal |last1=Zessin |first1=W. |last2=Krempien |first2=W. |title=Bemerkenswerte Saurier- Krokodil- und Fischfunde aus dem Lias von Grimmen, Vorpommern |journal=NABU Nachrichten Mecklenburg-Vorpommern |date=2010 |volume=8 |issue=1 |pages=5–18|url=https://www.researchgate.net/publication/284947400 |access-date=24 October 2021}}

|

Indeterminate

|

  • Klein Lehmhagen pit, Grimmen
  • Ahrensburg Erratic Assemblage

|

  • Presacral centrum
  • 2 ?sacral centrum
  • Isolated vertebrae

|Incertae sedis

|

Macrospondylus

|

  • M. bollensis

|

  • Klein Lehmhagen pit, Grimmen

|Incomplete skull and associated osteoderm

|A marine crocodrylomorph, member of the family Machimosauridae. GG 422/6 can be clearly referred to an immature individual.

|File:Steneosaurus bollensis, view 2, Early Jurassic, Toarcian Age, Posidonia Shale, Holzmaden, Baden-Wurttemberg, Germany - Houston Museum of Natural Science - DSC01846.JPG

Megalosauridae{{cite journal |last1=Von Huene |first1=F. |date=1966 |title=Ein Megalosauriden-Wirbel des Lias aus norddeutschem Geschiebe |url=https://www.archivportal-d.de/item/DO7Q54JSRGR2VEF2QI2PFUTN73NKZACG |journal=Neues Jahrbuch für Geologie und Paläontologie |volume=Monatshefte |issue=1 |pages=318–319}}

|Indeterminate

|

  • Forst Hagen gravel pit

|Dorsal vertebrae

|A theropod, member of the family Megalosauridae. Was referred to Megalosaurus. The vertebrae centrum measures 80 mm, implying a medium-sized theropod (~5 m long).

|File:Marshosaurus_restoration.jpg, example of basal Orionides]]

Mesoeucrocodylia

|Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|Cervical vertebra

|A crocodrilian, member of Mesoeucrocodylia. Is considered to come from an osteologically immature individual.

|

Meyerasaurus

|

  • M. sp.

|

  • Ahrensburg Erratic Assemblage

|

  • Incomplete coracoid
  • Associated rib

|A pliosauroid, member of the family Rhomaleosauridae.

|File:Meyerasaurus victor.jpg]]

Mystriosaurus

|

  • M. sp.

|

  • Klein Lehmhagen pit, Grimmen

|Partial rostrum with teeth

|A marine crocodrylomorph, member of the family Teleosauridae. Is indicative of a juvenile individual, with expected around 200–250 mm long skull and the entire animal about 1.50 m.

|

Neoichthyosauria{{cite journal |last1=Geinitz |first1=F.E |title=Ichthyosaurus von Dobbertin |journal=Archiv des Vereins der Freunde der Naturgeschichte in Mecklenburg |date=1900 |volume=54 |issue=1 |pages=382–383}}

|

Indeterminate

|

  • Clay Pit of Dobbertin

|

Four articulated tail vertebrae.

|

An indeterminate ichthyosaur, has been assigned to the species Stenopterygius longifrons.

|

Ophthalmosauridae?{{cite journal |last1=Zessin |first1=W. |title=Saurierfund im Lias epsilon von Grimmen, Kreis Nordvorpommern (Fundbericht). |journal=Geschiebekunde Aktuell |date=1995 |volume=11 |issue=4 |pages=1–10,113 |url=https://www.researchgate.net/publication/327338847 |access-date=24 October 2021}}

|

Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|

Partial skull and associated postcranial elements preserved in a concretion

|

It has an expanded basipterygoid process on the basisphenoid, only currently known in members of the Ophthalmosauridae

|

Plesiosauria

|Indeterminate

|

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, falciferum zone

|

  • Three caudal vertebrae and other postcranial elements
  • Gastralia
  • Phalanx
  • Cervical rib and other postcranial elements

|Incertae sedis

|

Plesiosauroidea

|Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|

  • Cervical centrum
  • Caudal centrum

|Incertae sedis

|

Pterosauria?{{Cite journal |last1=Schade |first1=Marco |last2=Ansorge |first2=Jörg |date=2024 |title=Enigmatic fragment possibly marks the first pterosaur record from the Lower Toarcian of Grimmen, NE Germany |journal=PalZ |doi=10.1007/s12542-024-00698-6 |doi-access=free}}

|Indeterminate

|

  • Klein Lehmhagen pit, Grimmen

|GG 510, fragmentary tibiofibular(?)

|A possible pterosaur bone, with features similar to the tibiofibular of coeval taxon Dorygnathus banthensis. Alternatively it can be another type of bone and belong to stem-turtles, plesiosaurs, dinosaurs or even anurans.{{Cite journal |last1=Schade |first1=Marco |last2=Ansorge |first2=Jörg |date=2024 |title=Enigmatic fragment possibly marks the first pterosaur record from the Lower Toarcian of Grimmen, NE Germany |journal=PalZ |doi=10.1007/s12542-024-00698-6 |doi-access=free}}

|File:Dorygnathus banthensis model.jpg

Seeleyosaurus

|

  • S.? sp.

|

  • Ahrensburg Erratic Assemblage

|

  • Three articulated dorsal vertebrae

|A plesiosaur, member of the family Plesiosauridae.

|File:SeeleysaurusDB.jpg]]

Stenopterygius{{cite journal |last1= Stumpf |first1=Sebastian |title=New information on the marine reptile fauna from the lower Toarcian (Early Jurassic) "Green Series" of North-Eastern Germany|journal=N. Jb. Geol. Paläont. Abh. 2|date=2016 |volume=280 |issue=1 |pages=87–105|doi=10.1127/njgpa/2016/0567 |bibcode=2016NJGPA.280...87S |url=https://epub.ub.uni-greifswald.de/frontdoor/deliver/index/docId/1941/file/STUMPF2017DISS.pdf|access-date=9 September 2021}}{{cite journal |last1=Maisch |first1=M. W. |last2=Ansorge |first2=J. |title=The Liassic ichthyosaur Stenopterygius cf. quadriscissus from the Lower Toarcian of Dobbertin (northeastern Germany) and some considerations on Lower Toarcian marine reptile palaeobiogeography |journal=Paläontologische Zeitschrift |date=2004 |volume=78 |issue=1 |pages=161–171 |doi=10.1007/BF03009136 |bibcode=2004PalZ...78..161M |s2cid=130196232 }}

|

  • S. cf. quadriscissus

|

  • Clay pit of Dobbertin

|

Posterior left half of the cranium.

|

An icthyosaur of the family Stenopterygiidae.

|File:Stenopterygius triscissus SW.png

Teleosauridae

|Indeterminate

|

  • Ahrensburg Erratic Assemblage

|

  • Osteoderms
  • Isolated non described remains

|Incertae sedis

|

Temnodontosaurus

|

  • T. cf. platyodon

|

  • Ahrensburg Erratic Assemblage

|

  • Skull
  • Coracoid
  • Associated rib fragment

|

An icthyosaur, type member of Temnodontosauridae. Assigned to Ichthyosaurus sp., but also suggested affinities to "Leptopterygius" (= Temnodontosaurus) platyodon.

|File:Temnodontosaurus DB.jpg

= Flora =

class="wikitable sortable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

style="background:#D1FFCF;" | Botryococcus

|style="background:#D1FFCF;" |

  • B. sp.

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Freshwater algae

|style="background:#D1FFCF;" | Indicates freshwater influx in palynofacies assemblage IV, associated with coastal or deltaic settings.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Equisetites{{Cite journal |last=Mantei |first=S. |date=2021 |title=Ein dolomitisierter Schachtelhalm (Equisetites sp.) in einem Liasgeschiebe (Toarcium) Vorpommerns |url=https://www.researchgate.net/publication/369088426 |journal=Geschiebekunde Aktuell |volume=37 |issue=4 |pages=106–113}}

|style="background:#D1FFCF;" |

  • E. sp.
  • E. cf. Münsteri
  • E. columnare

|style="background:#D1FFCF;" |

  • Glacial Erratic Near
  • Grimmen
  • Dobbertin
  • Ahrensburg area{{Cite journal |last=Ernst |first=W. |date=1920 |title=Jura- und marine Unterkreidegeschiebe aus dem Diluvium Schleswig-Holsteins (Vortrag) |journal=Zeitschrift der deutschen geologischen Gesellschaft |volume=72 |pages=285–289}}
  • Erratic deposit near Lüttow{{Cite journal |last=Zessin |first=W. |date=2014 |title=Kleine Mitteilungen - Ein interessantes Lias-Geschiebe mit Schachtelhalmen (Equisetites sp.) aus der Kiesgrube Lüttow bei Zarrentin, Mecklenburg |journal=Mitteilungen der Naturforschenden Gesellschaft Mecklenburg |volume=14 |issue=1 |pages=71–72}}

|style="background:#D1FFCF;" |Isolated stems, rhizomes

|style="background:#D1FFCF;" |Equisetalean stems of the family Equisetidae. Dominant in mono-to oligotypic stands near water bodies, indicative of high terrestrial input in palynofacies assemblages I and IV.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" | Erlansonisporites

|style="background:#D1FFCF;" |

  • E. sparassis
  • E. sp.

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Transported from northern sources (e.g., Fennoscandia), indicative of humid climates.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" | Hughesisporites

|style="background:#D1FFCF;" |

  • H. pustulatus

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Likely transported via suspension load plumes, reflecting freshwater influx.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" | Minerisporites

|style="background:#D1FFCF;" |

  • M. institus
  • M. volucris

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Associated with warm, humid conditions and high atmospheric CO2.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" |Pagiophyllum{{Cite journal |last=Salfeld |first=H. |date=1909 |title=Beiträge zur Kenntnis jurassischer Pflanzenreste aus Norddeutschland |url=https://www.schweizerbart.de/papers/palae/detail/56/66129/Beitrage_zur_Kenntnis_jurassischer_Pflanzenreste_aus_Norddeutschland_Pompeckj_J_F_Beitrage_zur_Palaontologie_und_Stratigraphie_des_nordwestdeutschen_Jura_I |journal=PALAEONTOGRAPHICA Beiträge zur Naturgeschichte der Vorzeit |volume=56 |issue=1 |pages=1–36}}{{Cite journal |last=Neuwald |first=HK. |date=2008 |title=Jura-Hölzer aus Vorpommern |journal=Fossilien |issue=1/2008 |pages=59–60}}

|style="background:#D1FFCF;" |

  • P. sp.
  • P. cf. Kurri

|style="background:#D1FFCF;" |Grimmen clay pit

|style="background:#D1FFCF;" |Leaves, driftwood

|style="background:#D1FFCF;" |Conifer macrofossils of the family Cheirolepidiaceae or Araucariaceae. High phytoclasts and cuticles in palynofacies assemblages I, II, and IV support conifer presence.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" | Paxillitriletes

|style="background:#D1FFCF;" |

  • P. phyllicus

|style="background:#D1FFCF;" |

  • Grimmen clay pit
  • Reinberg 1E

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with the Isoetales. Transported from Fennoscandia, indicates humid climate, biostratigraphic marker (Paxillitriletes phyllicus Zone, also seen in the Ciechocinek Formation)

|style="background:#D1FFCF;" |File:Isoetes_echinospora_NRCS-1.png, typical example of Isoetales. Paxillitriletes probably come from a similar or a related Plant]]

style="background:#D1FFCF;" | Striatriletes

|style="background:#D1FFCF;" |

  • S. excavatus

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with the Selaginellaceae. Reflects freshwater influx from northern sources.

|style="background:#D1FFCF;" |

style="background:#D1FFCF;" | Trileites

|style="background:#D1FFCF;" |

  • T. spp.

|style="background:#D1FFCF;" | Grimmen clay pit

|style="background:#D1FFCF;" | Megaspores

|style="background:#D1FFCF;" | Affinities with lycophytes or ferns. Indicates diverse megaspore flora transported from northern sources.

|style="background:#D1FFCF;" |

See Also

References