Pabdeh Formation

{{Infobox rockunit|name=Pabdeh Formation|image=|caption=|type=Formation|age=Late Eocene to Early Oligocene, {{fossilrange|37.7|28}}|period=Priabonian|prilithology=Marl, limestone|otherlithology=Oil shales|namedfor=|namedby=|region=|country=Iran|coordinates=|unitof=Zagros fold and thrust belt|subunits=|underlies=Asmari Formation|overlies=Gurpi Formation|thickness=|extent=|area=|map=|map_caption=}}The Pabdeh Formation is a Late Eocene to Early Oligocene-aged{{Cite journal |last1=Joudaki |first1=M. |last2=Baghbani |first2=D. |date=2018-09-01 |title=Biostratigraphy of Oligocene and Lower Miocene deposits, Anneh Anticline, Folded Zagros, SW of Iran |url=https://link.springer.com/article/10.1007/s13146-017-0362-5 |journal=Carbonates and Evaporites |language=en |volume=33 |issue=3 |pages=509–515 |doi=10.1007/s13146-017-0362-5 |issn=1878-5212|url-access=subscription }}{{Cite journal |last1=Joudaki |first1=Mohamad |last2=Asnavandi |first2=Hamed |last3=Panah |first3=Fatemeh Mahmoodi |last4=Baghbani |first4=Darioush |date=2020-03-12 |title=The regional facies analysis and depositional environments of the Oligocene and Lower Miocene deposits; Zagros Basin, SW of Iran |url=https://link.springer.com/article/10.1007/s13146-020-00575-8 |journal=Carbonates and Evaporites |language=en |volume=35 |issue=2 |pages=39 |doi=10.1007/s13146-020-00575-8 |issn=1878-5212|url-access=subscription }} geological formation in Iran. It outcrops along the Zagros Mountains. It was deposited in a deepwater environment in the upper bathyal zone of the Paratethys Sea, with these abyssal sediments being lifted high above sea level from the uplift of the Zagros fold and thrust belt as part of the Alpide orogeny.

A high number of fossil fishes are known from this formation, the majority of which appear to have pelagic and deepwater affinities. These are mostly known from two localities near Elam and Istehbanat.{{Cite book |last1=Muséum national d'histoire naturelle (France) |url=https://www.biodiversitylibrary.org/item/281269 |title=Notes et mémoires sur le Moyen-Orient |last2=naturelle (France) |first2=Muséum national d'histoire |date=1966 |publisher=Muséum national d'histoire naturelle |volume=t.8 (1966) |location=Paris}} The age of these localities have been disputed, and have previously been considered to be Early Oligocene, but are now generally thought to be middle to late Eocene in age, most likely Priabonian.{{Cite journal |last1=Bannikov |first1=A. F. |last2=Erebakan |first2=I. G. |date=2023-10-01 |title=On the Evolution of Some Groups of Marine Bony Fishes in the Cenozoic of the Tethys and Paratethys |url=https://link.springer.com/article/10.1134/S0031030123050015 |journal=Paleontological Journal |language=en |volume=57 |issue=5 |pages=475–490 |doi=10.1134/S0031030123050015 |issn=1555-6174|url-access=subscription }} Some fossils of migratory insects (a locust and ant alates), as well as fish and birds, are also known from presumed early Oligocene-aged strata near Babaheidar.

Paleobiota

= Cartilaginous fishes =

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!Genus

!Species

!Notes

!Images

Xiphodolamia

|X. serrata

|A xiphodolamiid shark.{{Cite journal |last1=Adnet |first1=S. |last2=Hosseinzadeh |first2=R. |last3=Antunes |first3=M. T. |last4=Balbino |first4=A. C. |last5=Kozlov |first5=V. A. |last6=Cappetta |first6=H. |date=2009-10-01 |title=Review of the enigmatic Eocene shark genus Xiphodolamia (Chondrichthyes, Lamniformes) and description of a new species recovered from Angola, Iran and Jordan |url=https://www.sciencedirect.com/science/article/abs/pii/S1464343X09000752 |journal=Journal of African Earth Sciences |volume=55 |issue=3 |pages=197–204 |doi=10.1016/j.jafrearsci.2009.04.005 |issn=1464-343X|url-access=subscription }}

|frameless

= Bony fishes =

Based primarily on Arambourg (1967), with taxonomic changes made where necessary:

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!Genus

!Species

!Locality

!Notes

!Images

Aluvarus

|A. praeimperialis

|Elam

|The larva of a percomorph of uncertain affinities, once thought to be a louvar relative.{{Cite journal |last1=Bannikov |first1=Alexandre F. |last2=Tyler |first2=James C. |date=1995 |title=Phylogenetic Revision of the Fish Families Luvaridae and Kushlukiidae (Acanthuroidei), with a New Genus and Two New Species of Eocene Luvarids |url=https://repository.si.edu/handle/10088/19149 |journal=Smithsonian Contributions to Paleobiology |language=en |issue=81 |pages=1–45 |doi=10.5479/si.00810266.81.1 |hdl=10088/19149}}

|frameless

Arambourgthurus

|A. scombrurus

|Istehbanat

|A surgeonfish. One of the only known surgeonfish to inhabit a pelagic ecosystem.{{Cite journal |last1=Carnevale |first1=Giorgio |last2=Tyler |first2=James C. |date=2018 |title=The caudal skeleton of Arambourgthurus scombrurus (Arambourg, 1967), a Paleogene oceanic surgeonfish |url=http://www.bioone.org/doi/10.2988/17-00023 |journal=Proceedings of the Biological Society of Washington |language=en |volume=131 |issue=1 |pages=101–110 |doi=10.2988/17-00023 |issn=0006-324X|url-access=subscription }}

|

?Astronesthes

|?A. simus

|Elam

|An apparent stareater, taxonomic affinity uncertain.

|frameless

Babelichthys

|B. olneyi

|Elam

|A bizarre crestfish.{{Cite journal |last=Davesne |first=Donald |date=2017-06-28 |title=A fossil unicorn crestfish (Teleostei, Lampridiformes, Lophotidae) from the Eocene of Iran |journal=PeerJ |language=en |volume=5 |pages=e3381 |doi=10.7717/peerj.3381 |issn=2167-8359 |pmc=5493034 |pmid=28674642 |doi-access=free}}

|frameless

Berycomorus

|B. firdoussi

|Elam

|A beryciform.

|

Bregmaceros

|B. filamentosus

|Elam

|A codlet.

|frameless

Caucasiganus

|C. sp.

|Istehbanat

|A relative of rabbitfish.{{Cite journal |last1=Bannikov |first1=Alexandre F. |last2=Tyler |first2=James C. |last3=Sorbini |first3=Chiara |date=2010 |title=Two new taxa of Eocene rabbitfishes (Perciformes, Siganidae) from the North Caucasus (Russia), with redescription of Acanthopygaeus agassizi (Eastman) from Monte Bolca (Italy) and a phylogenetic analysis of the family |url=https://www.researchgate.net/publication/266087721 |journal=Bollettino del Museo Civico di Storia Naturale di Verona |volume=34 |pages=3–21}}

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Cheilinus

|C. longifillis

|Istehbanat

|A wrasse.

|frameless

Cottopsis

|C. gaudryi

|Elam

|An oceanic basslet.{{Cite journal |last=Prokofiev |first=A. M. |date=2007-09-01 |title=The osteology of Bathysphyraenops symplex and the diagnosis of the Howellidae (Perciformes: Percoidei) family |url=https://link.springer.com/article/10.1134/S0032945207080036 |journal=Journal of Ichthyology |language=en |volume=47 |issue=8 |pages=566–578 |doi=10.1134/S0032945207080036 |issn=1555-6425|url-access=subscription }}

|

Dussumieria

|D. elami

|Elam

|A rainbow sardine.

|frameless

Eosternoptyx

|E. discoidalis

|Elam

|A deep-sea hatchetfish.{{Cite journal |last=Afsari |first=S. |date=2014 |title=A new deep-sea hatchetfish (Teleostei: Stomiiformes: Sternoptychidae) from the Eocene of Ilam, Zagros Basin, Iran |journal=Bollettino della Società Paleontologica Italiana |volume=53 |issue=1 |pages=27–37|s2cid=211198956 }}

|

Epinnula

|E. cancellata

|Elam

|A snake mackerel.

|frameless

Etrumeus

|E. hafizi

|Istehbanat

|A round herring.

|frameless

Grammatorcynus

|G. scomberoides

|Istehbanat, Elam

|A mackerel.

|frameless

Iraniplectus

|I. bakhtiari

|Babaheidar

|A pufferfish-like tetraodontiform.Tyler, J.C.; Mirzaie, M.; Nazemi, A. (2006). New genus and species of basal tetraodontoid puffer fish from the Oligocene of Iran, related to the Zignoichthyidae (Tetraodontiformes). Bollettino del Museo Civico di Storia Naturale di Verona Geologia Paleontologia Preistoria, 30, 49-58

|frameless

Maurolicus

|M. morgani

|Elam

|A pearlside.

|frameless

?Myroconger

|?M. rouslami

|Elam

|An apparent thin eel, taxonomic affinity uncertain.

|frameless

Opisthonema

|O. persicum

|Elam

|A thread herring.

|frameless

Palaeorhynchus

|P. altivelis

|Istehbanat

|A palaeorhynchid billfish.

|frameless

rowspan="2" |Palimphyes (=Dipterichthys)

|P. nematophorus

| rowspan="2" |Elam

| rowspan="2" |A euzaphlegid.{{Cite journal |last1=Monsch |first1=Kenneth A. |last2=Bannikov |first2=Alexandre F. |date=2011 |title=New taxonomic synopses and revision of the scombroid fishes (Scombroidei, Perciformes), including billfishes, from the Cenozoic of territories of the former USSR |url=https://www.cambridge.org/core/journals/earth-and-environmental-science-transactions-of-royal-society-of-edinburgh/article/abs/new-taxonomic-synopses-and-revision-of-the-scombroid-fishes-scombroidei-perciformes-including-billfishes-from-the-cenozoic-of-territories-of-the-former-ussr/37C191646AD6ECED0B473DE94B6971EC |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |language=en |volume=102 |issue=4 |pages=253–300 |doi=10.1017/S1755691011010085 |issn=1755-6910|url-access=subscription }}

| rowspan="2" |frameless

P. leptosomus
Paucaichthys

|P. elamensis

|Elam

|A pomfret.{{Cite journal |last=Přikryl |first=Tomáš |last2=Bannikov |first2=Alexandre F. |date=2014 |title=A new species of the Oligocene pomfret fish Paucaichthys (Perciformes; Bramidae) from Iran |url=https://www.researchgate.net/publication/262925988_A_new_species_of_the_Oligocene_pomfret_fish_Paucaichthys_Perciformes_Bramidae_from_Iran |journal=N. Jb. Geol. Paläont. Abh. |volume=272 |issue=3 |pages=325–330}}

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Praewoodsia

|P. mesogeae

|Elam

|A bristlemouth.

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Priacanthopsis

|P. crassispinus

|Elam

|A bigeye.

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Proserrivomer

|P. mecquenemi

|Elam

|A sawtooth eel.

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Propteridium

|P. douvillei

|Elam

|A viviparous brotula.

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Pristigenys

|P. macropus

|Istehbanat

|A bigeye.

|frameless

Protolophotus

|P. elami

|Elam

|A crestfish.

|frameless

?Psenes

|?P. macrolepis

|Elam

|A driftfish, taxonomic identity uncertain.

|frameless

Pseudoholocentrum

|P. cristatum

|Elam

|A holocentrid.

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Rybapina

|R. orientalis

|Elam

|A driftfish.{{Cite journal |last=Bannikov |first=Alexandre F. |date=1993 |title=FOSSIL STROMATEOID FISHES OF THE FAMILIES NOMEIDAE AND ARIOMMIDAE (PERCIFORMES) |url=https://www.researchgate.net/publication/268211561 |journal=Paleontological Journal |volume=27 |issue=4}}{{Cite journal |last=Bannikov |first=A. F. |date=2018-11-01 |title=A New Genus and Species of Stromateoid Fishes (Perciformes, Stromateoidei) from the Lower Oligocene of the Northern Caucasus |url=https://link.springer.com/article/10.1134/S0031030118060035 |journal=Paleontological Journal |language=en |volume=52 |issue=6 |pages=631–638 |doi=10.1134/S0031030118060035 |issn=1555-6174|url-access=subscription }}

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Scomber

|S. saadii

|Istehbanat

|A mackerel.

|frameless

Scomberomorus

|S. speciosus

|Istehbanat

|A mackerel.

|frameless

Scopeloides

|S. glarisianus

|Elam

|A bristlemouth.

|frameless

Sphyraena

|S. longimana

|Istehbanat

|A barracuda.

|frameless

Thyrsitoides

|T. zarathoustrae

|Elam

|A relative of the blacksail snake mackerel.

|frameless

?Urosphen

|?U. iranensis

|Elam

|A urosphenid syngnathiform.

|frameless

= Birds =

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!Genus

!Species

!Locality

!Notes

!Images

Diomedeoides (=Frigidafrons)

|D. babaheydariensis

|Babaheidar

|A diomedeoidid seabird.{{Cite journal |last1=Peters |first1=Dieter Stefan |last2=Hamedani |first2=Ali |date=2000-05-01 |title=Frigidafons babaheydariensis n. sp., ein Sturmvogel aus dem Oligozän des Irans (Aves: Procellariidae) |url=https://link.springer.com/article/10.1007/BF03043661 |journal=Senckenbergiana Lethaea |language=de |volume=80 |issue=1 |pages=29–37 |doi=10.1007/BF03043661 |issn=0037-2110|url-access=subscription }}

|frameless

= Arthropods =

== Crustaceans ==

Based on Garassino, Bahrami, Yazdi & Vega (2014):{{Cite journal |author1=Alessandro Garassino |author2=Ali Bahrami |author3=Medhi Yazdi |author4=Francisco J. Vega |year=2014 |title=Report on decapod crustaceans from the Eocene of Zagros Basin, Iran |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=274 |issue=1 |pages=43–54 |doi=10.1127/njgpa/2014/0442}}

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!Genus

!Species

!Locality

!Notes

!Images

Eogordonella

|E. iranianiensis

| rowspan="3" |Babaheidar

|A solenocerid shrimp.

|

Eopabdehus

|E. babaheydariensis

|A penaeid shrimp.

|

Parsacus

|P. eocenicus

|A slipper lobster.

|

== Insecta ==

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!Genus

!Species

!Locality

!Notes

!Images

Acrididae indet.

|

| rowspan="2" |Babaheidar

|A short-horned grasshopper of uncertain affinities, potentially in a migratory locust-like phase.{{Cite journal |last1=Mirzaie Ataabadi |first1=Majid |last2=Bahrami |first2=Ali |last3=Yazdi |first3=Mehdi |last4=Nel |first4=André |date=2019-05-28 |title=A locust witness of a trans-oceanic Oligocene migration between Arabia and Iran (Orthoptera: Acrididae) |url=https://www.tandfonline.com/doi/full/10.1080/08912963.2017.1378651 |journal=Historical Biology |language=en |volume=31 |issue=5 |pages=574–580 |doi=10.1080/08912963.2017.1378651 |issn=0891-2963}}

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Formicidae indet.

|

|A potentially myrmicine ant of uncertain affinities, migratory alates.

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References