Papilio memnon

{{Speciesbox

| name = Great Mormon

| image = Papilio-pjt1.jpg

| image_caption = Male

| image2 = Papilio-pjt2.jpg

| image2_caption = Female

| taxon = Papilio memnon

| authority = Linnaeus, 1758

| synonyms = Princeps memnon

| subdivision =

P. m. memnon
P. m. agenor Linnaeus, 1768
P. m. anceus Cramer, [1779]
P. m. thunbergi Siebold, 1824
P. m. lowii Druce, 1873
P. m. oceani Doherty, 1891
P. m. merapu Doherty, 1891
P. m. pryeri Rothschild, 1895
P. m. clathratus Rothschild
P. m. subclathratus Fruhstorfer
P. m. coeruleus van Eecke
P. m. perlucidus Fruhstorfer
P. m. heronus Fruhstorfer, 1902
P. m. tanahsahi Eliot, 1982

}}

Papilio memnon, the great Mormon, is a large butterfly native to southern Asia that belongs to the swallowtail family. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.

Range

Its range includes north-eastern India (including Sikkim, Assam and Nagaland), Nepal, Bangladesh, Myanmar, Nicobar Islands, Andaman Islands (stragglers only), western, southern and eastern China (including Hainan), Taiwan, southern Japan including Ryukyu Islands, Thailand, Laos, Vietnam, Kampuchea, Malaysia and Indonesia (Sumatra, Mentawai Islands, Nias, Batu, Simeulue, Bangka, Java, Kalimantan and the Lesser Sunda Islands).

Status

This species is common and not threatened. The cultivation of citrus all over Southern Asia provides an abundance of food plants.

Description and polymorphy

The butterfly is large with a {{convert|120|to|150|mm}} span. It has four male and many female forms, the females being highly polymorphic and many of them being mimics of unpalatable butterflies. This species has been studied extensively for understanding the genetic basis for polymorphy and Batesian mimicry. As many as twenty-six female forms are reported.{{cite journal|author=C. A. Clarke, P. M. Sheppard & I. W. B. Thornton|year=1968|title=The genetics of the mimetic butterfly Papilio memnon L.|journal=Philosophical Transactions of the Royal Society B|volume=254|issue=791|pages=37–89|doi=10.1098/rstb.1968.0013|jstor=2416804|bibcode=1968RSPTB.254...37C|doi-access=}}{{cite journal|author=C. A. Clarke & P. M. Sheppard|year=1971|title=Further studies on the genetics of the mimetic butterfly Papilio memnon L.|journal=Philosophical Transactions of the Royal Society B|volume=263|issue=847|pages=35–70|jstor=2417186|doi=10.1098/rstb.1971.0109 |bibcode=1971RSPTB.263...35C|doi-access=}}{{cite journal|author=C. A. Clarke & P. M. Sheppard|year=1973|title=The genetics of four new forms of the mimetic butterfly Papilio memnon L.|journal=Proceedings of the Royal Society B: Biological Sciences |volume=184|issue=1074|pages=1–14|jstor=76137|bibcode=1973RSPSB.184....1C|doi=10.1098/rspb.1973.0027|s2cid=86397695}}

=Typical form ''agenor''=

Male. Tailless, above deep blue to black. It may or may not have red streak on the forewing at the base of the cell.
Female. Tailless. Upperside forewing ground colour sepia, streaked with greyish white. The basal third part of the cell is red and is touched outwardly with white. Upper hindwing is blue black. It has five to seven yellow or white discal patches.

Great Mormon, dorsal view.jpg|Male agenor

Papilio memnon-agenor female.jpg|Female agenor, Thailand

Great mormon (Papilio memnon agenor) female Tai Lam.jpg|Female agenor, Hong Kong

Great mormon (Papilio memnon agenor) female underside Tai Lam.jpg|Female agenor, Hong Kong

=Female form ''butlerianus''=

Tailless. Resembles the typical male. Both wings are dark sepia. The forewing has a white area on the inner margin. The hindwing is scaled with blue.

Open wing nectaring position of Papilio memnon Linnaeus, 1758 – Great Mormon (Female) Form butlerianusDSC 2479.jpg|Female butlerianus, India

Close wing Nectaring of Papilio memnon Linnaeus, 1758 – Great Mormon (Female) Form butlerianus WLB DSC 9451.jpg|Female butlerianus, India

=Female form ''alcanor''=

Tailed. The sides of the abdomen are yellow.
Upperside forewing greyish brown with veins and streaks between them black. The cell is red at the base. There is a velvety black patch at the bases of veins 1 and 2 of the upperside forewing.
Upperside hindwing is black with part of the cell white. There are white streaks around it. The tornus is red with a large black spot. There is a row of red terminal spots between the vein.

Papilio memnon-alcanor female dorsal.jpg|Female alcanor, Thailand

Papilio memnon-alcanor female ventral.jpg|Female alcanor, Thailand

Great mormon (Papilio memnon alcanor) female Tai Lam.jpg|Female alcanor, Hong Kong

Great mormon (Papilio memnon alcanor) female underside Tai Lam.jpg|Female alcanor, Hong Kong

=Male and female form ''polymnestoroides''=

Tailless.
Male. Upperside hindwing and forewing have short blue discal stripes.
Female. The upperside forewing is sepia with pale grey streaks amongst the veins. The base of the cell is red. The upperside hindwing is velvety brown with a blue discal area and has black spots, as in the case of the blue Mormon (P. polymnestor), which it mimics.

Habitat

This species flies up to {{convert|2100|m}} in the Himalayas, but is most common at low elevations.

Behaviour

This butterfly is found in forest clearings. It is very common and is also seen amongst human habitation. It visits flowers of Poinsettia, Jasminum, Lantana, Canna and Salvia. It usually flies {{convert|2|to|4|m}} above the ground. The butterfly is known to mud-puddle. The males are much more common than females. The female forms butlerianus and alcanor are especially uncommon.