Butterflyfish

File:Schooling bannerfish.JPG of false Moorish idols, Heniochus diphreutes|alt=Fish bearing two strong black stripes separated by one strong white stripe with long white tendril as dorsal fin]]

Butterflyfish mostly range from {{Convert|12|to|22|cm|abbr=on}} in length. The largest species, the lined butterflyfish and the saddle butterflyfish, C. ephippium, grow to {{Convert|30|cm|abbr=on}}. The common name references the brightly coloured and strikingly patterned bodies of many species, bearing shades of black, white, blue, red, orange, and yellow. Other species are dull in colour. Butterflyfish are a boundless, different group of marine percoids with delegates on practically all coral reef frameworks and in every single tropical ocean. Their bright and color patterns have drawn in much consideration, creating an abundance of data about their conduct and environment.Smith, W. (2003). The evolution of the laterophysic connection with a revised phylogeny and taxonomy of butterflyfishes (Teleostei: Chaetodontidae). Cladistics the International Journal of the Willi Hennig Society., 19(4), 287–306. https://doi.org/10.1016/S0748-3007(03)00070-7 Many have eyespots on their flanks and dark bands across their eyes, not unlike the patterns seen on butterfly wings.

{{cite book |editor=Paxton, J.R. |editor2=Eschmeyer, W.N.|author1=Johnson, G.D. |author2=Gill, A.C. |year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 184|isbn= 978-0-12-547665-2}} Their deep, laterally narrow bodies are easily noticed through the profusion of reef life. The conspicuous coloration of butterflyfish may be intended for interspecies communication. Butterflyfish have uninterrupted dorsal fins with tail fins that may be rounded or truncated, but are never forked.

Generally diurnal and frequenting waters less than {{Convert|18|m|abbr=on}} deep (though some species descend to {{Convert|180|m|abbr=on}}, butterflyfish stick to particular home ranges. These corallivores are especially territorial, forming pairs and staking claim to a specific coral head. Contrastingly, the zooplankton feeders form large conspecific groups. By night, butterflyfish hide in reef crevices and exhibit markedly different coloration.

Their coloration also makes them popular aquarium fish. However, most species feed on coral polyps and sea anemones. Balancing the relative populations of prey and predator is complex, leading hobby aquarists to focus on the few generalists and specialist zooplankton feeders.

Butterflyfish are pelagic spawners; that is, they release many buoyant eggs into the water, which become part of the plankton, floating with the currents until hatching. The fry go through a tholichthys stage, wherein the body of the postlarval fish is covered in large, bony plates extending from the head. They lose their bony plates as they mature. Only one other family of fish, the scats (Scatophagidae) express such an armored stage.

The Chaetodontidae can be, but are not usually, divided into two lineages that arguably are subfamilies. The subfamily name Chaetodontinae is a little-used leftover from the period when the Pomacanthidae and Chaetodontidae were united under the latter name as a single family. Hence, Chaetodontinae is today considered a junior synonym of Chaetodontidae. In any case, one lineage of Chaetodontidae (in the modern sense) contains the "typical" butterflyfish around Chaetodon, while the other unites the bannerfish and coralfish genera. As the Perciformes are highly paraphyletic, the precise relationships of the Chaetodontidae as a whole are badly resolved.Fessler & Westneat (2007)

Chaetodontidae is classified within the suborder Percoidei by the 5th edition of Fishes of the World, but they are placed in an unnamed clade which sits outside the superfamily Percoidea. This clade contains 7 families which appear to have some relationship to Acanthuroidei, Monodactylidae, and Priacanthidae.{{cite book |title=Fishes of the World |edition=5th |author1=J. S. Nelson |author2=T. C. Grande |author3=M. V. H. Wilson |year=2016 |pages=453 |publisher=Wiley |isbn=978-1-118-34233-6 |url=https://sites.google.com/site/fotw5th/ |access-date=2020-11-19 |archive-date=2019-04-08 |archive-url=https://web.archive.org/web/20190408194051/https://sites.google.com/site/fotw5th/ |url-status=dead }} Other authorities have paced the family in the order Chaetodontiformes alongside the family Leiognathidae.{{cite journal | author1 = R. Betancur-Rodriguez | author2 = E. Wiley | author3 = N. Bailly | author4 = A. Acero | author5 = M. Miya | author6 = G. Lecointre | author7 = G. Ortí | doi = 10.1186/s12862-017-0958-3 | title = Phylogenetic Classification of Bony Fishes – Version 4 | year = 2017 | journal = BMC Evolutionary Biology | volume = 17 | issue = 162| page = 162 | pmid = 28683774 | pmc = 5501477 | doi-access = free }} Presently, based on the most recent phylogenetic analyses, Eschmeyer's Catalog of Fishes places it in the large, diverse order Acanthuriformes.{{Cite web |last1=Fricke |first1=R. |last2=Eschmeyer |first2=W. N. |last3=Van der Laan |first3=R. |date=2025 |title=ESCHMEYER'S CATALOG OF FISHES: CLASSIFICATION |url=https://www.calacademy.org/eschmeyers-catalog-of-fishes-classification |access-date=2025-02-10 |website=California Academy of Sciences |language=en}} Molecular analyses suggest that the Chaetodontidae shared a common ancestor with the Scatophagidae and Pomacanthidae that lived during the Early Eocene (54 Ma).{{Cite journal |last1=Marrama' |first1=Giuseppe |last2=Giusberti |first2=Luca |last3=Carnevale |first3=Giorgio |date=2022-04-28 |title=A Rupelian Coral Reef Fish Assemblage from the Venetian Southern Alps (Berici Hills, Ne Italy) |journal=Rivista Italiana di Paleontologia e Stratigrafia |language=en |volume=128 |issue=2 |doi=10.54103/2039-4942/16601 |issn=2039-4942|doi-access=free }}

Before DNA sequencing, the taxonomy was confused about whether to treat these as species or subspecies. Also, numerous subgenera have been proposed for splitting out of Chaetodon, and it is becoming clear how to subdivide the genus if that is desired.Fessler & Westneat (2007), Hsu et al. (2007)

The fossil record of this group is disputed and marginal. Their restriction to coral reefs means their carcasses are liable to be dispersed by scavengers, overgrown by corals, and any that do fossilize will not long survive erosion. Many Eocene-aged fish such as Pygaeus have been variously referred to Chaetodontidae in the past; if used for calibration, it can be deduced that most living genera were probably distinct by the end of the Paleogene 23 Mya.Sepkoski (2002), Fessler & Westneat (2007) However, these Eocene genera cannot be reliably referred to Chaetodontidae.

The earliest confirmed fossil specimens of Chaetodontidae are an indeterminate larval specimen from Frauenweiler, Germany, two tholichthys specimens from the Menilite Formation of Poland, as well as the extinct species Chaetodon (Blumchaetodon) wattsi from Italy; all date to the Early Oligocene. The occurrence of the first confirmed butterflyfish in the Early Oligocene is supported by molecular phylogenies indicating that the family may have arose during the Late Eocene.

The bannerfish-coralfish lineage can be further divided in two groups; these might be considered tribes, but have not been formally named. Genera are listed in order of the presumed phylogeny, from the most ancient to the youngest:{{FishBase family|family=Chaetodontidae|month=February|year=2013}}

Bannerfish/coralfish lineage 1:

• Amphichaetodon Burgess, 1978
• Coradion Kaup, 1860
• Chelmon Cloquet, 1817
• Chelmonops Bleeker, 1876

Bannerfish/coralfish lineage 2:

• Forcipiger Jordan & McGregor, 1898
• Hemitaurichthys Bleeker, 1876
• Heniochus Cuvier, 1816
• Johnrandallia Nalbant, 1974

The "typical" butterflyfishes may eventually come to contain more genera; see Chaetodon:

• Chaetodon Linnaeus, 1758
• Parachaetodon Bleeker, 1874
• Prognathodes Gill, 1862
• Roa Jordan, 1923

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Image:Seattle Aquarium, 1.JPG|Copperband butterflyfish, Chelmon rostratus

Image:Johnrandallia nigrirostris.jpg|The enigmatic Johnrandallia nigrirostris

Image:Chaetodon kleinii.jpg|Sunburst butterflyfish, (sometimes placed in Lepidochaetodon)

Image:Bep chaetodon bennetti.jpg|Bluelashed butterflyfish, Chaetodon bennetti (sometimes placed in Megaprotodon'')

• Pratchett, Morgan S. & Berumen, Michael L. & Kapoor, B.G. [Editors] : Biology of Butterflyfishes. CRC Press, 2014. {{ISBN|978-1-4665-8290-3}}

{{Commons category|Chaetodontidae}}

;General

• {{Aut|Fessler, Jennifer L. & Westneat, Mark W.}} (2007): Molecular phylogenetics of the butterflyfishes (Chaetodontidae): Taxonomy and biogeography of a global coral reef fish family. Mol. Phylogenet. Evol. 45(1): 50–68. {{doi|10.1016/j.ympev.2007.05.018}} (HTML abstract)
• {{Aut|FishBase}} [2008]: [http://www.fishbase.org/Summary/FamilySummary.cfm?ID=343 Family Chaetodontidae – Butterflyfishes]. Retrieved 2008-SEP-02.
• {{Aut|Hsu, Kui-Ching; Chen, Jeng-Ping & Shao, Kwang-Tsao}} (2007): Molecular phylogeny of Chaetodon (Teleostei: Chaetodontidae) in the Indo-West Pacific: evolution in geminate species pairs and species groups. Raffles Bulletin of Zoology Supplement 14: 77–86. [http://rmbr.nus.edu.sg/rbz/biblio/s14/s14rbz10_Hsu-pp77-86.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20070811042056/http://rmbr.nus.edu.sg/rbz/biblio/s14/s14rbz10_Hsu-pp77-86.pdf |date=2007-08-11 }}
• {{Aut|Sepkoski, Jack}} (2002): [Chaetodon]. In: A compendium of fossil marine animal genera. Bulletins of American Paleontology 364: 560. [https://web.archive.org/web/20090220223520/http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class HTML database excerpt]

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Category:Articles which contain graphical timelines

Category:Taxa named by Constantine Samuel Rafinesque

Category:Extant Rupelian first appearances