Dove Spring Formation

{{Short description|Geologic formation in the Mojave Desert, California, U.S.}}

{{Infobox rockunit

| name = Dove Spring Formation

| image =

| caption =

| type = Geological formation

| age = Miocene
~{{Fossilrange|13.5|7.3}}

| period = Neogene

| prilithology =

| otherlithology =

| namedfor =

| namedby =

| region = California

| country = United States

| coordinates =

| unitof = Ricardo Group

| subunits =

| underlies =

| overlies = Cudahy Camp Formation

| thickness = 1,800 meters

| extent =

| area =

| map =

| map_caption =

}}

The Dove Spring Formation (formerly the Ricardo Formation) is a geologic formation in the western Mojave Desert of California.{{Cite journal |last1=Whistler |first1=David P. |last2=Burbank |first2=D. |date=1992 |title=Miocene biostratigraphy and biochronology of the Dove Spring Formation, Mojave Desert, California, and characterization of the Clarendonian mammal age (late Miocene) in California |journal=Geological Society of America Bulletin |language=en |volume=104 |issue=6 |pages=644–658|doi=10.1130/0016-7606(1992)104<0644:MBABOT>2.3.CO;2 |bibcode=1992GSAB..104..644W |s2cid=130575409 }} It preserves fossils dating back to the Miocene epoch of the Neogene period.

Fossil content

{{Paleobiota-key-compact}}

=Mammals=

==Bats==

class="wikitable" align="center"
colspan="6" align="center" | Bats reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Chiroptera

|Gen. et. sp. undetermined

| Kern County, California.

|

| A bat.

|

==Carnivorans==

class="wikitable" align="center"
colspan="6" align="center" | Carnivorans reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
rowspan="3" |Barbourofelis

|style="background:#E6E6E6;"|B. osborni

|style="background:#E6E6E6;"|Kern County, California.

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|Junior synonym of B. whitfordi.

|style="background:#E6E6E6;"|

B. whitfordi

|Red Canyon & other localities, Kern County, California.{{Cite journal |last1=Tseng |first1=Zhijie J. |last2=Takeuchi |first2=Gary T. |last3=Wang |first3=Xiaoming |date=2010-01-29 |title=Discovery of the upper dentition of Barbourofelis whitfordi (Nimravidae, Carnivora) and an evaluation of the genus in California |url=https://www.researchgate.net/publication/236961603 |journal=Journal of Vertebrate Paleontology |volume=30 |issue=1 |pages=244–254 |doi=10.1080/02724630903416001 |s2cid=21713508 |issn=0272-4634|doi-access=free }}

|Teeth & mandible.

|A barbourofelid.

|File:Barbourofelisdigitalreconstruction.jpg

B. sp.

|LACM localities 6260 & 1108, Kern County, California.

|Left partial humerus (LACM 140853) & left distal humerus (LACM 59336).

|A barbourofelid also found in the Green Valley Formation.

|

Bassariscus

|B. sp.

|Kern County, California.

|

|A procyonid.

|

Borophagus

|B. littoralis

|Red Rock Canyon, Kern County, California.{{Cite journal |last1=Wang |first1=Xiaoming |last2=Tedford |first2=Richard H. |last3=Taylor |first3=Beryl E. |date=1999 |title=Phylogenetic systematics of the Borophaginae (Carnivora, Canidae). |url=https://digitallibrary.amnh.org/handle/2246/1588 |journal=Bulletin of the AMNH |language=en-US |issue=243 |pages=9–391|hdl=2246/1588 }}

|Left maxillary (LACM 143520).

|A borophagine dog also found in the Green Valley, Santa Margarita & Chanac formations.

|File:Borophagus littoralis.jpg

Carpocyon

|C. robustus

|Kern County, California.

|Partial skeleton (UCMP 33569).

|A borophagine dog.

|

cf. Eomellivora

|cf. E. sp.

|Kern County, California.

|

|A large mustelid similar to the honey badger.

|File:Eomellivora piveteaui mandible - Batallones 3 fossil site, Torrejón de Velasco, Madrid, Spain.jpg

rowspan="3" |Epicyon

|style="background:#E6E6E6;"|E. aphobus

|style="background:#E6E6E6;"|Kern County, California.

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|Junior synonym of E. haydeni.

|style="background:#E6E6E6;"|

E. haydeni

|Ricardo Fauna, Kern County, California.

|Skull elements.

|A borophagine dog.

|File:Epicyon haydeni.jpg

E. saevus

|Ricardo Fauna, Kern County, California.

|Ramal elements.

|A borophagine dog.

|File:Epicyon saevus mandible.jpg

"Felis"

|"F." sp.

|Kern County, California.

|

|A small felid.

|

rowspan=2|Ischyrocyon

|I. gidleyi

|Last Chance Gulch, Kern County, California.{{Cite journal |last=Hunt |first=Robert |date=1998-01-01 |title=Amphicyonidae |url=https://digitalcommons.unl.edu/museummammalogy/275 |journal=Mammalogy Papers: University of Nebraska State Museum |pages=196–227}}

|

|A bear-dog.

|

style="background:#E6E6E6;"|I. mohavensis

|style="background:#E6E6E6;"|Kern County, California.

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|Junior synonym of I. gidleyi.

|style="background:#E6E6E6;"|

Leptocyon

|L. vafer

|Kern County, California (lower & upper parts of the formation).{{Cite book |last1=Tedford |first1=Richard H. |url=https://digitallibrary.amnh.org/handle/2246/5999 |title=Phylogenetic systematics of the North American fossil Caninae (Carnivora, Canidae). (Bulletin of the American Museum of Natural History, no. 325) |last2=Wang |first2=Xiaoming |last3=Taylor |first3=Beryl E. |date=2009 |publisher=[New York] : American Museum of Natural History. |hdl=2246/5999 |language=en-US}}

|Skull elements.

|A canine dog.

|File:Leptocyon head restoration.jpg

rowspan=2|Martinogale

|M. faulli

|Red Rock Canyon, Kern County, California.{{Cite journal |last1=Wang |first1=Xiaoming |last2=Whistler |first2=David P. |last3=Takeuchi |first3=Gary T. |date=2005 |title=A New Basal Skunk Martinogale (Carnivora, Mephitinae) from Late Miocene Dove Spring Formation, California, and Origin of New World Mephitines |url=https://www.jstor.org/stable/4524520 |journal=Journal of Vertebrate Paleontology |volume=25 |issue=4 |pages=936–949 |doi=10.1671/0272-4634(2005)025[0936:ANBSMC]2.0.CO;2 |jstor=4524520 |s2cid=21747985 |issn=0272-4634|url-access=subscription }}

|A nearly-complete skull (LACM 56230).

|A skunk.

|

M. sp.

|Kern County, California.

|

|A skunk.

|

"Mephitis"

|"M." sp.

|Kern County, California.

|

|A skunk.

|File:Striped Skunk - Desert Museum - Tucson - AZ - 2015-10-12at10-56-241 (22241714622).jpg

Metalopex

|M. macconnelli

|Red Rock Canyon, Kern County, California.

|Skull elements.

|A fox also found in the Milk Creek Formation & Malheur County, Oregon.

|

Mustela

|?M. buwaldi

|Kern County, California.

|

|A weasel.

|

style="background:#E6E6E6;"|Osteoborus

|style="background:#E6E6E6;"|O. diabloensis

|style="background:#E6E6E6;"|Kern County, California.

|style="background:#E6E6E6;"|

|style="background:#E6E6E6;"|Junior synonym of Borophagus littoralis.

|style="background:#E6E6E6;"|

Pseudaelurus

|P. sp.

|Kern County, California.

|

|A felid.

|

rowspan=2|Tomarctus

|style="background:#fbdddb;"|T. robustus

|style="background:#fbdddb;"|Kern County, California.

|style="background:#fbdddb;"|

|style="background:#fbdddb;"|Moved to the genus Carpocyon.

|style="background:#fbdddb;"|

T. sp.

|Kern County, California.

|

|A borophagine dog.

|

"Vulpes"

|"V." sp.

|Kern County, California.

|

|A fox.

|

==Eulipotyphlans==

class="wikitable" align="center"
colspan="6" align="center" | Eulipotyphlans reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Alluvisorex

|A. chasseae

|Kern County, California.

|

|A shrew.

|

Erinaceidae

|Erinaceidae "A"

|Kern County, California.

|

|An erinaceid.

|

Lanthanotherium

|L. sp.

| Kern County, California.

|

| An erinaceid.

|

Limnoecus

|L. sp.

|Kern County, California.

|

|A shrew.

|

Scapanus

|S. (Xeroscapheus) shultzi

|Kern County, California.

|

|A mole.

|

"Sorex"

|"S." sp.

|Kern County, California.

|

|A shrew.

|

==Lagomorphs==

class="wikitable" align="center"
colspan="6" align="center" | Lagomorphs reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Hesperolagomys

|H. sp.

|Kern County, California.

|

|A pika.

|

Hypolagus

|H. sp.

|Kern County, California.

|

|A leporid.

|File:Hypolagus peregrinus.jpg

==Proboscideans==

class="wikitable" align="center"
colspan="6" align="center" | Proboscideans reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Gomphotherium

|G. sp.

|Kern County, California.

|

|A gomphothere.

|

Serbelodon

|S. burnhami

|Ricardo, Kern County, California.{{Cite journal |last1=Osborn |first1=Henry Fairfield |last2=Blick |first2=John C. |last3=Frick |first3=Childs |date=June 1933 |title=Serbelodon burnhami, a new shovel-tusker from California. |url=https://digitallibrary.amnh.org/handle/2246/2061 |journal=American Museum Novitates |language=en-US |issue=639 |pages=1–5|hdl=2246/2061 }}

|Anterior portion of a mandible (F:AM 18228) & 2 tusk fragments (F:AM 18228A & 18228B).

|An amebelodontid.

|

==Rodents==

class="wikitable" align="center"
colspan="6" align="center" | Rodents reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Acrolophomys

|A. rhodopetros

|

|Teeth & dentary.{{cite journal |author1=Thomas S. Kelly |author2=David P. Whistler |year=2014 |title=New late Miocene (latest Clarendonian to early Hemphillian) cricetid rodents from the upper part of the Dove Spring Formation, Mojave Desert, California |journal=Paludicola |volume=10 |issue=1 |pages=1–48 |url=https://paludicolavertpaleo.files.wordpress.com/2018/05/10-1-kelly-and-whistler-2014.pdf }}

|A cricetid.

|

Ammospermophilus

|A. sp.

|Kern County, California.

|

|An antelope squirrel.

|

Antecalomys

|A. coxae

|

|Many teeth.

|A sigmodontine.

|

Bensonomys

|B. sp.

|Kern County, California.

|Maxillae, a dentary & teeth.

|A sigmodontine.

|

rowspan="5" |Copemys

|C. dentalis

|Kern County, California.

|Dentaries & teeth.

|A cricetid.

|

C. cf. C. longidens

|Kern County, California.

|

|A cricetid.

|

C. russelli

|Kern County, California.

|

|A cricetid.

|

C. sp.

|Kern County, California.

|

|A cricetid.

|

C. sp., cf. C. dentalis

|

|Teeth & jaw elements.

|A cricetid.

|

rowspan="3" |Cupidinimus

|C. avawatzensis

|Kern County, California.

|

|A heteromyid also found in the Avawatz Formation.

|

C. tertius

|Kern County, California.

|

|A heteromyid.

|

C. sp.

|Kern County, California.

|

|A small heteromyid.

|

Eucastor

|E. sp.

|Kern County, California.

|

|A castorid.

|

Leptodontomys

|L. sp.

|Kern County, California.

|

|An eomyid.

|

rowspan="4" |Lindsaymys

|L. takeuchii

|Kern County, California.

|Multiple specimens.

|A cotton rat.

|

L. sp., cf. L. takeuchii

|Kern County, California.

|Multiple teeth.

|A cotton rat.

|

L. sp. A

|Kern County, California.

|Molars.

|A cotton rat.

|

L. sp. B

|Kern County, California.

|Molars.

|A cotton rat.

|

Parapliosaccomys

|P. sp.

|Kern County, California.

|

|A gopher.

|

Paronychomys

|P. spp.

|Kern County, California.

|

|A cricetid.

|

rowspan=2|Perognathus

|P. furlongi

|Kern County, California.

|

|A pocket-mouse.

|

P. minutus

|Kern County, California.

|

|A pocket-mouse.

|

Postcopemys

|P. sp., cf. P. valensis

|

|Teeth & dentary.

|A cricetid.

|

Protospermophilus

|P. sp.

|Kern County, California.

|

|A squirrel.

|

rowspan=2|Repomys

|"R." sp.

|Kern County, California.

|

|A cricetid.

|

cf. R. sp.

|

|M2 (LACM 156378).

|A cricetid.

|

Tamias

|T. ateles

|Kern County, California.

|

|A chipmunk.

|

"Thomomys"

|"T." sp.

|Kern County, California.

|

|A gopher.

|

==Ungulates==

class="wikitable" align="center"
colspan="6" align="center" | Ungulates reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Aphelops

|A. sp.

|Kern County, California.

|

|A rhinoceros.

|

rowspan=2|Cormohipparion

|style="background:#fbdddb;" |C. occidentale

|style="background:#fbdddb;" |Kern County, California.

|style="background:#fbdddb;" |Isolated teeth & a lower mandible.{{Cite journal |last=Woodburne |first=Michael O. |date=2007 |title=Phyletic diversification of the Cormohipparion occidentale complex (Mammalia, Perissodactyla, Equidae), late Miocene, North America, and the origin of the Old World Hippotherium datum ; Bulletin of the American Museum of Natural History, no. 306 |hdl=2246/5872 |url=https://digitallibrary.amnh.org/handle/2246/5872 |language=en-US}}

|style="background:#fbdddb;" |Remains reassigned to C. sp.

|style="background:#fbdddb;" |

C. sp.

| El Paso Basin.

|Isolated teeth & a lower mandible.

| An equid.

|File:Cormohipparion occidentale.jpg

Cosoryx

|C. sp.

|Kern County, California.

|

|An antilocaprid.

|

Dinohippus

|D. cf. D. leardi

|Kern County, California.

|

|An equid.

|

?Hemiauchenia

|?H. sp.

|Kern County, California.

|

|A camelid.

|

rowspan=2|Hipparion

|H. forcei

|Kern County, California.

|

|An equid.

|

H. tejonensis

|Kern County, California.

|

|An equid.

|

Ilingoceros

|I. sp.

|Kern County, California.

|

|An antilocaprid.

|File:Ilingoceros.jpg

Megahippus

|M. cf. M. matthewi

|Kern County, California.

|

|An equid.

|File:Megahippus sp.JPG

?Megatylopus

|?M. sp.

|Kern County, California.

|

|A camelid.

|File:Titanotylopus.JPG

?Michenia

|?M. sp.

|Kern County, California.

|

|A camelid.

|

Paracosoryx

|P. furlongi

|Kern County, California.{{Cite book |last1=Prothero |first1=Donald R. |url=https://books.google.com/books?id=qO8H_alEofAC&dq=paracosoryx+furlongi&pg=PA229 |title=The Evolution of Artiodactyls |last2=Foss |first2=Scott E. |date=2007-12-17 |publisher=JHU Press |isbn=978-0-8018-8735-2 |pages=229 |language=en}}

|

|An antilocaprid.

|

Peraceras

|P. sp.

|Kern County, California.

|

|A rhinoceros.

|File:Peraceras superciliosus.jpg

Plioceros

|P. sp.

|Kern County, California.

|

|An antilocaprid.

|

rowspan=2|Pliohippus

|P. tantulus

|Kern County, California.

|

|An equid.

|

P. tejonensis

|Kern County, California.

|

|An equid.

|

?Procamelus

|?P. sp.

|Kern County, California.

|

|A camelid.

|File:Procamelus AMNH.jpg

?Prosthennops

|?P. sp.

|Kern County, California.

|

|A peccary.

|

Teleoceras

|T. cf. T. meridianum

|Kern County, California.

|

|A rhinoceros.

|File:Teleoceras.jpg

rowspan=2|Ustatochoerus

|U. californicus

|Kern County, California.

|

|A merycoidodontid.

|

U. cf. U. profectus

|Kern County, California.

|

|A merycoidodontid.

|

=Reptiles=

==Birds==

class="wikitable" align="center"
colspan="6" align="center" | Birds reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Branta

|B. howardae

| Kern County, California.

|

| A goose.

|

Neophrontops

|N. ricardoensis

|Kern County, California.

|

|A gypaetine vulture.

|

==Squamates==

class="wikitable" align="center"
colspan="6" align="center" | Squamates reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Anguidae

|Gen. indet.

|Kern County, California.

|

|A very large anguid lizard.

|

Callisaurus

|C. sp.

|Kern County, California.

|

|A zebra-tailed lizard.

|File:ZebraTailed-Lizard-640px.jpg

Gerrhonotus

|G. cf. G. kingi

|Kern County, California.

|

|An alligator lizard.

|

Lichanura

|L. sp.

|Kern County, California.

|

|A rosy boa.

|File:Lichanura orcutti.jpg

Paleoheterodon

|P. sp.

|Kern County, California.

|

|A colubrid snake.

|

Paracoluber

|P. sp.

|Kern County, California.

|

|A colubrid snake.

|

Paragerrhonotus

|P. ricardensis

|Kern County, California.

|

|An anguid lizard.

|

Proptychophis

|P. achoris

|Red Rock Canyon State Park, Kern County, California.{{Cite journal |last1=Whistler |first1=David P. |last2=Wright |first2=John W. |date=1989 |title=A Late Miocene Rear-Fanged Colubrid Snake from California with Comments on the Phylogeny of North American Snakes |url=https://www.jstor.org/stable/3892892 |journal=Herpetologica |volume=45 |issue=3 |pages=350–367 |jstor=3892892 |issn=0018-0831}}

|Numerous vertebrae & the posterior portion of a right maxilla.

|A colubrid snake.

|

Sceloporus

|S. sp.

|Kern County, California.

|

|A large spiny lizard.

|File:Sceloporus variabilis variabilis, Rose-bellied Lizard, Tamaulipas.jpg

"Thamnophis"

|"T." sp.

|Kern County, California.

|

|A garter snake.

|File:Giant Garter Snake 1.jpg

Uma

|U. sp.

|Locality LACM 4702.{{Cite journal |last=Scarpetta |first=Simon G. |date=2019-09-06 |title=The first known fossil Uma: ecological evolution and the origins of North American fringe-toed lizards |journal=BMC Evolutionary Biology |volume=19 |issue=1 |pages=178 |doi=10.1186/s12862-019-1501-5 |issn=1471-2148 |pmc=6729053 |pmid=31492110 |doi-access=free }}

|A partial & fused premaxilla (LACM 159892).

|A fringe-toed lizard.

|File:Uma-scoparia.jpg

?Uta

|?U. sp.

|Kern County, California.

|

|A side-blotched lizard.

|File:Joshua Tree NP - Desert Side-blotched Lizard - 1.jpg

Xantusia

|X. sp.

|Kern County, California.

|

|A night lizard.

|File:Xantusia henshawi.jpg

==Testudines==

class="wikitable" align="center"
colspan="6" align="center" | Testudines reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Clemmys

|C. sp.

|Kern County, California.

|

|An emydine turtle.

|File:Spotted Turtle (Clemmys guttata) (captive specimen) (36331198042).jpg

Geochelone

|G. sp.

|Kern County, California.

|

|A tortoise.

|

?Gopherus

|?G. sp.

|Kern County, California.

|

|A gopher tortoise.

|File:Gopherus polyphemus Tomfriedel.jpg

=Amphibians=

class="wikitable" align="center"
colspan="6" align="center" | Amphibians reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Batrachoseps

|B. sp.

| Kern County, California.

|

| A lungless salamander.

|

Bufonidae

|

|Kern County, California.

|

|A true toad.

|

cf. Ensatina

|cf. E. sp.

|Kern County, California.

|

|A lungless salamander.

|

Ranidae

|

|Kern County, California.

|

|A true frog.

|

=Fish=

class="wikitable" align="center"
colspan="6" align="center" | Fish reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Empetrichthys

|E. sp.

| Kern County, California.

|

| A splitfin.

|

=Plants=

class="wikitable" align="center"
colspan="6" align="center" | Plants reported from the Dove Spring Formation
GenusSpeciesPresence

!Material

NotesImages
Acacia

|A. sp.

|Kern County, California.

|

|An acacia.

|

Ceanothus

|C. sp.

|Kern County, California.

|

|A buckbrush.

|

Cupressus

|C. sp.

|Kern County, California.

|

|A cypress tree.

|

Lycium

|L. sp.

|Kern County, California.

|

|A box thorn.

|

Palmoxylodon

|P. mohavensis

| Kern County, California.

|

| A palm.

|

Pinus

|P. kelloggii

|Kern County, California.

|

|A pine.

|

Quercus

|Q. ricardensis

|Kern County, California.

|

|An oak tree.

|

Robinia

|R. alexanderi

|Kern County, California.

|

|A legume.

|

See also

References

{{Reflist}}

  • {{cite web|title= Fossilworks: Gateway to the Paleobiology Database|author= ((Various Contributors to the Paleobiology Database))|url= https://www.fossilworks.org|access-date= 17 December 2021}}

Category:Neogene California