Hasle Formation

{{Short description|Geologic formation on the island on Bornholm, Denmark}}

{{distinguish|Hasler Formation}}

{{infobox rockunit

| name = Hasle Formation

| image = HasleSandstone.jpg

| caption = Outcrop (Yellow layers)

| age = Lower to Middle Pliensbachian
~{{fossil range|189|184}}Jamesoni–Margaritatus Chronozone

| period = Pliensbachian

| unitof = Bornholm Group

| overlies = Rønne Formation

| underlies = Sorthat Formation

| thickness = {{convert|80|-|140|m|ft|abbr=on}}

| prilithology = Siltstone, sandstone

| namedfor = Hasle, Bornholm

| namedby = Gry{{cite journal |last1=Gry |first1=H. |last2=Jørgart |first2=T. |last3=Poulsen |first3=V. |title=Lithostratigraphy and sedimentary evolution of the Triassic, Jurassic and Lower Cretaceous of Bornholm, Denmark |journal=Mineralogisk Mus. |date=1969 |volume=6 |issue=1 |access-date=}}

| year_ts = 1969

| region = Bornholm

| country = Denmark

| coordinates = {{coord|55.2|N|14.7|E|display=inline,title}}

| paleocoordinates = {{coord|45.0|N|18.5|E|display=inline}}

| map = {{Location map+ | Denmark

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Denmark

| lat_deg = 55.2

| lon_deg = 14.7

| mark = Blue pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

The Hasle Formation is a geologic formation on the island on Bornholm, Denmark. It is of early to late Pliensbachian age. Vertebrate fossils have been uncovered from this formation.{{cite journal|last=Rees|first=J.|title=Early Jurassic selachians from the Hasle Formation on Bornholm, Denmark|journal=Acta Palaeontologica Polonica|year=1998|series=43|issue=3|pages=43–439|url=http://www.app.pan.pl/archive/published/app43/app43-439.pdf}}{{cite journal|last1=Donovan |first1=Desmond T. |last2=Surlyk |first2=Finn|title=Lower Jurassic (Pliensbachian) ammonites from Bornholm, Baltic Sea, Denmark|journal=Geological Survey of Denmark and Greenland Bulletin|series=1|pages=555–583|url=http://www.geus.dk/publications/bull/nr1_p555-583.pdf}} The type section of the formation is found at the south of the costal Hasle Town, and it is composed by rusty yellow to brownish siltstones and very fine-grained sandstones. The southernmost arch, Hvjdoddebuen, is not as fossil-bearing as the type unit in Hasle.{{cite journal |last1=Malling |first1=C. |title=De Jespersenske Buelag i Lias paa Bornholm |journal=Meddelelser Fra Dansk Geologisk Forening) |date=1914 |volume=4 |issue=1 |pages=265–270 |url=https://2dgf.dk/xpdf/bull-1914-4-3-265-270.pdf |access-date=20 October 2021}} The formation can be separated in two different petrographic types: type 1 sandstones are friable with layers and lenses of concretionary siderite and type 2 well-cemented sandstones.{{cite journal |last1=Larsen |first1=Ole Hede |last2=Friis |first2=Henrik |title=Petrography, diagenesis and pore-water evolution of a shallow marine sandstone (Hasle Formation, Lower Jurassic, Bornholm, Denmark) |journal=Sedimentary Geology |date=1991 |volume=72 |issue=3–4 |pages=269–284 |doi=10.1016/0037-0738(91)90015-6 |bibcode=1991SedG...72..269L |url=https://www.sciencedirect.com/science/article/abs/pii/0037073891900156 |access-date=6 October 2021|url-access=subscription }} Both types where deposited in a relatively high-energy marine environment with a diagenetic pattern that demonstrates a close relation to various phases of subsidence and uplift in the tectonically unstable Fennoscandian Border Zone.

Most of its deposition happened on a storm-dominated shoreface, with the exposed parts deposited in an open marine shelf within 1–2 km distance from the fault-controlled coastlines.{{cite journal |last1=Michelsen |first1=O. |last2=Nielsen |first2=L. H. |last3=Johannessen |first3=P. N. |last4=Andsbjerg |first4=J. |last5=Surlyk |first5=F. |title=Jurassic lithostratigraphy and stratigraphic development onshore and offshore Denmark |journal=Geological Survey of Denmark and Greenland (GEUS) Bulletin |date=2003 |volume=1 |issue=1 |pages=145–216 |doi=10.34194/geusb.v1.4651 |s2cid=126907584 |doi-access=free}} However, recent works have recovered terrestrial fauna from it, including a footprint, suggesting easterly winds and low tide could have exposed the inner parts of the upper shoreface, and create long-lasting Floodplain-type environments. Field works since 1984 have shown a mostly hummocky cross-stratified deposition, with great complexity of the sediments that suggests very complicated and variable flow conditions, with Megaripples derived from storm events.{{cite journal |last1=Surlyk |first1=Finn |last2=Noe-Nygaard |first2=Nanna |title=Hummocky cross-stratification from the Lower Jurassic Hasle Formation of Bornholm, Denmark |journal=Sediment. Geol. |date=1986 |volume=46 |issue=3–4 |pages=259–273 |doi=10.1016/0037-0738(86)90062-X |bibcode=1986SedG...46..259S |url=https://www.sciencedirect.com/science/article/abs/pii/003707388690062X |access-date=6 October 2021|url-access=subscription }} Storms were frequent and the coastline faced a wide epeiric sea with a fetch towards the west of possibly 1000 kilometers. The Jamesoni–Ibex Chronozone in the Central European Basin represents a clear sea Transgression, due to the appearance of ammonites from Thuringia and southern Lower Saxony, showing a full marine ingression towards the west.{{cite journal |last1=Barth |first1=G. |last2=Pieńkowski |first2=G. |last3=Zimmermann |first3=J. |last4=Franz |first4=M. |last5=Kuhlmann |first5=G. |title=Palaeogeographical evolution of the Lower Jurassic: high-resolution biostratigraphy and sequence stratigraphy in the Central European Basin |journal=Geological Society, London, Special Publications |date=2018 |volume=469 |issue=1 |pages=341–369 |doi=10.1144/SP469.8 |bibcode=2018GSLSP.469..341B |s2cid=134043668 |url=https://www.researchgate.net/publication/322268601 |access-date=8 September 2021}} This rise in the sea level is also measured in the north, as is proven by the presence of Uptonia jamesoni in Kurremölla (Röddinge Formation, Skåne) and Beaniceras centaurus plus

Phricodoceras taylori on the Hasle Formation. The whole Hasle Sandstones are a result of this rise in the sea level, where the marine sediments cover the deltaic layers of the Rønne Formation. The rise in the sea level is observed on palynology, as on the Hasle Formation Nannoceratopsis senex (Dinoflajellate) and Mendicodinium reticulaturn (Algae Acritarch) appear, indicating a transition from paralic and restricted marine to fully marine.{{cite journal |last1=Surlyk |first1=F. |last2=Arndorff |first2=L. |last3=Hamann |first3=N. E. |last4=Hamberg |first4=L. |last5=Johannessen |first5=P. N. |last6=Koppelhus |first6=E. B. |last7=Petersen |first7=H. I. |title=High-resolution sequence stratigraphy of a Hettangian-Sinemurian paralic succession, Bornholm, Denmark |journal=Sedimentology |date=1995 |volume=42 |issue=2 |pages=323–354|doi=10.1111/j.1365-3091.1995.tb02105.x |bibcode=1995Sedim..42..323S }}

Fossil content

= Annelida =

class = "wikitable"
Genus

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! Material

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Pentaditrupa{{cite journal |last1=Malling |first1=C. |last2=Grönwall |first2=K. A. |title=En fauna i Bornholms Lias |journal=Geologiska Föreningen i Stockholm Förhandlingar |date=1912 |volume=34 |issue=3 |pages=366–367 |doi=10.1080/11035891209448698 |url=https://zenodo.org/record/2214689 }}

|

  • P. quinquesulcata

|

  • Stampeå stream

|

Two tubes

|

A sessile, marine annelid tube worm of the family Serpulidae

|

Serpula?{{Cite journal|last1=Kočí |first1=T. |last2=Milàn |first2=J. |last3=Jakobsen |first3=S. L. |last4=Bashforth |first4=A. R. |year=2024 |title=Serpula? alicecooperi sp. nov. – a new serpulid from the Lower Jurassic (Pliensbachian) Hasle Formation of Bornholm, Denmark |journal=Bulletin of the Geological Society of Denmark |volume=73 |pages=41–56 |doi=10.37570/bgsd-2024-73-02 |doi-access=free }}

|

  • S.? alicecooperi

|

  • Stampeå stream

|

One well-preserved tube and several fragmentary tubes

|

A sessile, marine annelid tube worm of the family Serpulidae.

|

File:Serpula vermicularis 2 Linnaeus, 1767.jpg

=Brachiopoda=

class = "wikitable"
Genus

! Species

! Location

! Material

! Notes

! Images

Rhynchonella{{cite journal |last1=Malling |first1=C. |last2=Grönwall |first2=K. A. |title=En fauna i Bornholms Lias |journal=Meddel. Dansk Geol. Foren. |date=1909 |volume=3 |issue=1 |pages=271–308 |url=https://docplayer.dk/3279236-En-fauna-i-bornholms-lias.html |access-date=6 October 2021}}{{cite journal |last1=Malling |first1=C. |title=Hasle-Sandstenens alder |journal=Meddelelser Fra Dansk Geologisk Forening |date=1911 |volume=3 |issue=1 |pages=629–631 |url=https://2dgf.dk/xpdf/bull-1911-3-4-619-633.pdf?fbclid=IwAR3YmILcYUKW_fJ3jiaNS-N0KOCgYOfv2hn5Ng0GkynrSH4EcQNP0tIXFOM |access-date=20 October 2021}}

|

  • R. jurensis
  • R. variabilis

|

  • Stampeå stream
  • South of Hasle

|Shells

|

A saltwater Brachiopodan, member of Rhynchonellata inside Rhynchonellida. Found associated with Plicatula on long-term well-oxygenated conditions within the substrate and bottom waters.

|

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Waldheimia

|

  • W. sp.

|

  • Stampeå stream

|Shells

|

A Saltwater lamp shell Brachiopodan, member of Terebratulidae inside Terebratulida. The specimens are rather incomplete. The genus is a possible junior synonym of Terebratula.

|

File:FMIB 48838 Waldheimia (Magellania) flavascens.jpeg

=Bivalvia=

class="wikitable" align="center"
Genus

! Species

! Location

! Material

! Notes

! Images

Astarte

|

  • A. obsoleta
  • A. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Clam, type member of the family Astartidae inside Carditida. It is found on small, smooth Clam associations, with the exception of one does section, with less well-preserved stone cores and imprints.

|

155px

Avicula

|

  • A. inaequivalvis
  • A. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater pearl oyster, member of the family Pteriidae inside Ostreida. Some specimens are big and found associated with Burrow-filled surfaces. This well-known species occurs extremely frequently; Copies occur at all ages, as well as right-handers Left shells have been found.

|

Cardinia

|

  • C. multicostatum
  • C. crassiuscula
  • C. infera

|

  • Stampeå stream
  • South of Hasle
  • Hvideoddebuen
  • Nebbeodde

|

  • Shells

|

A saltwater Clam, type member of the family Cardiidae inside Carditida. Is a frequently occurring Clam.

|

155px

Cucullaea

|

  • C. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Clam, type member of the family Cucullaeidae inside Arcida.

|

Cyprina

|

  • C. cardioides

|

  • Stampeå stream
  • Hvideoddebuen
  • Nebbeodde

|

  • Shells

|

A saltwater Clam, member of the family Arcticidae inside Veneroidei. Is a frequently occurring Clam.

|

Cypricardia

|

  • C. cf.laevigata
  • C. cf.tetragona

|

  • South of Hasle

|

  • Shells

|

A saltwater Clam, member of the family Trapezidae inside Veneroidei.

|

Dendostrea

|

  • D. sandvichensis

|

  • Stampeå stream

|

  • Shells

|

A saltwater Oyster, member of the family Ostreinae inside Ostreida. Identified originally as Ostrea semiplicata

|

Gervilleia

|

  • G. sjögreni
  • G. cf. hagenovii
  • G. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Oyster, member of the family Bakevelliidae inside Ostreida. Found on a series of incomplete Stone Cores filled with specimens, that by Size and Form can be possibly attributable to Gervilleia aerosa.

|

155px

Hinnites

|

  • H. tumidus

|

  • Stampeå stream

|

  • Shells

|

A saltwater Scallop, member of the family Pectinoidae inside Pectinida. A Pair of incomplete specimens of a Hinnites are known and probably match with the species Hinnites tumidus; however, the state of preservation is bad, with the assignment rather uncertain.

|

155px

Lima

|

  • L. succincta

|

  • Stampeå stream

|

  • Shells

|

A saltwater file Clam, type member of the family Limidae inside Pectinida.

|

155px

Limea

|

  • L. acuticosfa

|

  • Stampeå stream

|

  • Shells

|

A saltwater file Clam, member of the family Limidae inside Pectinida. Mostly know due to a single young individual.

|

155px

Luciniola

|

  • L. pumila

|

  • Stampeå stream

|

  • Shells

|

A saltwater Clam, type member of the family Lucinidae inside Lucinida.

|

Modiolus

|

  • M. scalprum
  • M. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Mussel, type member of the family Mytilidae inside Mytilida.

|

Myoconcha

|

  • M. stampensis
  • M. jespersenii

|

  • Stampeå stream

|

  • Shells

|

A saltwater Clam, member of the family Kalenteridae inside Cardiida. One of the most frequent and characteristic Fossils in the formation

|

155px

Mytilus

|

  • M. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Mussel, type member of the family Mytilidae inside Mytilida.

|

Nucula

|

  • N. distinguenda
  • N. (Leda?) dr. omaliusi?
  • N. pinguis

|

  • Stampeå stream

|

  • Shells

|

A saltwater nut Clam, type member of the family Nuculidae inside Nuculida. Two copies of a Nucula (leda?) so closely in accordance with Moberg's depictions and description of occurring at Kurremolla.

|

155px

Nuculana

|

  • N. bornholmiensis
  • N. subovalis
  • N. complanata

|

  • Stampeå stream

|

  • Shells

|

A saltwater pointed nut Clam, type member of the family Nuculanidae inside Nuculida. Nuculana bornholmiensis is a very common genus only found locally.

|

155px

Palaeoneilo

|

  • P. bornholmiensis
  • P. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater pointed nut Clam, type member of the family Nuculanidae inside Nuculida.

|

Parallelodon

|

  • P. cypriniformis
  • P. pullus

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Clam, type member of the family Parallelodontidae inside Arcida. This species occurs frequently and varies a lot

|

Pecten

|

  • P. priscus
  • P. aequivalvis

|

  • Stampeå stream
  • South of Hasle
  • Hvideoddebuen
  • Nebbeodde

|

  • Shells

|

A saltwater Scallop, type member of the family Pectinoidae inside Pectinida. This species has been exceedingly frequent, and numerous Fragments are available.

|

155px

Perna

|

  • P. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Mussel, member of the family Mytilidae inside Mytilida.

|

Pleuromya

|

  • P. forchhammeri
  • P. dunkeri

|

  • Stampeå stream
  • South of Hasle
  • Hvideoddebuen
  • Nebbeodde

|

  • Shells

|

A saltwater Clam, type member of the family Pleuromyidae inside Pholadida. Three specimens of this species have been found.

|

Plicatula

|

  • P. spinosa

|

  • Stampeå stream

|

  • Shells

|

A saltwater Scallop, type member of the family Plicatulidae inside Pectinida. Two poorly preserved specimens have been found.

|

Spondylus

|

  • S. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Scallop, member of the family Pectinoidae inside Pectinida.

|

155px

Tancredia

|

  • T. lineata
  • T. elegans
  • T. tenera
  • T. johnstrupi

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Clam, type member of the family Tancrediidae inside Carditida. Tancredia lineata Is based on six shell cores, of which one complete, the others a little defective at the front end.

|

Tellina

|

  • T. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater tellin Clam, member of the family Tellinidae inside Cardiida.

|

155px

=Scaphopoda=

class="wikitable" align="center"
Genus

! Species

! Location

! Material

! Notes

! Images

Laevidentalium

|

  • L. etalense
  • L. elongatum
  • L. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater tusk shell (Scaphopoda), member of the family Dentaliidae inside Dentaliida

|

155px

=Gastropoda=

class="wikitable" align="center"
Genus

! Species

! Location

! Material

! Notes

! Images

Actaeonina

|

  • A. nathorsti

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Snail, type member of the family Acteoninidae inside Prosobranchia. Of this small species there are quite a few specimens, both stone cores and imprints that perfectly allow identifying the species.

|

Amberlya

|

  • A. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater Snail, type member of the family Spirostylidae inside Murchisoniina.

|

Chemnitzia

|

  • C.? undulata
  • C. craticia
  • C. citharella
  • C. blainvillei

|

  • Stampeå stream

|

  • Shells

|

A saltwater Snail, member of the family Pyramidellidae inside Pyramidelloidea. Two Specimens of a Turritella-like Gastropod found locally have been refer to the Genus. Chemnitzia citharella includes ten specimens, the largest of the genus found locally. However, most of the specimens are shell cores.

|

155px

Eucyclus

|

  • E. sp.

|

  • Stampeå stream

|

  • Shells

|

A saltwater Snail, type member of the family Eucyclidae inside Seguenzioidea. The material available is rather poor.

|

Margarites

|

  • M. solarium

|

  • Stampeå stream

|

  • Shells

|

A saltwater Snail, member of the family Turbinidae inside Turbinoidea. Occurs frequently, especially on marine ingression surfaces.

|

Natica

|

  • N. sp.

|

  • South of Hasle

|

  • Shells

|

A saltwater moon Snail, type member of the family Naticidae inside Naticoidea.

|

155px

Neritina

|

  • N. cf.liasina

|

  • South of Hasle

|

  • Shells

|

A saltwater Snail, type member of the family Neritidae inside Neritoidea.

|

155px

Pleurotomaria

|

  • P. expansa
  • P. sp.

|

  • Stampeå stream
  • South of Hasle

|

  • Shells

|

A saltwater Snail, type member of the family Pleurotomariidae inside Pleurotomarioidea.

|

155px

Trochus

|

  • T. laevis
  • T. subsulcatus

|

  • Stampeå stream

|

  • Shells

|

A saltwater top Snail, type member of the family Trochidae inside Trochoidea. Rare compared with other species found locally.

|

=Belemnites=

class="wikitable" align="center"
Genus

! Species

! Location

! Material

! Notes

! Images

Nannobelus

|

  • N. acutus

|

  • Stampeå stream
  • South of Hasle

|

  • Phragmocones

|

A belemnite, incertae sedis member of the family Belemnitida. Frequently shows the embryonic chamber preserved as a bladder, slightly bent forward-towards the ventral side.

|

File:Nannobelus acutus.1 - Jurasico.JPG

Subhastites

|

  • S. paxillosus
  • S. sp.

|

  • Stampeå stream

|

  • Phragmocones

|

A Belemnite, member of the family Hastitidae inside Belemnitida.

|

= Ammonites =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Acanthopleuroceras?

|

A.? sp. indet.

| rowspan="24" |

  • Rønne Lervarefabrik
  • Stampeå stream
  • South of Hasle
  • Hvideoddebuen
  • Nebbeodde

|Shells

|

A possible Ammonite of the family Polymorphitidae inside Ammonitida.

|

rowspan="5" |

Aegoceras

|

A. armatum

| rowspan="4" |Shells

| rowspan="4" |

An Ammonite of the family Liparoceratidae inside Ammonitida. Later renamed Beaniceras centaurus, senior synonym of Aegoceras centaurus

| rowspan="5" |

File:Aegoceras lataecosta (fossil ammonite) (Lower Jurassic).jpg

style="background:#fbdddb;" |

A. centaurus

A. capraius
A. mangenesti
style="background:#fbdddb;" |

A. pettos

|style="background:#fbdddb;" |Shells

|style="background:#fbdddb;" |

Junior synonym of Coeloceras pettos, possible synonym of Coeloceras grenouillouxi

Apoderoceras

|

A. aculeatum

|Shells

|

An Ammonite of the family Polymorphitidae inside Ammonitida.

|

Arnioceras

|

A. falcaries

|Shells

|

An Ammonite of the family Arietitidae inside Ammonitida.

|

File:Arnioceras acuticarinatu - Naturhistorisches Museum, Braunschweig, Germany - DSC05129 cropped.jpg

Beaniceras

|

B. centaurus

|Shells

|

An Ammonite of the family Liparoceratidae inside Ammonitida. Senior synonym of Aegoceras centaurus

|

rowspan="2" |

Coeloceras

|

C. pettos

|Shells

|

An Ammonite of the family Coeloceratidae inside Psilocerataceae. Senior synonym of Aegoceras pettos. Possible junior synonym of Coeloceras grenouillouxi

|

style="background:#fbdddb;" |

C. grenouillouxi

|style="background:#fbdddb;" |Shells

|style="background:#fbdddb;" |

Possible senior synonym of Coeloceras pettos

|style="background:#fbdddb;" |

rowspan="2" |

Paramicroderoceras

|

P. fila

| rowspan="2" |Shells

|rowspan="2" |

An Ammonite of the family Juraphyllitidae inside Phylloceratina.

|

P.? sp indet

|

Phylloceras

|

P. loscombii

|Shells

|

An Ammonite of the family Juraphyllitidae inside Phylloceratina.

|

File:Phylloceras NT.jpg

Phricodoceras

|

P. taylori

|Shells

|

An Ammonite of the family Phricodoceratidae inside Psilocerataceae.

|

rowspan="4" |

Platypleuroceras

|

P. brevispina

| rowspan="4" |Shells

|rowspan="4" |

An Ammonite of the family Polymorphitidae inside Ammonitida.

|

P. submulticum

|

P. caprarium

|

P. spp. indet

|

Radstockiceras

|

R. hechingense

|Shells

|

An Ammonite of the family Oxynoticeratidae inside Psilocerataceae.

|

rowspan="2" |

Tragophylloceras

|

T. numismale

| rowspan="2" |Shells

|rowspan="2" |

An Ammonite of the family Juraphyllitidae inside Phylloceratina.

|

T. sp

|

rowspan="2" |

Uptonia

|

U. lata

|Shells

|rowspan="2" |

An Ammonite of the family Polymorphitidae inside Ammonitida.

|

File:Uptonia jamesoni ESV.jpg

U. sp indet

|Shells

|

= Crustacea =

class="wikitable" align="center"
Genus

! Species

! Location

! Material

! Notes

! Images

Goniodromites?{{Cite journal |last=Beurlen |first=Karl |date=1932 |title=Brachyurenreste aus dem Lias von Bornholm mit Beiträgen zur Phylogenie und Systematik der Brachyuren Decapoden |url=https://doi.org/10.1007/bf03041617 |journal=Paläontologische Zeitschrift |volume=14 |issue=1-2 |pages=52–66 |doi=10.1007/bf03041617 |issn=0031-0220|url-access=subscription }}

|

  • G.? liasicus

|

  • Stampeå stream

|

  • 2 partially preserved specimens

|A possible early Brachyuran, then recovered as not related with Goniodromites, and suggested to be from younger strata, what was contested latter.{{Cite journal |last=Schweitzer |first=Carrie E. |last2=Feldmann |first2=Rodney M. |date=2010-04-01 |title=The Oldest Brachyura (Decapoda: Homolodromioidea: Glaessneropsoidea) Known to Date (Jurassic) |url=https://academic.oup.com/jcb/article-abstract/30/2/251/2419233 |journal=Journal of Crustacean Biology |volume=30 |issue=2 |pages=251–256 |doi=10.1651/09-3231.1 |issn=0278-0372}}{{Cite journal |last=Krobicki |first=Michał |last2=Zatoń |first2=Michał |date=2016-09-01 |title=A New Homolodromioid Crab (Brachyura: Dromiacea: Tanidromitidae) from the Bajocian of Central Poland and a Review of the Stratigraphical Distribution and Paleoenvironments of the Known Middle Jurassic Homolodromioids |url=https://academic.oup.com/jcb/article-abstract/36/5/695/2735719?redirectedFrom=fulltext |journal=Journal of Crustacean Biology |volume=36 |issue=5 |pages=695–715 |doi=10.1163/1937240X-00002463 |issn=0278-0372}}

|

= Chondrichthyes =

Unidentified fin spines are known from this formation.

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Acrodus

| A. minimus

|Stampeå stream

|Large number of teeth

|A marine shark, member of the family Acrodontidae inside Hybodontiformes.

|

Agaleus

|A. dorsetensis

|Hasle Harbour

|Four teeth

|A marine shark, type member of the family Agaleidae inside Euselachii. This genus may have been a suction feeder as its anterior teeth are similar to those of extant nurse sharks.

|

Hybodontidae

|Indeterminate

|Hasle Harbour

|One incomplete spine

|A marine shark, incertae sedis inside Hybodontiformes.

|

rowspan="2" |Hybodus

|H. reticulatus

|Hasle Harbour

|21 incomplete crowns

|A marine shark, type member of the family Hybodontidae inside Hybodontiformes.

| rowspan="2" | File:Hybodus NT.jpg

H. delabechei

|Hasle Harbour

|Three complete crowns

|A marine shark, type member of the family Hybodontidae inside Hybodontiformes. Its teeth suggest that it may have been feeding on shelly invertebrates.

Lissodus

|L. hasleensis

|Hasle Harbour

|Four complete teeth

|A marine/brackish shark, member of the family Lonchidiidae inside Hybodontiformes.

|File:Lissodus hasleensis.jpg

Myriacanthidae{{Cite journal |last1=Duffin |first1=Christopher |last2=Milàn |first2=Jesper |date=November 2022 |title=Further holocephalian remains from the Hasle Formation (Early Jurassic) of Denmark |url=https://www.researchgate.net/publication/365870659 |journal=Bulletin of the Geological Society of Denmark |volume=70 |pages=139–149 |doi=10.37570/bgsd-2022-70-10 |s2cid=254356926 |doi-access=free}}

|Indeterminate

|Hasle Harbour

|Tooth plates

|Indeterminate tooth plates of marine chimaeras.

|

rowspan="2" |MyriacanthusBonde, N., Andersen, S., Hald, N., & Jakobsen, S.L. 2008: Danekræ – Danmarks bedste fossiler, 225 pp. Gyldendal, Copenhagen.

|M. paradoxus

|Cliff section 100 m south of Hasle

|Tooth plates

|A marine chimaera.

|

M. sp.

|Hasle Harbour

|Isolated tooth plate

|A marine Chimaera, type member of the family Myriacanthidae inside Chimaeriformes.

|

Oblidens{{Cite journal|author1=Christopher J. Duffin |author2=Jesper Milàn |year=2017 |title=A new myriacanthid holocephalian from the Early Jurassic of Denmark |journal=Bulletin of the Geological Society of Denmark |volume=65 |pages=161–170 |doi=10.37570/bgsd-2017-65-10 |url=http://2dgf.dk/xpdf/bull65-161-170.pdf }}

|O. bornholmensis

|Cliff section 100 m south of Hasle

|Multiple tooth plates.

|A marine Chimaera, member of the family Myriacanthidae inside Chimaeriformes.

|

Paraorthacodus

|P. sp

|Hasle Harbour

|Two incomplete teeth

|A marine shark, member of the family Palaeospinacidae inside Synechodontiformes.

|

Synechodus

|S. occultidens

|Hasle Harbour

|Four incomplete teeth

|A marine shark, type member of the family Palaeospinacidae inside Synechodontiformes.

|

=Actinopteri=

class = "wikitable"
Genus

! Species

! Stratigraphic position

! Material

! Notes

! Images

ArchaeotolithusStolley, E.: Über mesozoische Fischotolithen aus Norddeutschland. — 3. Jahresber. d. Niedersächs. Geol. Vereins, 1910

|

  • A. bornholmiensis
  • A. sp.

|

  • Stampeå stream

|

Otoliths

|

Interpreted as sagittal otoliths of a Palaeonisciformes Indet.{{cite journal |last1=Schwarzhans |first1=W. |title=A review of Jurassic and Early Cretaceous otoliths and the development of early morphological diversity in otoliths. |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |date=2018 |volume=287 |issue=1 |pages=75–121 |doi=10.1127/njgpa/2018/0707 |url=https://www.schweizerbart.de/papers/njgpa/detail/287/88402/A_review_of_Jurassic_and_Early_Cretaceous_otoliths_and_the_development_of_early_morphological_diversity_in_otoliths |access-date=13 August 2021|url-access=subscription }} Fish Otoliths, originally assigned to the extant family Sciaenidae, then from members of Palaeoniscidae and some other research suggest affinities with Pholidophoriformes.

|

Saurichthys?

|

  • S. acuminata?
  • S. longidens?

|

  • Stampeå stream

|

Teeth

|

A marine/brackish Osteichthyes, representative of the family Saurichthyidae inside Chondrostei. The referral to the genus must be taken with caution, as currently it´s geological range is limited to the Triassic.

|

File:Saurichthys (Mensch & Natur-Museum, München).jpg model]]

= Sauropterygia =

class = "wikitable"
GenusSpeciesLocationMaterialNotes

!Images

Plesiosauria

|

Indeterminate

|

  • Stampeå stream
  • Hasle Harbour

|

  • Complete right ischium
  • Complete isolated humerus
  • Complete ulna/fibula
  • Isolated flattened lunate epipodial bone
  • Fragment of jaw
  • Sacral Vertebra
  • Sacral Rib
  • Single Fragment of Rib

|

Indeterminate Plesiosauroidea Material

|

Plesiosauroidea{{cite journal |last1=Rees |first1=J. |last2=Bonde |first2=N. |title=Plesiosaur remains from the Early Jurassic Hasle Formation, Bornholm, Denmark |journal=Secondary Adaptation to Life in Water |date=1999 |pages=13–17}}{{cite journal |last1=Smithy |first1=A. S. |title=Plesiosaurs from the Pliensbachian (Lower Jurassic) of Bornholm, Denmark |journal=Journal of Vertebrate Paleontology |date=2008 |volume=28 |issue=4 |pages=1213–1217 |doi=10.1671/0272-4634-28.4.1213 |jstor=20491054 |bibcode=2008JVPal..28.1213S |s2cid=128999502 |url=https://www.jstor.org/stable/20491054 |access-date=6 October 2021}}

|

Indeterminate

|Hasle Harbour

|

Tooth

|

Indeterminate Plesiosauroidea Material

|

Pliosauroidea

|

Indeterminate

|Hasle Harbour

|

Fourteen isolated teeth

|

A marine plesiosaur, member of the family Pliosauroidea inside Sauropterygia. Resembles Attenborosaurus.

|

Rhomaleosauridae

|

Indeterminate

|Hasle Harbour

|

Tooth

|

A marine plesiosaur, member of the family Rhomaleosauridae inside Sauropterygia. Resembles Rhomaleosaurus.

|File:Rhomaleosaurus crampt12DB.jpg ]]

= Thalattosuchia =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Teleosauridae{{cite journal |last1=Milàn |first1=J. |last2=Mueller-Töwe |first2=I. |title=En havskrokodille fra Hasles fjerne fortid |journal=Natur på Bornholm |date=2021 |volume=19 |issue=1 |pages=14–16 |url=http://publicationslist.org/data/jesper.milan/ref-220/(75)%20Mil%C3%A0n%20_%20Mueller-T%C3%B6we%202021%20-%20En%20havkrokodille%20fra%20Hasles%20fjerne%20fortid.pdf |access-date=6 October 2021}}

|

Indeterminate

|

Hasle Harbour

|

DK-1072, a dermal plate

|

A Teleosauridae Thalattosuchian. One of the northernmost fossil finds of the group and one of the few of the pliensbachian worldwide, it represents a genus whose dermal armor was similar to the genus Macrospondylus of the Toarcian.

|

File:Steneosaurus bollensis, view 1, Early Jurassic, Toarcian Age, Posidonia Shale, Holzmaden, Baden-Wurttemberg, Germany - Houston Museum of Natural Science - DSC01844.JPG example of Teleosauroid]]

= Pterosauria =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Pterosauria?{{cite book |last1=G. Olsen |first1=Ernst |title=Bornholms Historie: landbruget og dets biografier |date=1950 |publisher=Ego |location=RØNNE |page=15 |url=https://bornholmske-samlinger.dk/wp-content/uploads/2014/10/Bornholms-Historie-Landbruget-og-dets-Biografier.pdf?fbclid=IwAR1V_S6i4rkb9vL-dOLd4ZuddRdMPqYlNT-mNOr6p3nZACKOg-rFjj-L_vU |access-date=26 October 2021}}

|

Indeterminate

|

Hasle Harbour

|

Uncertain Remains

|

A possible Pterosaur. Is quoted the find of a "Flying Reptile" on the Hasle Sandstone by C. Malling at the beginning of the 20th century. No further data related to the find has been published.

|

File:Dimorphodon-macronyx jconway.jpg]]

= Theropods =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Neotheropoda{{cite journal |last1=Molin |first1=E. |title=Rare terrestrial vertebrate remains from the Pliensbachian (Lower Jurassic) Hasle Formation on the Island of Bornholm, Denmark |journal=Dissertations in Geology at Lund University |date=2021 |volume=1 |issue=1 |url=https://lup.lub.lu.se/student-papers/search/publication/9049640 |access-date=6 October 2021}}

|

Indeterminate

|

Hasle Harbour

|

Propodial bone, possibly a radius

|

Theropod, incertae sedis inside Neotheropoda. Extremely hollow shaft suggests that it most likely belonged to a juvenile member of a theropod dinosaur.

|

File:Segisaurus.jpg a coeval North American genus]]

style="background:#FEF6E4;" |

Stenonyx{{cite journal |last1=Milàn |first1=Jesper |last2=Surlyk |first2=Finn |title=An enigmatic, diminutive theropod footprint in the shallow marine Pliensbachian Hasle Formation, Bornholm, Denmark |journal=Lethaia |date=2014 |volume=48 |issue=1 |pages=429–435 |doi=10.1111/let.12115 |url=http://publicationslist.org/data/jesper.milan/ref-151/Mil%C3%A0n%20_%20Surlyk%202015%20-%20Small%20theropod%20track%20Bornholm.pdf |access-date=6 October 2021}}{{cite web |last1=Milán |first1=J. |title=Hasle Formationen |url=http://www.geocenter.dk |website=Geocenter DK |publisher=Geological Survey of Denmark and Greenland |access-date=6 October 2021}}

| style="background:#FEF6E4;" |

S. isp.

| style="background:#FEF6E4;" |

Hasle Harbour

| style="background:#FEF6E4;" |

MGUH – 30889, single Footprint

| style="background:#FEF6E4;" |

Theropod Tracks, member of the ichnofamily Anchisauripodidae, incertae sedis inside Theropoda. Considered a very Small Dinosaur, probably a juvenile. The Foot resembles those of young Coelophysis from Ghost Ranch Quarries. Stenonyx footprints had been described from the Early Jurassic Soltykow (Hettangian) and an identical one from Szydlowek (Pliensbachian), linking southern Sweden, Bornholm and Poland, contiguous during the Early Jurassic and dinosaurs could thus freely roam this large area.

| style="background:#FEF6E4;" |

Theropoda{{Cite journal |last=Molin |first=E. |last2=Hall |first2=S. A. |last3=Heingård |first3=M. |last4=Milàn |first4=J. |last5=Lindgren |first5=J. |date=2024 |title=The first tritylodontid (Synapsida, Cynodontia) fossil from Scandinavia |url=https://2dgf.dk/xpdf/bull73-221-230.pdf |journal=Bulletin of the Geological Society of Denmark |volume=73 |issue=1 |pages=221–230}}

|Indeterminate

|Hasle Harbour

|Isolated Teeth

|Theropod, incertae sedis

|

= Sauropodomorpha =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Massopoda{{cite journal |last1=Milàn |first1=J. |last2=Cuny |first2=G. |title=Knogle fra mini sauropod fundet ved Hasle |journal=Geomuseum Faxe Danmark, University of Lyon Frankrig |date=2019 |url=http://publicationslist.org/data/jesper.milan/ref-208/70%20-%20Mil%C3%A0n%20_%20Cuny%202019%20-%20Sauropodomorf%20fra%20Hasle%20-%20Natur%20p%C3%A5%20Bornholm%202018.pdf |access-date=6 October 2021}}

|

Indeterminate

|

Hasle Harbour

|

DK966, metatarsal, metapodial or a propodial bone

|

A Sauropodomorph of uncertain placement, probably a rather basal genus.

|

File:Mussaurus patagonicus life restoration.png, a possible relative of DK966]]

Sauropoda{{cite journal |last1=Schnetler |first1=I. |title=Medlemsblad for Jysk Stenklub |journal=Geomuseum Faxe Danmark. University of Lyon Frankrig |date=2019 |url=http://www.jyskstenklub.dk/pdf-filer/2018%20Stenhuggeren%20december.pdf |access-date=6 October 2021}}

|

Indeterminate

|

Hasle Harbour

|

DK976, spinous process from a cervical vertebrae

|

A Sauropod Sauropodomorph. Assigned to an early actual Sauropods, which also have quite powerful lateral processes and whose cervical vertebrae are relatively long.

|

Turiasauria{{cite journal |last1=Milàn |first1=J. |last2=Mateus |first2=O. |title=A Turiasaurian (Dinosauria, Sauropoda) Tooth from the Pliensbachian Hasle Formation of Bornholm, Denmark, Shows an Early Jurassic Origin of the Turiasauria |journal=Diversity |date=2024 |volume=16 |issue=1 |page=12|doi=10.3390/d16010012 |doi-access=free |hdl=10362/172148 |hdl-access=free }}

|

Indeterminate

|

Hasle Harbour

|

NHMD 1185136, Isolated tooth

|

A possible Turiasaurian Eusauropod. The oldest geological record of the family, likely represents a new unnamed taxon, estimated to have been 10–14 m long based on related taxa.

|File:Turiasaur tooth from Bornholm.png

= Synapsida =

class ="wikitable" align="center"
GenusSpeciesLocationMaterialNotesImages
Tritylodontidae

|

Indeterminate

|

Hasle Harbour

|NHMD 1725979: an isolated tooth crown

|

A Tritylodontoidea Cynognathian. Hasle Formation fossil cannot be confidently placed within any known genus, it can be either a primitive Tritylodont related to specimens that lack post-canines (Yunnanodon, Dianzhongia), or an early derived genus of the Stereognathus group. It is the oldest occurrence of a mammaliamorph in Scandinavia.

|File:Bienotherium yunnanenseDB24.jpg, a coeval genus]]

See also

References