Tendaguru Formation

{{Short description|Geological formation and paleontological site in Tanzania}}

{{Infobox Rockunit

| name = Tendaguru Formation

| image =Tendaguru Tendaguru3 (cropped).jpg

| caption =

| type = Geological formation

| age =
?Callovian-Hauterivian
~{{fossil range|165|130}}

| period = Late Jurassic

| prilithology = Sandstone

| otherlithology = Shale, siltstone, clay, conglomerate, limestone

| namedfor = Tendaguru Hill

| namedby = Janensch & Hennig

| year_ts = 1914

| region = Lindi Region

| country = Tanzania

| coordinates = {{coord|9.7|S|39.2|E|display=inline,title}}

| paleocoordinates = {{coord|29.4|S|16.7|E|display=inline}}

| unitof =

| subunits = See text

| underlies = Makonde Formation

| overlies = Neoproterozoic gneiss basement

| thickness = >{{convert|110|m|ft|abbr=on}}

| extent = Mandawa Basin

| area =

| map = {{Location map+ | Tanzania

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Tanzania

| lat_deg = -9.7

| lon_deg = 39.2

| mark = Green-blue pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

File:Tendaguru location map.jpg

The Tendaguru Formation, or Tendaguru Beds are a highly fossiliferous formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represents the oldest sedimentary unit of the Mandawa Basin, overlying Neoproterozoic basement, separated by a long hiatus and unconformity. It reaches a total sedimentary thickness of more than {{convert|110|m|ft}}. The formation ranges in age from the late Middle Jurassic to the Early Cretaceous, Oxfordian to Hauterivian stages, with the base of the formation possibly extending into the Callovian.

The Tendaguru Formation is subdivided into six members; these are from oldest to youngest the Lower Dinosaur Member, the Nerinella Member, the Middle Dinosaur Member, the Indotrigonia africana Member, the Upper Dinosaur Member, and the Rutitrigonia bornhardti-schwarzi member. The succession comprises a sequence of sandstones, shales, siltstones, conglomerates with minor oolitic limestones, deposited in an overall shallow marine to coastal plain environment, characterized by tidal, fluvial and lacustrine influence with a tsunami deposit occurring in the Indotrigonia africana member. The climate of the Late Jurassic and Early Cretaceous was semi-arid with seasonal rainfall and the eustatic sea level was rising in the Late Jurassic from low levels in the Middle Jurassic. Paleogeographical reconstructions show the Tendaguru area was located in the subtropical southern hemisphere during the Late Jurassic.

The Tendaguru Formation is considered the richest Late Jurassic strata in Africa. The formation has provided a wealth of fossils of different groups; early mammaliaforms, several genera of dinosaurs, crocodyliforms, amphibians, fish, invertebrates and flora. More than {{convert|250|t}} of material were shipped to Germany during excavations in the early 20th century. The faunal assemblage of the Tendaguru is similar to the Morrison Formation of the central-western United States, with an additional marine interbed fauna not present in the Morrison.

The dinosaur fauna found in the formation is similar to that of other highly fossiliferous stratigraphic units of the Late Jurassic; among others the Kimmeridge and Oxford Clays of England, the Sables de Glos, Argiles d'Octeville, Marnes de Bléville of France, the Alcobaça, Guimarota and Lourinhã Formations of Portugal, the Villar del Arzobispo Formation of Spain, the Shishugou, Kalazha and Shangshaximiao Formations in China, the Toqui Formation of Chile and Cañadón Calcáreo Formation and the Morrison Formation, with the presence of dinosaurs with similar counterparts, e.g., Brachiosaurus and Stegosaurus in the Morrison, and Giraffatitan and Kentrosaurus in the Tendaguru.Mateus, 2006, pp.223–232

Description

File:Tendaguru Formation - map and stratigraphic column.jpg

The Tendaguru Formation represents the oldest sedimentary unit in the Mandawa Basin, directly overlying Neoproterozoic basement consisting of gneiss. The contact contains a large hiatus, a missing sequence of stratigraphy, spanning the Paleozoic, Triassic and Early Jurassic. The formation is unconformably overlaid by late Early Cretaceous sediments of the Makonde Formation that forms the top of several plateaus; Namunda, Rondo, Noto, and Likonde-Kitale.Bussert et al., 2009, p.154

Based on extended geological and paleontological observations, the "Tendaguruschichten" (Tendaguru Beds) were defined by Werner Janensch as expedition leader and Edwin Hennig in 1914 referring to a sequence of Late Jurassic to Early Cretaceous strata, exposed in the Tendaguru area, which is named after the Tendaguru Hill.Bussert et al., 2009, p.142

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= Stratigraphy =

The Tendaguru is divided into 6 members, which represent different depositional environments, with the 'Dinosaur Beds' representing terrestrial facies while the beds with genus/species names represent marine interbeds with shallow marine to lagoonal facies. In ascending order these are: the Lower Dinosaur Member, the Nerinella Member, the Middle Dinosaur Member, Indotrigonia africana Member, the Upper Dinosaur Member, and the Rutitrigonia bornhardti-schwarzi Member.Schwarz-Wings & Böhm, 2014, p.82

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class="wikitable" style="font-size:80%"

|+ Stratigraphy of the Tendaguru FormationBussert et al., 2009, p.152

Formationwidth=120px | Time periodwidth=120px | MemberLithologywidth=120px | ThicknessImage
Makonde

| style="background-color: {{period color|albian}};" | Early Albian
Aptian

|

| Fine to medium grained sandstones, intercalated conglomerates, siltstones and claystones

| ~{{convert|200|m|ft|abbr=on}}

|

bgcolor=lightgrey |

| style="background-color: {{period color|barremian}};" | Barremian

colspan="4" align=center bgcolor=lightgrey | Unconformity
rowspan="8" | Tendaguru

| style="background-color: {{period color|hauterivian}};" | Hauterivian
Valanginian

| Rutitrigonia bornhardti-schwarzi

| Fine to medium grained sandstones with basal conglomerate

| {{convert|5

70|m|ft|abbr=on}}

| File:Tendaguru Formation - Rutitrigonia bornhardti-schwarzi Member.jpg

style="background-color: {{period color|berriasian}};" | Berriasiancolspan="4" align=center bgcolor=lightgrey | Unconformity
style="background-color: {{period color|tithonian}};" rowspan=2 | Tithonian

| Upper Dinosaur

| Ripple cross bedded fine grained sandstones and siltstones with intercalated claystone and micritic carbonates

| ~{{convert|32|m|ft|abbr=on}}

| File:Tendaguru Formation - Upper Dinosaur Member.jpg

rowspan=2 | Indotrigonia africana

| rowspan=2 | Calcite cemented sandstones, conglomerate beds, thin clay and silt layers with sandy limestones

| rowspan=2 | {{convert|20

50|m|ft|abbr=on}}

| rowspan=2 | File:Tendaguru Formation - Indotrigonia africana Member.jpg

style="background-color: {{period color|kimmeridgian}};" rowspan=2 | Late Kimmeridgian
Middle Dinosaur

| Ripple cross bedded fine grained calcareous sandstones and siltstones and massive to crudely bedded silt and claystones

| {{convert|13

30|m|ft|abbr=on}}

| File:Tendaguru Formation - Middle Dinosaur Member.jpg

style="background-color: {{period color|oxfordian}};" | Kimmeridgian
Oxfordian

| Nerinella

| Trough cross bedded sandstone to massive sandstone

| {{convert|5

45|m|ft|abbr=on}}

| File:Tendaguru Formation - Nerinella Member.jpg

style="background-color: {{period color|callovian}};" | Mid Oxfordian
?Callovian

| Lower Dinosaur

| Cross bedded fine grained sandstones and siltstones, with Interbedded clay-rich siltstones

| >{{convert|20|m|ft|abbr=on}}

|

rowspan=3 bgcolor=lightgrey |style="background-color: {{period color|pliensbachian}};" | Early Jurassicrowspan=3 colspan="6" bgcolor=lightgrey align=center | Hiatus
style="background-color: {{period color|carnian}};" | Triassic
style="background-color: {{period color|paleozoic}};" | Paleozoic
Basement

| style="background-color: {{period color|neoproterozoic}};" | Neoproterozoic

|

| Gneiss

|

|

= Paleogeography and depositional environment =

== Paleogeography ==

File:Paleogeography and paleoclimate of the Late Jurassic - 150 Ma with dinosaur fossil localities.png with S1.]]

The Tendaguru Formation was deposited in the Mandawa Basin, a post-Karoo,Muhongo, 2013, p.28 Mesozoic rift basin located between the Ruvu Basin and Rufiji Trough to the north and the Ruvuma Basin to the south.Muhongo, 2013, p.8 To the west of the basin, Archean and Early Proterozoic basement rocks crop out.Muhongo, 2013, p.33 The main rift phase in present-day southeastern Africa led to the separation of Madagascar and the then-connected Indian subcontinent that happened during the Early Cretaceous.Muhongo, 2013, p.3 The Songo Songo and Kiliwani gas fields are located just offshore the basin.Muhongo, 2013, p.17Muhongo, 2013, p.22

At time of deposition the area was undergoing a semi-arid climate with coastal influences that maintained somewhat higher moisture levels than seen inland.Noto & Grossmann, 2010, p.7 The upper parts of the formation, the Middle Dinosaur and Rutitrigonia bornhardti-schwarzi Members in particular, showed prevailing semiarid conditions with pronounced dry seasons, based on palynologic analysis.Schrank, 1999, p.181 The Tendaguru fauna was stable through the Late Jurassic.Noto & Grossmann, 2010, p.9

During the Late Jurassic and Early Cretaceous, the Gondwana paleocontinent was breaking up and the separation of the Laurasian and Gondwana supercontinents resulted from the connection of the Tethys Ocean with the proto-Atlantic and the Pacific Ocean. In addition, the South Atlantic developed towards the end of the Late Jurassic with the separation of South America and Africa. Africa became increasingly isolated from most other continents by marine barriers from the Kimmeridgian into the Early Cretaceous, but retained a continental connection with South America. Global sea levels dropped significantly in the Early Jurassic and remained low through the Middle Jurassic but rose considerably towards the Late Jurassic, deepening the marine trenches between continents.Arratia et al., 2002, p.227

== Depositional environment ==

File:Tendaguru Formation - Generalized depositional environment.jpg

The sedimentary rocks and fossils record a repeated shift from shallow marine to tidal flat environments indicating that the strata of the Tendaguru Formation were deposited near an oscillating strandline which was controlled by sea level changes. The three dinosaur-bearing members are continental to marginal marine and the three sandstone-dominated members are marginal marine in origin.Bussert et al., 2009, p.168

;Nerinella Member

The composition of benthic molluscs and foraminifera, euhaline to mesohaline ostracods, and dinoflagellate assemblages indicate marine, shallow water conditions for the Nerinella Member, in particular for the lower part. Sedimentation occurred as tidal channel fills, subtidal and tidal sand bars, minor storm layers (tempestites), and beach deposits. Overall, the Nerinella Member represents a variety of shallow subtidal to lower intertidal environments influenced by tides and storms.Bussert et al., 2009, p.167

;Middle Dinosaur Member

The sedimentological characteristics of the basal part of the Middle Dinosaur Member suggest deposition on tidal flats and in small tidal channels of a lagoonal paleoenvironment. The ostracod Bythocypris sp. from the member indicates polyhaline to euhaline conditions. Slightly higher up, a faunal sample dominated by the bivalve Eomiodon and an ostracod assemblage composed of brackish to freshwater taxa is indicative of a brackish water paleoenvironment with distinct influx of freshwater as revealed by the nonmarine ostracod genus Cypridea, charophytes, and other freshwater algae. The paleoenvironment of the ostracod assemblages of the Middle Dinosaur Member changed upsection from a marine setting in the basal parts through alternating marine-brackish conditions to freshwater conditions in the higher parts of this member. The highly sporadic occurrence, in this part of the section, of molluscs typical of marginal marine habitats indicates only a very weak marine influence, at sabkha-like coastal plains with ephemeral brackish lakes and ponds are recorded in the upper part of the Middle Dinosaur Member. This part also contains pedogenic calcretes indicating subaerial exposure and the onset of soil formation. The calcrete intraclasts within adjacent sandstone beds testify to erosive reworking of calcrete horizons.Aberhan et al., 2002, p.32 The presence of crocodyilforms indicates freshwater to littoral environments and adjacent terrestrial areas.Aberhan et al., 2002, p.33

;Indotrigonia africana Member

The coarse-grained sandstone of the lower part of the Indotrigonia africana Member that shows highly variable transport directions is interpreted as deposits of large tidal channels. Grain-size, large-scale sedimentary structures, and the lack of both trace fossils and epifaunal and infaunal body fossils suggest high water energy and frequent reworking. This basal succession passes upward in cross-bedded sandstone and minor siltstone and claystone with flaser or lenticular bedding that are interpreted as tidal flat and tidal channel deposits. Horizontal to low-angle cross-bedded, fine-grained sandstone with intercalated bivalve pavements indicates tidal currents that operated in small flood and ebb tidal deltas and along the coast. Stacked successions of trough cross-bedded, medium- to coarse-grained sandstone of the upper part of the Indotrigonia africana Member are interpreted as tidal channel and sand bar deposits. At some places in the surroundings of Tendaguru Hill, these sediments interfinger with oolitic limestone layers that represent high-energy ooid shoals.

In the Tingutinguti stream section, the Indotrigonia africana Member exhibits several up to {{convert|20|cm|in}} thick, poorly sorted, conglomeratic sandstone beds. They contain mud clasts, reworked concretions and/or accumulations of thick-shelled bivalves (mainly Indotrigonia africana and Seebachia janenschi), and exhibit megaripple surfaces. These conglomeratic sandstone layers are interpreted as storm deposits. In the Dwanika and Bolachikombe stream sections, and in a small tributary of the Bolachikombe creek, a discrete, up to {{convert|70|cm|ft}} thick conglomerate in the lower portion of the Indotrigonia africana Member displays evidence of a tsunami deposit. Overall, lithofacies and the

diverse macroinvertebrate and microfossil assemblages of the Indotrigonia africana Member suggest a shallow marine environment. Based on the diverse mesoflora and the abundance of Classopollis, a nearby vegetated hinterland is postulated that was dominated by xerophytic conifers.

;Upper Dinosaur Member

The small-scale trough and ripple cross-bedded fine-grained sandstone at the base of the Upper Dinosaur Member is interpreted as tidal flat deposits. Unfossiliferous sandstone in the upper part

was most likely deposited in small fluvial channels in a coastal plain environment, whereas argillaceous deposits were laid down in still water bodies such as small lakes and ponds. Rare occurrences of the ostracod Cypridea and charophytes signal the influence of freshwater, whereas the sporadic occurrence of marine invertebrates suggests a depositional environment close to the sea.

;Rutitrigonia bornhardti-schwarzi Member

Fining upward sequences of the basal part of the Rutitrigonia bornhardti-schwarzi Member are interpreted as tidal channel fills, the overlying fine-grained sandstone, silt- and claystone as tidal flat deposits. From the immediate surroundings of Tendaguru Hill, invertebrates and vertebrates are poorly known and limit the palaeoenvironmental interpretation of this member. The composition of the land-derived sporomorph assemblage suggests a terrestrial vegetation which was dominated by cheirolepidiacean conifers in association with ferns.

Excavation history

File:Geologic map Tendaguru Formation members, Tanzania.jpg

The Tendaguru Beds as a fossil deposit were first discovered in 1906, when German pharmacist, chemical analyst and mining engineer Bernhard Wilhelm Sattler, on his way to a mine south of the Mbemkure River in German East Africa in today's Tanzania, was shown by his local staff enormous bones weathering out of the path near the base of Tendaguru Hill, {{convert|10|km|mi}} south of Mtapaia (close to Nambiranji village, Mipingo ward, {{convert|60|km|mi}} northwest of Lindi town).Maier, 2003 Because of its morphology, the hill was locally known as "steep hill" or "tendaguru" in the language of the local Wamwera people. Sattler forwarded his report on the giant bones to the Kommission für die landeskundliche Erforschung der deutschen Schutzgebiete (“Commission for the Geographical Investigation of the German Protectorates”), headed by the geographer Hans Meyer.{{Cite book |last=Maier |first=Gerhard |title=African dinosaurs unearthed: the Tendaguru expeditions |date=2003 |publisher=Indiana University Press |isbn=978-0-253-34214-0 |series=Life of the past |location=Bloomington |pages=48-53}} Meyer, in turn, wrote to the palaeontologist Eberhard Fraas,{{Cite web |title=Tanzania dinosaurs: amazing discoveries of ancient giants |url=https://altezzatravel.com/articles/dinosaurs-tanzania |access-date=2025-04-23 |website=Altezza Travel |language=en}} who was in East Africa at that time, and urged him to hasten to the Tendaguru site.{{Cite book |last=Tamborini |first=Marco |title=Deconstructing dinosaurs: the history of the German Tendaguru Expedition and its finds, 1906-2022 |last2=Ohl |first2=Michael |last3=Sadock |first3=Musa |last4=Mapunda |first4=Bertram |last5=Vennen |first5=Mareike |last6=Stoecker |first6=Holger |last7=Heumann |first7=Ina |date=2025 |publisher=Brill |isbn=978-90-04-69105-6 |location=Leiden ; Boston}} The latter visited the site in 1907 and, with the aid of Sattler, recovered two partial skeletons of enormous size.Fraas, 1908

Following the discovery in 1906, teams from the Museum für Naturkunde, Berlin (1907–1913), and later the British Museum (Natural History), London (1924–1931) launched a series of collecting expeditions that remain unequalled in scope and ambition. Led by the vision and influence of geologist Wilhelm von Branca, the German expeditions were particularly successful. This was largely due as the project was regarded as a matter of national ambition of the German Empire and enjoyed the benevolence of wealthy patrons. Eventually, nearly 250 tons of fossils were shipped to Berlin, representing an entirely new dinosaur fauna that remains the best understood assemblage from all of former Gondwana.Cifelli, 2003, p.608

From there, the material was transported to Fraas' institution, the Royal Natural History Collection in Stuttgart, Germany. Fraas described two species in the badly known genus "Gigantosaurus"; G. robustus and G. africanus (today Janenschia robusta and Tornieria africana, respectively.

= German Tendaguru Expedition =

The Berlin Natural History Museum excavated at Tendaguru hill and in the surroundings for four years. From 1909 through 1911, Werner Janensch as expedition leader and Edwin Hennig as assistant directed excavations, while Hans Reck and his wife Ina Reck led the 1912 field season. Other European participants included Hans von Staff. In the rainy seasons the scientists explored the geology of German East Africa on long safaris.

= Public discussion about provenance and restitution =

In the context of international discussion about the provenance and possible restitution of colonial heritage, as discussed for example in the 2018 report on the restitution of African cultural heritage, both German as well as Tanzanian commentators have called the claim to rightful ownership by the Berlin museum into question. The Tanzanian government has, however, not submitted any official demand for repatriation. German authorities have preferred to offer information on the provenance and research by increasing cooperation between Tanzanian paleontologists and museums with their German counterparts.{{Cite web|last=Vogel|first=Gretchen|date=2019-03-27|title=Countries demand their fossils back, forcing natural history museums to confront their past|url=https://www.science.org/content/article/countries-demand-their-fossils-back-forcing-natural-history-museums-confront-their-past|access-date=2021-06-15|website=Science {{!}} AAAS|language=en}}

Paleontological significance

Possible dinosaur eggs have been recovered from the formation.Weishampel et al., 2004, p.552 Further, the fauna of the Tendaguru Formation has been correlated with the Morrison Formation of the central-western United States,Taylor, 2009, p.790 several formations in England, such as the Kimmeridge Clay and Oxford Clay, and in France (Sables de Glos, Argiles d'Octeville, Marnes de Bléville), the Alcobaça, Guimarota and Lourinhã Formations in Portugal,Mateus, 2006, p.1 the Villar del Arzobispo Formation in Spain, the Shishugou, Kalazha and Shangshaximiao Formations in China, the Toqui Formation in the Magallanes Basin, Chile and the Cañadón Calcáreo Formation in the Cañadón Asfalto Basin in central Patagonia, Argentina.Noto & Grossmann, 2010, p.3

= Fossil content =

{{paleobiota-key-compact}}

== Mammaliaformes ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Mammaliaformes reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! Notes

! Images

Allostaffia

| A. aenigmatica

| Quarry Ig

| Middle Dinosaur

| Three isolated molars

| Originally described as Staffia, later renamed Allostaffia as Staffia was preoccupied bya foraminifer.Heinrich, 2004
Assigned to Haramiyida (though possibly a gondwanathere instead).Chimento et al., 2016

|

Brancatherulum

| B. tendagurense

| Unspecified

| Upper Dinosaur

| Dentary without teeth

| Either a stem-zatherian or dryolestidan.Averianov & Martin, 2015, p.327

|

Tendagurodon

| T. janenschi

| Quarry Ig

| Middle Dinosaur

| Single tooth

| One of the earliest amphilestidsHeinrich, 1998, p.269

|

Tendagurutherium

| T. dietrichi

| Quarry Ig

| Middle Dinosaur

| Partial dentary with damaged last molar

| Either a peramurid or an australosphenidan

|

== Squamates ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Squamates reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! Notes

! Images

Paramacellodidae?

| Indeterminate

|

|

| An osteoderm

| Unconfirmed{{Citation |last=Evans |first=Susan E. |title=The Origin and Early Diversification of Squamates |date=2022-08-11 |url=https://www.cambridge.org/core/product/identifier/9781108938891%23CN-bp-2/type/book_part |work=The Origin and Early Evolutionary History of Snakes |pages=7–25 |editor-last=Gower |editor-first=David J. |access-date=2024-01-11 |edition=1 |publisher=Cambridge University Press |doi=10.1017/9781108938891.004 |isbn=978-1-108-93889-1 |editor2-last=Zaher |editor2-first=Hussam|url-access=subscription }}

|

== Pterosaurs ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Pterosaurs reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! Notes

! Images

Tendaguripterus

| T. recki

| Quarry Ig

| Middle Dinosaur

| A partial mandible with teeth

| align=center | Barrett et al., 2008

| rowspan="99" | File:Tendaguru pterosaurs.jpg

?Indeterminate archaeopterodactyloid

|

| Mkoawa Mtwara

|

| Humerus

| align=center | Costa & Kellner, 2009, p.814

Indeterminate azhdarchid

|

| Mkoawa Mtwara

|

|

| align=center |

Indeterminate dsungaripteroid

|

|

| Upper Dinosaur

| Humerus

| align=center |

rowspan="3" style="background:#f3e9f3;" | Pterodactylus

| style="background:#E6E6E6;" | P. maximus

| style="background:#E6E6E6;" | Mkoawa Mtwara

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" | Later determined to be an indeterminate pterodactyloid

style="background:#E6E6E6;" | P. brancai

| style="background:#E6E6E6;" | Mkoawa Mtwara

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" | TibiotarsiGalton, 1980

| style="background:#E6E6E6;" | Later determined to be an indeterminate dsungaripteroid

style="background:#E6E6E6;" | P. arningi

| style="background:#E6E6E6;" | Mkoawa Mtwara

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" | Later determined to be an indeterminate pterosaur

style="background:#f3e9f3;" | Rhamphorhynchus

| style="background:#E6E6E6;" | R. tendagurensis

| style="background:#E6E6E6;" | Mkoawa Mtwara

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" | Later determined to be an indeterminate "rhamphorhynchoid"

== Ornithischians ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Ornithischians reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! Notes

! Images

Dysalotosaurus

| D. lettowvorbecki

| Quarry Ig

| Middle Dinosaur

| "Large number of mostly disassociated cranial and postcranial elements"

| align=center | A dryosaurid"Table 19.1," in Weishampel et al., 2004, p.414

| File:Dryosaurus lettowvorbecki skeleton.jpg

Kentrosaurus

| K. aethiopicus

| Quarry Q, Ig, St, S, Ny, Li, XX, r, y, d, Ng, X, H, IX, Om, bb, Ha, XIV, II, IV, V, VIII, G, e, g, Ki

| Lower, Middle & Upper Dinosaur

| "[Two] composite mounted skeletons, [four] braincases, [seven] sacra, more than [seventy] femora, approximately 25 isolated elements, juvenile to adult"

| align=center | A stegosaur"Table 16.1," in Weishampel et al., 2004, p.344

| File:Fossil Kentrosaurus aethiopicus in Museum für Naturkunde Berlin 001.JPG

== Sauropods ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Sauropods reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! width="40%" | Notes

! Images

Australodocus

| A. bohetii

| Quarry G

| Upper Dinosaur

| Two neck vertebrae; more undescribed remains destroyed during World War II

| A SomphospondylanRemes, 2007

| File:Australodocus LM.png

rowspan="2" | Dicraeosaurus

| D. hansemanni

| Quarry Q, m, St, dd, Sa

| Lower, Middle & Upper Dinosaur

| "Skeleton lacking skull and forelimbs, [two] partial skeletons, isolated vertebrae, and limb elements"

| rowspan = 2|A Dicraeosaurid"Table 13.1," in Weishampel et al., 2004, p.264

| File:Dicraeosaurus hansemanni22.jpg

D. sattleri

| Quarry La, s, O, ab, E, M, o, Ob, bb, XIV, G, GD

| Middle & Upper Dinosaur

| "[Two] partial skeletons without skulls, isolated postcranial remains"

| File:Dicraeosaurus sattleri Skeletal.svg

Giraffatitan

| G. brancai

| Quarry Q, J, Ig, Y, St, dd, S, TL, XX, Ma, JR, Ng, Bo, To, p, t, Lw, D, N, ab, cc, X, IX, Z, T, Aa, l, E, XIV, II, G, e, Ki, No, R, F, XII, GD, XV, Sa, U, i

| Lower, Middle & Upper Dinosaur

|

| Brachiosaurid. The new genus Giraffatitan was erected to hold the former Brachiosaurus species, B. brancai after scientists concluded that it was distinct enough from the Brachiosaurus type species, B. altithorax, to warrant such a reclassification.Taylor, M.P., 2009, pp.787-806

| File:Museum für Naturkunde (36556352434).jpg

Janenschia

| J. robusta

| Quarry dd, P, IX, B, G, Oa, NB

| Middle & Upper Dinosaur

| Known from hindlimb and forelimb material, left pubis and two right ischia

| align=center | non-neosauropod eusauropod

| File:Janenschia.jpg

Tendaguria

| T. tanzaniensis

| Nambango site

| Upper Dinosaur

| "[Two] associated cranial dorsal vertebrae"

| A turiasaur"Table 13.1," in Weishampel et al., 2004, p.261

|

Tornieria

| T. africana

| Quarry St, k, MD, A, e, Sa

| Middle & Upper Dinosaur

| "More than [three] partial skeletons, a few skull elements, [and] many isolated postcranial elements"

| align=center | Diplodocid"Table 13.1," in Weishampel et al., 2004, p.265

|

Wamweracaudia

| W. keranjei

|

|

| A sequence of caudal vertebrae

| align=center | MamenchisauridMannion et al., 2019

|

rowspan="2" style="background:#f3e9f3;" | Brachiosaurus

| style="background:#fbdddb;" | B. brancai

| style="background:#fbdddb;" | Mkoawa Mtwara

| style="background:#fbdddb;" |

| style="background:#fbdddb;" | "[Five] partial skeletons, more than [three] skulls, [and] isolated limb elements"

| style="background:#fbdddb;" | B. brancai was distinct enough from the non-Tendaguru Brachiosaurus type species B. altithorax that it was moved to its own genus, Giraffatitan."Table 13.1," in Weishampel et al., 2004, p.267

|

style="background:#E6E6E6;" | B. fraasi

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" |

| style="background:#E6E6E6;" | Remains attributed to B. fraasi were later referred to B. brancai, and thus now Giraffatitan

|

Diplodocinae indet.

| Indeterminate

| Kijenjere

| Upper Dinosaur

| Partial skull

| Belonging to a form that is closely related to DiplodocusRemes, 2009, p.26

| File:Diplodocinae indet. - Tendaguru Formation.jpg

Diplodocidae indet.

| Indeterminate

| Kijenjere

| Upper Dinosaur

| Caudal vertebrae and metatarsal

| Originally referred to as "Barosaurus africanus"Remes, 2009, p.28

| File:Diplodocidae indet. (Barosaurus africanus) - Tendaguru Formation.jpg

Diplodocidae indet.

| Indeterminate

| Trench XIV

| Upper Dinosaur

| Articulated pedes

| Possibly representing two different taxaRemes, 2009, p.30

| File:Diplodocidae indet. - Tendaguru Formation.jpg

Flagellicaudata indet.

| Indeterminate

|

| Upper Dinosaur

| Braincase

| Referred to Flagellicaudata indet. based on the derived characters shared with this groupRemes, 2009, p.34

| File:Flagellicaudata indet. - braincase - Tendaguru Formation, Tanzania.jpg

"The Archbishop"

|

|

|

|

| align=center | Brachiosaur, distinct from Giraffatitan{{refn|group=note|Formal description in preparation by Michael Taylor in 2019}}

| File:The Archbishop.jpeg

== Theropods ==

class="wikitable sortable" align="center"
colspan="7" align="center" | Theropods reported from the Tendaguru Formation
Genus

! Species

! Location

! Member

! Material

! Notes

! Images

?Abelisauridae indet.

|Indeterminate

|Quarry TL

|Upper Dinosaur

|A left tibia, a right tibia, and a femur{{Cite journal|last=Rauhut|first=Oliver W. M.|date=2011|title=Theropod dinosaurs from the Late Jurassic of Tendaguru, Tanzania|journal=Special Papers in Palaeontology|volume=86|pages=195–239}}

|Possibly an indeterminate abelisaurid.

|

Megalosauroidea indet.

|Indeterminate

|Quarry MW

|Upper Dinosaur

|left tibia and left astragalus

|A large indeterminate megalosauroid.

|

Elaphrosaurus

| E. bambergi

| Quarry Ig, dd, ?RD

| Middle Dinosaur, ?Upper Dinosaur

| "Postcranial skeleton""Table 3.1," in Weishampel et al., 2004, p.48

| An elaphrosaurine noasaurid

| File:Elaphrosaurus mount MfN Berlin 2018 2.jpg

style="background:#E6E6E6;" | Ostafrikasaurus

| style="background:#E6E6E6;" | O. crassiserratus

| style="background:#E6E6E6;" | Quarry Om

| style="background:#E6E6E6;" | Upper Dinosaur

| style="background:#E6E6E6;" | "Tooth"

| style="background:#E6E6E6;" | A tooth of controverisial affinitiies, had either been suggested to have been a spinosaurid or a ceratosaurid.Buffetaut, 2012, p.2{{cite journal |last1=Soto |first1=Matías |last2=Toriño |first2=Pablo |last3=Perea |first3=Daniel |title=Ceratosaurus (Theropoda, Ceratosauria) teeth from the Tacuarembó Formation (Late Jurassic, Uruguay) |journal=Journal of South American Earth Sciences |date=2020 |volume=103 |page=102781 |doi=10.1016/j.jsames.2020.102781|bibcode=2020JSAES.10302781S |s2cid=224842133 }}

| File:Ostafrikasaurus holotype tooth by PaleoGeek.png

Veterupristisaurus

| V. milneri

| Quarry St

| Middle Dinosaur

| "Vertebrae"

| The earliest known carcharodontosaurid.

| File:Veterupristisaurus Skeletal Diagram.png

style="background:#E6E6E6;" | ?Allosaurus

| style="background:#E6E6E6;" | ?A. tendagurensis

| style="background:#E6E6E6;" | Quarry TL

| style="background:#E6E6E6;" | Middle Dinosaur

| style="background:#E6E6E6;" | A tibia"Table 4.1," in Weishampel et al., 2004, p.75

| style="background:#E6E6E6;" | Remains now considered "Tetanurae indet." Possibly a megalosauroid or carcharodontosaurid. Originally referred to Allosauridae'

| File:Allosaurus tendagurensis.jpg

style="background:#E6E6E6;" | Ceratosaurus

| style="background:#E6E6E6;" | C. roechlingi

| style="background:#E6E6E6;" | Quarry St, MW

| style="background:#E6E6E6;" | Middle & Upper Dinosaur

| style="background:#E6E6E6;" | Caudal vertebra

| style="background:#E6E6E6;" | Tentatively referred to Ceratosauridae. Originally referred to Allosauridae.

| File:Ceratosaurus nasicornis DB 2.jpg

style="background:#E6E6E6;" | Labrosaurus

| style="background:#E6E6E6;" | L. stechowi

| style="background:#E6E6E6;" | Quarry St, MW

| style="background:#E6E6E6;" | Middle Dinosaur

| style="background:#E6E6E6;" | Teeth

| style="background:#E6E6E6;" | A ceratosaurid, a species of Ceratosaurus. Originally referred to Allosauridae.

|

?Torvosaurus

| T. sp

| Quarry St, MW

| Upper Dinosaur

| Teeth

| Includes remains previously referred to "Megalosaurus" ingens - now known as "Torvosaurus sp".{{cite journal |last1=Soto |first1=Matías |last2=Toriño |first2=Pablo |last3=Perea |first3=Daniel |title=A large sized megalosaurid (Theropoda, Tetanurae) from the late Jurassic of Uruguay and Tanzania |journal=Journal of South American Earth Sciences |date=2020 |volume=98 |page=102458 |doi=10.1016/j.jsames.2019.102458|bibcode=2020JSAES..9802458S |s2cid=213672502 }}

|

== Crocodyliformes ==

class="wikitable sortable"

! Genus

! Species

! Location

! Member

! Material

! Notes

! Images

Bernissartia

| B. sp

|

| Upper & Middle Dinosaur

|

| align=center | Bussert et al., 2009, p.164

|

== Amphibians ==

class="wikitable sortable"

! Genus !! Species !! Member !! Notes !! Images

?Salientia indetindeterminateMiddle Dinosauralign=center | Aberhan et al., 2002, p.30

== Fish ==

class="wikitable sortable"

! Genus !! Species !! Member !! Notes !! Images

EngaibatisEngaibatis schultzeiUpper Dinosauralign=center | Arratia et al., 2002, p.219File:Engaibatis schultzei - Tendaguru Formation, Tanzania.jpg
rowspan=2 | LepidotesLepidotes tendaguruensisMiddle Dinosauralign=center |
L. sp.Upper & Middle Dinosauralign=center | Arratia et al., 2002, p.224File:Lepidotes sp. - Tendaguru Formation, Tanzania.jpg
HybodusHybodus sp.Upper Dinosauralign=center | Arratia et al., 2002, p.213File:Hybodus sp. - Tendaguru Formation, Tanzania.jpg
LonchidionLonchidion sp.Upper Dinosauralign=center | Arratia et al., 2002, p.216File:Lonchidion sp. - Tendaguru Formation, Tanzania.jpg
SphenodusSphenodus sp.Upper Dinosauralign=center | Arratia et al., 2002, p.218File:Sphenodus sp. - Tendaguru Formation, Tanzania.jpg

== Invertebrates ==

===Gastropods===

class="wikitable sortable"

! Genus !! Species !! Member
bold is defining !! Notes !! Images

PseudomelaniaPseudomelania dietrichiMiddle Dinosauralign=center |
PromathildiaPromathildia sp.Middle Dinosauralign=center |
NerinellaNerinella cutleriNerinellaalign=center | Bussert et al., 2009, p.159rowspan=5 | File:Tendaguru Formation - invertebrates.jpg

===Bivalves===

class="wikitable sortable"

! Genus !! Species !! Member
bold is defining !! Notes !! Images

EomiodonEomiodon cutleriUpper Dinosauralign=center | Bussert et al., 2009, p.165
rowspan=2 | IndotrigoniaIndotrigonia africanaIndotrigonia africanaalign=center | Bussert et al., 2009, p.162
| I. dietrichiLower Dinosauralign=center |
rowspan=2 | RutitrigoniaRutitrigonia bornhardtirowspan=2 | Rutitrigonia bornhardti-schwarzialign=center |
| R. schwarzialign=center |
AcestaAcesta cutleriLower Dinosauralign=center | Aberhan et al., 2002, p.27
ActinostreonActinostreon hennigiIndotrigonia africanaalign=center |
EntoliumEntolium corneolumLower Dinosauralign=center |
FalcimytilusFalcimytilus dietrichiMiddle Dinosauralign=center |
GrammatodonGrammatodon irritansLower Dinosauralign=center |
rowspan=2 | LiostreaLiostrea dubiensisIndotrigonia africanaalign=center | Aberhan et al., 2002, p.34
| L. kindopeensisIndotrigonia africanaalign=center |
LithophagaLithophaga suboblongaIndotrigonia africanaalign=center |
MeleagrinellaMeleagrinella radiataLower Dinosauralign=center |
NanogyraNanogyra nanaLower Dinosauralign=center |
ProtocardiaProtocardia schenkiLower Dinosauralign=center |
SeebachiaSeebachia janenschiIndotrigonia africanaalign=center |

===Coral===

class="wikitable sortable"

! Genus !! Species !! Member
bold is defining !! Notes !! Images

AstrocoeniaAstrocoenia bernensisIndotrigonia africanaalign=center |
MeandrophylliaMeandrophyllia oolithotithonicaIndotrigonia africanaalign=center |
ThamnoserisThamnoseris sp.Indotrigonia africanaalign=center |

===Ostracods===

class="wikitable sortable"

! Genus !! Species !! Member
bold is defining !! Notes !! Images

BythocyprisBythocypris sp.Middle Dinosauralign=center |
CyprideaCypridea sp.Middle & Upper Dinosauralign=center |

=Flora=

class="wikitable sortable"

! Group !! Taxa !! Member !! Notes !! Images

AraucariaceaeAraucariacitesLower Dinosauralign=center | Aberhan et al., 2002, p.25
CheirolepidiaceaeClassopollisIndotrigonia africana
Lower Dinosaur
align=center |
CupressaceaeCupressinoxylon sp.Rutitrigonia bornhardti-schwarzialign=center | Kahlert et al., 1999, p.192
CycadaceaeCycadoxylon sp.Indotrigonia africanaalign=center | Kahlert et al., 1999, p.188
GinkgoaceaeGinkgoxylon sp.Rutitrigonia bornhardti-schwarzialign=center | Kahlert et al., 1999, p.190
TaxodiaceaeGlyptostroboxylon sp.Middle Dinosauralign=center |
TaxaceaeTaxaceoxylon sp.Rutitrigonia bornhardti-schwarzialign=center |
PrasinophytaCymatiosphaera sp.Indotrigonia africanaalign=center | Schrank, 1999, p.173
ZygnemataceaeOvoidites parvusMiddle Dinosauralign=center |
DinoflagellatesvariousIndotrigonia africana
Middle Dinosaur
align=center |
Gymnosperm pollenvariousIndotrigonia africana
Middle Dinosaur
align=center |
Pteridophytic and bryophytic sporesvariousIndotrigonia africana
Middle Dinosaur
align=center |

See also

{{div col|colwidth=30em}}

{{div col end}}

Notes and references

{{Commons category|Tendaguru Formation}}

= Notes =

{{reflist|group=note}}

= References =

{{reflist|20em}}

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  • {{citation |last=Taylor |first=M.P |year=2009 |title=A Re-evaluation of Brachiosaurus altithorax Riggs 1903 (Dinosauria, Sauropod) and its generic separation from Giraffatitan brancai (Janensch 1914) |url=https://pdfs.semanticscholar.org/25d1/6d91f6fd7ab47da795a299c4b8595d7edf08.pdf |archive-url=https://web.archive.org/web/20190402200148/https://pdfs.semanticscholar.org/25d1/6d91f6fd7ab47da795a299c4b8595d7edf08.pdf |archive-date=2019-04-02 |journal=Journal of Vertebrate Paleontology |volume=29 |issue=3 |pages=787–806 |doi=10.1671/039.029.0309 |bibcode=2009JVPal..29..787T |s2cid=15220647 |accessdate=2019-04-01}}
  • {{citation |last1=Weishampel |first1=David B. |author2-link=Peter Dodson |last2=Dodson |first2=Peter |author3-link=Halszka Osmólska |last3=Osmólska |first3=Halszka |year=2004 |title=The Dinosauria, 2nd edition |url=https://books.google.com/books?id=vtZFDb_iw40C |publisher=Berkeley: University of California Press |pages=1–880 |accessdate=2019-02-21 |isbn=0-520-24209-2 |author1-link=David B. Weishampel }}

{{Lindi}}

{{National Historic Sites of Tanzania}}

Category:Jurassic System of Africa

Category:Lower Cretaceous Series of Africa

Category:Sandstone formations

Category:Shale formations

Category:Siltstone formations

Category:Conglomerate formations

Category:Limestone formations

Category:Shallow marine deposits

Category:Tidal deposits

Category:Deltaic deposits

Category:Lacustrine deposits

Category:Lagoonal deposits

Category:Paleontology in Tanzania

Category:Rufiji-Ruvuma languages