Bahariya Formation

{{Short description|Fossiliferous beds in North Africa}}

{{Infobox rockunit

| name = Bahariya Formation

| image = Bahariya Formation McAfee.jpg

| caption = Restoration of the environment and animals of the formation

| type = Geological formation

| period = Cenomanian

| age = Early Cenomanian
~{{fossil range|100|95}}

| prilithology = Sandstone

| otherlithology = Mudstone, siltstone

| namedfor = Bahariya Oasis

| namedby = Said

| year_ts = 1962

| region = Western Desert

| country = {{EGY}}

| coordinates = {{coord|28|24|20|N|28|48|20|E|region:EG_source:kolossus-frwiki|display=title}}

| underlies = El Heiz Fm., El Naqb Fm.Catuneany et al., 2006, p.122

| overlies = Basement

| thickness = ≈{{convert|100|m|ft|abbr=on}}

| map = {{Location map+ | Egypt

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Egypt

| lat_deg = 28.405556

| lon_deg = 28.805556

| mark = Lightgreen pog.svg

| marksize = 12

}}

}}

}}

The Bahariya Formation (also transcribed as Baharija Formation) is a fossiliferous geologic formation dating back to the early Cenomanian, which outcrops within the Bahariya depression in Egypt, and is known from oil exploration drilling across much of the Western Desert where it forms an important oil reservoir.{{cite journal | url=http://www.wuestenschiff.de/dateien/bahariya.pdf | title=Sequence stratigraphy of the Lower Cenomanian Bahariya Formation, Bahariya Oasis, Western Desert, Egypt | author=Catuneanu O., Khalifa M.A. & Wanas H.A. | journal=Sedimentary Geology | year=2006 | volume=190 | issue=1–4 | pages=121–137 | doi=10.1016/j.sedgeo.2006.05.010| bibcode=2006SedG..190..121C }}Weishampel, David B; et al (2004). "Dinosaur distribution (Late Cretaceous, Africa)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 604. {{ISBN|0-520-24209-2}}.{{cite book | chapter-url=http://sp.lyellcollection.org/content/132/1/201.abstract | title=Petroleum geology of North Africa|chapter=Mesozoic and Cenozoic petroleum systems of North Africa | publisher=Geological Society | author=Macgregor D.S. & Moody R.T.G. |series=Special Publications|volume=132|editor=Macgregor D.S. |editor2=Moody R.T.G. |editor3=Clark-Lowes D.D.| year=1998 | pages=201–216 | isbn=9781862390041}}

Extent

File:Geological map of the Bahariya Oasis.PNG

The Bahariya Formation forms the base of the depression, the lower part of the enclosing escarpment and all of the small hills within.{{cite journal | title=Sedimentology of the fluvial and fluvio-marine facies of the Bahariya Formation (Early Cenomanian), Bahariya Oasis, Western Desert, Egypt | author=Khalifa M.A. & Catuneanu O. | journal=Journal of African Earth Sciences | year=2008 | volume=51 | issue=2 | pages=89–103 | doi=10.1016/j.jafrearsci.2007.12.004| bibcode=2008JAfES..51...89K }} The type section for the formation is found at Gebel El-Dist, a hill at the northern end of the Bahariya depression.{{cite journal|title=Origin of ferricretes in fluvial-marine deposits of the Lower Cenomanian Bahariya Formation, Bahariya Oasis, Western Desert, Egypt |author=Tanner L.H. & Khalifa M.A. |journal=Journal of African Earth Sciences |year=2010 |volume=56 |issue=4–5 |pages=179–189 |doi=10.1016/j.jafrearsci.2009.07.004 |bibcode=2010JAfES..56..179T }}

Stratigraphy and sedimentology

Four depositional sequences have been recognised in the Bahariya Formation in the Bahariya depression, separated by three sub-aerial unconformities. The formation was deposited during a period of relative rise in sea level, with each unconformity representing a relative fall in sea level. Each of the individual sequences contains sediments deposited under fluvial, shoreline and shallow marine conditions.

[[Microfauna]] and [[Meiofauna]]

= Foraminifera =

class="wikitable" align="center" width="100%"
colspan="7" align="center" |Foraminifera of the Bahariya Formation
Genus

! Species

Charentia

| C. cuvillieri

Favusella

| F. washitensis

Mayncina

| M. orbignyi

Rotalipora

| R. cushmani
R. reicheli

Thomasinella

| T. aegyptia
T. fragmentaria
T. punica

Whiteinella

| W. archaeocretacea

= Other microorganisms =

class="wikitable" align="center" width="100%"
colspan="7" align="center" |Other microorganisms of the Bahariya Formation
Genus

! Species

! Images

Botryococcus

|

| rowspan="15" |File:Botryococcus_braunii.jpg]]File:Pediastrum_duplex_wagner.jpg]]
File:Scenedesmus_bijunga_EPA.jpg]]

Coronifera

| C. oceanica

Cyclonephelium

| C. edwardsii
C. vannophorum

Dynopterigium

| D. cladoides

Exochosphaeridium

|

Florentinia

| F. cooksoniae
F. mantlii

Kallosphaeridium

|

Mudrongia

| M. simplex

Palaeoperidinium

| P. cretaceum

Pediastrum

|

Pseudoceratium

| P. anaphrisum
P. securigerum

Scenedesmus

|

Spiniferites

|

Subtilisphaera

| S. perlucida
S. senegalensis

Xiphophoridium

| X. alatum

Invertebrates

= Molluscs =

class="wikitable" align="center" width="100%"
colspan="7" align="center" |Molluscs of the Bahariya Formation
Genus

! Species

! Notes

! Images

Baculites

|

| A heteromorph ammonite with a nearly straight shell.

| rowspan="7" | File:Baculites_grandis_shell.JPG.]]

Cardium

|

| A cockle.

Exogyra

|

| A reef-forming true oyster associated with solid substrates and warm temperatures.

Gastrochaena

|

| A saltwater clam.

Neolobites

| N. vibrayeanus

| A typical rolled ammonite.

Nucula

|

| A small saltwater clam.

Ostrea

| O. flabeata

| An edible oyster.

= Crustaceans (Ostracoda)=

class="wikitable" align="center" width="100%"
colspan="7" align="center" |Crustaceans of the Bahariya Formation
Genus

! Species

! Notes

! Images

Amphicytherura

| A. sexta

|

Anticythereis

| A. gaensis

|

Bairdia

| B. bassiounii
B. elongata

|

Brachycythere

| B. ledaforma porosa

|

Bythoceratina

| B. avnonensis
B. tamarae

|

Bythocypris

| B. eskeri

|

Cythereis

| C. algeriana
C. bicornis levis
C. canteriolata

|

Cytherella

| C. ovata
C. paenovata
C. parallela
C. sulcata

|

Fabanella

|

|

Looneyella

| L. sohni

|

Loxoconcha

| L. clinocosta
L. fletcheri

|

Metacytheropteron

| M. berbericum

|

Ovocytheridea

| O. caudata
O. producta
O. reniformis

|

Paracypris

| P. acutocaudata
P. angusta
P. mdaouerensis
P. triangularis

|

Pterygocythere

| P. raabi

|

Veeniacythereis

| V. jezzineensis

|

Xestoleberis

| X. obesa

|

= Insects =

Direct fossils are sparse, though plant leaves with extensive damage from folivorous insects have been documented.

Vertebrates

= Cartilaginous fish =

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Chondrichthyes of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Aegyptobatus

| A. kuehnei

|

| A sclerorhynchiform.

| rowspan="20" |File:Gymnura_aaustralis_csiro-nfc.jpg.]]
File:Onchopristis_numidus_052013.JPG tooth.]]
File:Rhinoptera_steindachneri.jpg.]]

File:Scapanorhynchus raphiodon.jpg]]

File:Squalicorax2DB.jpg]]

File:Squatina australis.jpg.]]

Asteracanthus

| A. aegyptiacus

|

| A hybodont shark.

Baharipristis

| B. bastetiae

|

| A sclerorhynchiform.

Cretodus

| C. longiplicatus

|

| A shark.

Cretolamna

| C. appendiculata

|

| A mackerel shark.

Distobatus

| D. nutiae

|

| A sclerorhynchiform.

Gymnura

| G. laterialata

|

| A butterfly ray.

Haimirichia

| H. amonensis

|

| A shark previously classified in the genera Odontaspis, Serratolamna, and Carcharias.

Isidobatus

| I. tricarinatus

|

| A sclerorhynchiform.

Marckgrafia

| M. lybica

|

| A sclerorhynchiform.

Onchopristis

| O. numida

| One complete cranium and associated vertebrae.

| A sclerorhynchid rajoid.

Peyeria

| P. libyca

|

| A sclerorhynchiform.

Ptychotrygon

| P. henkeli

|

| A sclerorhynchiform.

Renpetia

| R. labiicarinata

|

| A sclerorhynchiform.

Rhinoptera

|

|

| A batoid ray.

Schizorhiza

| S. stromeri

|

| An unusual sclerorhynchid sawfish that retained its rostral teeth after they were replaced.

Squalicorax

| S. baharijensis

|

| A large shark.

Scapanorhynchus

| S. subulatus

|

| A mitsukurinid similar to the modern goblin shark.

Squatina

|

|

| An angelshark.

Tribodus

| T. aschersoni

|

| A hybodont.

= Bony fish =

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Osteichthyes of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Bawitius

| B. bartheli

|

| A giant bichir.

| rowspan="15" |File:Ceratodus_BW.jpg]]
File:Enchodus_petrosus.jpg]]
File:Lepidotes_elvensis.JPG]]
File:MawsoniaDB16.jpg]]
File:Barramunda.jpg]]
File:Saurodon_leanusDB15.jpg]]

style="background:lightgrey;"

| Concavotectum

| C. moroccensis

|

| Possibly synonymous with Paranogmius doederleini.

Ceratodus

|

|

| A lungfish.

Coelodus

|

|

| A pycnodontid.

Enchodus

|

| One tooth.

| A predatory fish.

Lepidotes

|

| Isolated scales.

| Possibly misidentified from Bawitius bartheli.

Mawsonia

| M. lybica

| Considered a "signature taxon" of the formation.

| A giant freshwater coelacanth. Species assignation deemed provisional due to the lack of neotype.

Neoceratodus

| N. africanus

|

| A lungfish related to the living Australian species.

Obaichthys

|O. africanus

|

|An obaichthyid lepisosteiform. Remains formerly attributed to "Stromerichthys".{{Cite journal |last1=Pimentel |first1=Ricardo |last2=Barroso-Barcenilla |first2=Fernando |last3=Berrocal-Casero |first3=Mélani |last4=Callapez |first4=Pedro Miguel |last5=Ozkaya de Juanas |first5=Senay |last6=dos Santos |first6=Vanda F. |date=2023 |title=On the Occurrence of the Gar Obaichthys africanus Grande in the Cretaceous of Portugal: Palaeoecological and Palaeobiogeographical Implications |journal=Geosciences |language=en |volume=13 |issue=12 |pages=372 |doi=10.3390/geosciences13120372 |doi-access=free |bibcode=2023Geosc..13..372P |issn=2076-3263}}

Palaeonotopterus

| P. greenwoodi

|

| Nomen conservandum of the two Plethodus species previously described.

Paranogmius

| Paranogmius doederleini

| One vertebra.

| Holotype lost in World War II. Could be synonymous with Concavotectum moroccensis.

style="background:lightgrey;"

| Plethodus

| P. libycus
P. tibniensis

|

| Holotypes destroyed in World War II, but now believed to have been misidentified Palaeonotopterus greenwoodi.

Retodus

| R. tuberculatus

|

| A lungfish species identified from remains previously assigned to Neoceratodus.

style="background:#f3e9f3;"

| Saurodon

|

|

| Identified by Stromer in 1936, but now rejected due to this genus appearing only in post-Cenomanian Europe and North America. Now listed as Ichthyodectidae incertae sedis.

style="background:lightgrey;"

| Stromerichthys

| S. aethiopicus

|

| A fish initially identified as a bowfin relative, but now thought to be a chimera consisting of remains assignable to Bawitius, Obaichthys, and others.{{Cite journal |last1=Cavin |first1=Lionel |last2=Boudad |first2=Larbi |last3=Tong |first3=Haiyan |last4=Läng |first4=Emilie |last5=Tabouelle |first5=Jérôme |last6=Vullo |first6=Romain |date=2015 |title=Taxonomic composition and trophic structure of the continental bony fish assemblage from the early late cretaceous of Southeastern Morocco |journal=PLOS ONE |volume=10 |issue=5 |pages=e0125786 |doi=10.1371/journal.pone.0125786 |doi-access=free |issn=1932-6203 |pmc=4446216 |pmid=26018561|bibcode=2015PLoSO..1025786C }}

= Testudines =

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Testudines of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Apertotemporalis

| A. baharijensis

|

| A small pleurodiran turtle, likely belonging to Bothremydidae. Other unnamed species are also present.

|

= Squamates =

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Squamates of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Simoliophis

|

| Abundant

| First known sea snake, with functional hind legs. Now believed to include elements from different species and at least one of a different, unnamed genus.

|

= Plesiosaurs =

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Plesiosaurs of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

style="background:#f3e9f3;"

| Leptocleidus

| L. capensis

|

| A small plesiosaur that visited brackish or fresh water systems. Though known from both South Africa and England, the Egyptian material lacks diagnostic traits of the genus and is now referred as Polycotylidae incertae sedis.

| File:Leptocleidus1DB.jpg]]

= Crocodyliformes =

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Crocodyliformes of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Aegyptosuchus

| A. peyeri

|

| A poorly known, possibly stomatosuchid crocodile.

| rowspan="5" |File:August_1,_2012_-_Hamadasuchus_rebouli_Skull_on_Display_at_the_Royal_Ontario_Museum_(ROM_52620).jpg]]
File:Stomatosuchus2.jpg]]

Hamadasuchus

|H. rebouli

|

| A terrestrial, dog-like peirosaurid. It is possible that some material previously attributed to Libycosuchus actually belongs to this species.

Libycosuchus

| L. brevirostris

|

| A terrestrial crocodile of uncertain affinities, possibly related to Notosuchus.

Stomatosuchus

| S. inermis

| A complete cranium

| A large, flat-headed stomatosuchid with multiple small conical teeth, and possibly a pelican-like throat pouch. The only remains were destroyed in World War II.

style="background:lightgrey;"

| Stromerosuchus

| S. aegyptiacus

| Fragmentary remains

| Nomen dubium assigned to material found in 1911, that were badly damaged in 1922 while being shipped to Germany for study. Some material may belong to Aegyptosuchus and other to Stomatosuchus.

= Pterosaurs =

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Pterosaurs of the Bahariya Formation
Genus

! Species

! Material

! Notes

! Images

Pterosauria indet.{{Cite journal |last1=Salem |first1=Belal S. |last2=Sallam |first2=Hesham M. |last3=El-Sayed |first3=Sanaa |last4=Thabet |first4=Wael |last5=Antar |first5=Mohammed |last6=Lamanna |first6=Matthew C. |date=October 2019 |title=NEW DINOSAUR, PTEROSAUR, AND CROCODYLIFORM FOSSILS FROM THE UPPER CRETACEOUS (CENOMANIAN) BAHARIYA FORMATION OF THE BAHARIYA OASIS, EGYPT |url=https://www.researchgate.net/publication/344722122 |journal=Society of Vertebrate Paleontology (SVP) – Brisbane, Queensland, Australia}}

| Indeterminate

| Left first wing phalanx

| A medium-sized pterosaur. The first record of a pterosaur from Egypt.

= Dinosaurs =

== Sauropods ==

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Sauropods of the Bahariya Formation
Genus

! Species

! Material

! Notes

! Images

Aegyptosaurus

| A. baharijensis

| Partial postcranial skeleton"Table 13.1," in Weishampel, et al. (2004). Page 267.

| A titanosaur of about 15 meters. All remains were destroyed in World War II.

| rowspan="3" |File:Aegyptosaurus LM.png]]

File:dicraeosaurus hansemanni22.jpg]]


File:Paralititan stromeri.jpg]]

style="background:#f3e9f3;"

| Dicraeosaurus

| D. hansemanni

| Isolated scapula and vertebra

| Identified by Stromer in 1932. Subsequently considered a rebbachisaurid, or a third, unnamed titanosaur.

Paralititan

| P. stromeri

| Partial postcranial skeleton"Table 13.1," in Weishampel, et al. (2004). Page 269.

| One of the largest titanosaurs of the Cretaceous, with a 1.69 meters long humerus and an estimated total length of 26 meters.

== Theropods ==

{{paleobiota-key-compact}}

class="wikitable" align="center" width="100%"
colspan="5" align="center" |Theropods of the Bahariya Formation
Genus

! Species

! Abundance

! Notes

! Images

Abelisauridae indet.{{Cite journal |last1=Salem |first1=Belal S. |last2=Lamanna |first2=Matthew C. |last3=O'Connor |first3=Patrick M. |last4=El-Qot |first4=Gamal M. |last5=Shaker |first5=Fatma |last6=Thabet |first6=Wael A. |last7=El-Sayed |first7=Sanaa |last8=Sallam |first8=Hesham M. | year=2022 |title=First definitive record of Abelisauridae (Theropoda: Ceratosauria) from the Cretaceous Bahariya Formation, Bahariya Oasis, Western Desert of Egypt |journal=Royal Society Open Science |volume=9 |issue=6 |pages=220106 |doi=10.1098/rsos.220106|pmid=35706658 |pmc=9174736 |doi-access=free |bibcode=2022RSOS....920106S }}

|Indeterminate

|"MUVP 477, an isolated caudal (tenth) cervical vertebra"

|A medium-sized abelisaurid, estimated around 5.77 meters (~18.9 feet) long. The first unambiguous abelisaurid known from the Bahariya Formation.

| rowspan="9" |File:Bahariya abelisaurid McAfee.png

File:Bahariasaurus ingens, like megaraptora.jpg]]

Image:Tameryraptor markgrafi.png]]

File:Spinosaurus aegyptiacus.png]]

Bahariasaurus

| B. ingens

|

| A medium-sized theropod of uncertain affinities. Possibly a megaraptoran, a relative of Deltadromeus, or both.

Deltadromeus?

|Deltadromeus agilis?

|

|

Ceratosauria?

|

|

|

style="background:#f3e9f3;"

| Elaphrosaurus

|E. bambergi or aff

|

| Material now considered to be indeterminate theropod remains.

Erectopus?

|Erectopus sauvagei?

|

|

style="background:pink"

| Sigilmassasaurus

| S. brevicollis

|

| Previously considered a species of Spinosaurus, or a synonym of S. aegyptiacus.

Spinosaurus

| S. aegyptiacus

| Most common dinosaur of the formation.

| A large spinosaurid.

Tameryraptor{{Cite journal |last1=Kellermann |first1=Maximilian |last2=Cuesta |first2=Elena |last3=Rauhut |first3=Oliver W. M. |date=2025-01-14 |title=Re-evaluation of the Bahariya Formation carcharodontosaurid (Dinosauria: Theropoda) and its implications for allosauroid phylogeny |url=https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0311096 |journal=PLOS One |language=en |volume=20 |issue=1 |pages=e0311096 |doi=10.1371/journal.pone.0311096 |issn=1932-6203 |doi-access=free|pmc=11731741 }}

| T. markgrafi

|

| A large carcharodontosaurid theropod, originally assigned to Carcharodontosaurus

In addition, there are isolated theropod teeth disputedly assigned to dromaeosaurids, or to abelisaurids.

Flora

Thirty different genera are known from Bahariya, including megaflora. Much of the material is yet to be described.{{Cite journal |last1=Lejal-Nicol |first1=A |last2=Dominik |first2=W. |date=1990 |title=Sur la paleoflore a Weichseliaceae et a angiospermes du Cenomanien de la region de Bahariya (Egypte du Sud-Ouest) |journal=Berliner geowissenschaftliche Abhandlungen A |volume=120 |issue=5 |pages=957–991}}Ijouhier, Jamale (2016) A reconstruction of the palaeoecology and environmental dynamics of the Bahariya Formation of Egypt. PeerJ Preprints, https://doi.org/10.7287/peerj.preprints.2470v1 Other taxa include Sapindales, Piperaceae, Lauraceae, Platanaceae, Magnoliopsida, Nymphaeaceae, Cornaceae, Proteaceae and Vitaceae not identified at genus level; and miospore and pollen species.{{Cite journal |last1=El Atfy |first1=Haytham |last2=Coiffard |first2=Clément |last3=El Beialy |first3=Salah Y. |last4=Uhl |first4=Dieter |date=2023-01-30 |title=Vegetation and climate change at the southern margin of the Neo-Tethys during the Cenomanian (Late Cretaceous): Evidence from Egypt |journal=PLOS ONE |volume=18 |issue=1 |pages=e0281008 |doi=10.1371/journal.pone.0281008 |doi-access=free |pmid=36716334 |pmc=9886267 |bibcode=2023PLoSO..1881008E |issn=1932-6203}}{{Cite journal |last1=El Atfy |first1=Haytham |last2=Coiffard |first2=Clément |last3=Uhl |first3=Dieter |last4=Spiekermann |first4=Rafael |last5=El Khoriby |first5=Essam M. |last6=Aleraky |first6=Heba |last7=Mohamed |first7=Ahmed |date=2023 |title=A new florula dominated by angiosperms from the Cenomanian of Egypt |url=http://dx.doi.org/10.1016/j.cretres.2023.105554 |journal=Cretaceous Research |volume=149 |pages=105554 |doi=10.1016/j.cretres.2023.105554 |bibcode=2023CrRes.14905554E |issn=0195-6671}}

class="wikitable" align="center"
colspan="7" align="center" | Vascular Plants
GenusSpeciesAbundanceNotesImages
style="background:#D1FFCF;" | Agathis{{Cite journal |last1=Lyon |first1=M. A. |last2=Johnson |first2=K. R. |last3=Wing |first3=S. L. |last4=Nichols |first4=D. J. |last5=Lacovara |first5=K. J. |last6=Smith |first6=J. B. |date=2001 |title=Late Cretaceous equatorial coastal vegetation: new megaflora associated with dinosaur finds in the Bahariya Oasis, Egypt |url=https://www.researchgate.net/publication/235708368 |journal=Geological Society of America, Annual Meeting |pages=198}}

|style="background:#D1FFCF;" |A/W spp.

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" | An Araucarian conifer, now restricted to Australasia.

| rowspan="18" style="background:#D1FFCF;" |

File:Cladophlebis_asiatica_IMG_5157_Beijing_Museum_of_Natural_History.jpg]]

File:Sapindopsis anhouryi.JPG]]

File:Weichselia reticulata.jpg]]

style="background:#D1FFCF;" |Araliaephyllum?

|style="background:#D1FFCF;" |Indeterminate ("Morphotype-14")

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Suggested to be related with Lauraceae

style="background:#D1FFCF;" | Cladophlebis

|style="background:#D1FFCF;" |C. spp.

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" | Fern Laflets

style="background:#D1FFCF;" |Cornophyllum

|style="background:#D1FFCF;" |C. distense

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Suggested to be related with Cornaceae

style="background:#D1FFCF;" |Cinnamophyllum?

|style="background:#D1FFCF;" |Indeterminate ("Morphotype-12")

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Suggested to be related with Lauraceae

style="background:#D1FFCF;" |Eucalyptolaurus/"Myrtophyllum”(?)

|style="background:#D1FFCF;" |Indeterminate ("Morphotype-08")

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Specimens of the family Lauraceae

style="background:#D1FFCF;" |Laurophyllum

|style="background:#D1FFCF;" |L. africanum

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Specimens of the family Lauraceae

style="background:#D1FFCF;" | Liriophyllum

|style="background:#D1FFCF;" |L. farafraense

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Specimens of the family Magnoliaceae

style="background:#D1FFCF;" |Marsilea

|style="background:#D1FFCF;" |aff. Marsilea spp.

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" | Water fern.

style="background:#D1FFCF;" |Magnoliid

|style="background:#D1FFCF;" |

  • "Morphotype-01"
  • "Morphotype-04"
  • "Morphotype-09"
  • "Morphotype-11"
  • "Morphotype 21"?

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Shows festooned brochidodromous venation

style="background:#D1FFCF;" |Magnoliaephyllum

|style="background:#D1FFCF;" |

  • M. auriculatum
  • M. bahariyense
  • M. isbergiana

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Possible affinities with Lauraceae

style="background:#D1FFCF;" | Nelumbites

|style="background:#D1FFCF;" |

  • N. schweinfurthi
  • N. giganteum

|style="background:#D1FFCF;" |Common occurrence in the lower shale bed

|style="background:#D1FFCF;" | Typical leaves of the faimily Nelumbonaceae

style="background:#D1FFCF;" |Paradoxopteris

|style="background:#D1FFCF;" |P. stromeri

|style="background:#D1FFCF;" |Co-Dominant plant

|style="background:#D1FFCF;" |Xerophytic tree fern suggestive of a dry tropical climate.

style="background:#D1FFCF;" |Plumafolium?

|style="background:#D1FFCF;" |Indeterminate ("Morphotype-18")

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |A Monocot, probably related with Liliopsida

style="background:#D1FFCF;" |Rogersia

|style="background:#D1FFCF;" |R. longifolia

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Likely lobes of a much bigger leaf of Sapindopsis type

style="background:#D1FFCF;" |Typhaephyllum

|style="background:#D1FFCF;" |cf. T. sp.

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Interpreted as a monocot, probably related to Typhaceae

style="background:#D1FFCF;" |Vitiphyllum

|style="background:#D1FFCF;" |V. aff. multifidum

|style="background:#D1FFCF;" |Few Specimens

|style="background:#D1FFCF;" |Some similarities with Pabiania of the family Lauraceae

style="background:#D1FFCF;" | Weichselia

|style="background:#D1FFCF;" | W. reticulata

|style="background:#D1FFCF;" |Dominant plant

|style="background:#D1FFCF;" | Xerophytic tree fern suggestive of a dry tropical climate.

See also

References