Haplogroup O-M119

The Haplogroup O-M119 branch is believed to have evolved during the Late Pleistocene (Upper Paleolithic) in China mainland.{{Citation needed|date=December 2012}}

{{harvtxt|Karafet|Hallmark|Cox|Sudoyo|2010}} suggest haplogroup O-M119 was part of a four-phase colonization model in which Paleolithic migrations of hunter-gatherers shaped the primary structure of current Y-Chromosome diversity of Maritime Southeast Asia. Approximately 5000 BCE, Haplogroup O-M119 coalesced at Sundaland and migrated northwards to as far as Taiwan, where O-M50 constitutes some 90% of the Aboriginal Y-DNA, being the main haplogroup that can be directly linked to the Austronesian expansion in phase 3.

The Liangdao man, an 8,000 year old skeleton found on Liang Island in the Republic of China off the coast of Fujian, is believed to belong to Haplogroup O-M119, specifically under branch O-CTS5726.http://www.taipeitimes.com/News/taiwan/archives/2014/12/13/2003606642|archiveurl=https://web.archive.org/web/20200811140358/http://www.taipeitimes.com/News/taiwan/archives/2014/12/13/2003606642M{{cite web | url=https://discover.familytreedna.com/y-dna/O-CTS5726/ancient | title=Welcome to FamilyTreeDNA Discover }}

{{harvtxt|Li|Wen|Chen|Su|2008}} concluded that in contrast to the Taiwan homeland hypothesis, Island Southeast Asians do not have a Taiwan origin based on their paternal lineages. According to their results, lineages within Maritime Southeast Asia did not originate from Taiwanese aborigines as linguistic studies suggest. Taiwan aborigines and Indonesians were likely to have been derived from the Tai–Kadai-speaking populations based on their paternal lineages, and thereafter evolved independently of each other.

The strongest positive correlation between Haplogroup O-M119 and ethno-linguistic affiliation is that which is observed between this haplogroup and the Austronesians. The peak frequency of Haplogroup O-M119 is found among the aborigines of Taiwan, precisely the region from which linguists have hypothesized that the Austronesian language family originated. A slightly weaker correlation is observed between Haplogroup O-M119 and the Han Chinese populations of southern China, as well as between this haplogroup and the Tai–Kadai-speaking populations of southern China and Southeast Asia. The distribution of Tai–Kadai languages in Thailand and other parts of Southeast Asia outside of China has long been believed, for reasons of traditional linguistic geography, to reflect a recent invasion of Southeast Asia by Tai–Kadai-speaking populations originating from southeastern China, and the somewhat elevated frequency of Haplogroup O-M119 among the Tai–Kadai populations, coupled with a high frequency of Haplogroup O-M95, which is a genetic characteristic of the Austroasiatic-speaking peoples of Southeast Asia, suggests that the genetic signature of the Tai–Kadai peoples' affinity with populations of southeastern China has been weakened due to extensive assimilation of the earlier Austroasiatic residents of the lands which the Tai–Kadai peoples invaded.

Haplogroup O-M119 lineages are found primarily in Southeast Asian populations of Malaysia, Vietnam, Indonesia, the Philippines, southern China and Taiwan {{harv|ISOGG|2010}}. High frequencies of this haplogroup have been found in populations spread in an arc through southeastern China, Taiwan, the Philippines, and Indonesia. It has been found with generally lower frequency in samples from Oceania, mainland Southeast Asia, Southwest China, Northwest China, North China, Northeast China, Korea, Japan, North Asia, and Central Asia.

A 2008 study by Li suggested that the admixture analyses of Tai–Kadai-speaking populations showed a significant genetic influence in a large proportion of Indonesians. Most of the population samples contained a high frequency of haplogroup O-M119.{{harv|Li|Wen|Chen|Su|2008}}

The frequencies of Haplogroup O-M119 among various East Asian and Austronesian populations suggest a complex genetic history of the modern Han populations of southern China.{{citation needed|date=December 2012}} Although Haplogroup O-M119 occurs only at an average frequency of approximately 4% among Han populations of northern China and peoples of southwestern China and Southeast Asia who speak Tibeto-Burman languages, the frequency of this haplogroup among the Han populations of southern China nearly quadruples to about 15-23%.{{cite journal|last1=Yan|first1=Shi|title=An updated tree of Y-chromosome Haplogroup O and revised phylogenetic positions of mutations P164 and PK4|journal= European Journal of Human Genetics|pmc=3179364|pmid=21505448|doi=10.1038/ejhg.2011.64|volume=19|issue=9|date=September 2011|pages=1013–5}} The frequency of Haplogroup O-M119 among the Southern Han has been found to be slightly greater than the arithmetic mean of the frequencies of Haplogroup O-M119 among the Northern Han and a pooled sample of Austronesian populations. This suggests that modern Southern Han populations may possess a non-trivial number of male ancestors who were originally affiliated with some Austronesian-related culture, or who at least shared some genetic affinity with many of the ancestors of modern Austronesian peoples.{{cite journal|last1=HUGO Pan-Asian SNP Consortium|first1=HUGO Pan-Asian SNP Consortium|title=Mapping Human Genetic Diversity in Asia|journal=Science|volume=326|issue=5959|pages=1541–1545|doi=10.1126/science.1177074|pmid=20007900|bibcode=2009Sci...326.1541.|year=2009|s2cid=34341816}}{{cite journal|last1=HUGO Pan-Asian SNP Consortium|first1=HUGO Pan-Asian SNP Consortium|title=Mapping Human Genetic Diversity in Asia|url=https://www.science.org/doi/10.1126/science.1177074|journal=Science|year=2009 |volume=326 |issue=5959 |pages=1541–1545 |publisher=Science Magazine|doi=10.1126/science.1177074 |pmid=20007900 |bibcode=2009Sci...326.1541. |s2cid=34341816 |access-date=11 December 2009}}

This lineage is found frequently in Austronesians, southern Han Chinese, and Kra-Dai peoples.{{cite web|title=Major East–West Division Underlies Y Chromosome Stratification across Indonesia|url=http://mbe.oxfordjournals.org/content/suppl/2010/02/28/msq063.DC1|archive-url=https://archive.today/20140608191525/http://mbe.oxfordjournals.org/content/suppl/2010/02/28/msq063.DC1|url-status=dead|archive-date=2014-06-08|publisher=Karafet et al|access-date=15 May 2010}} This lineage is presumed to be a marker of the prehistoric Austronesian expansion, with possible origins encompassing the regions along the southeastern coast of China and neighboring Taiwan, and is found among modern populations of Maritime Southeast Asia and Oceania {{harv|Karafet|2005}}.

Haplogroup O-M119 Y-chromosomes also have been found to occur at low frequency among various populations of Siberia, such as the Nivkhs (one of 17 sampled Y-chromosomes), Ulchi/Nanai (2/53), Yenisey Evenks (1/31), and especially the Buryats living in the Sayan-Baikal uplands of Irkutsk Oblast (6/13) {{harv|Lell|2002}}.

O-P203 was found in 86.7% (52/60) of a sample from Nias, 70.8% (34/48) of Taiwanese Aboriginals, 28.4% (21/74) of Mentawai, 11.4% (73/641) of Balinese, 9.8% (6/61) of a sample from Java, 9.1% (36/394) of a sample from Flores, 9.1% (15/165) of Han Chinese, 8.3% (1/12) of a sample from Western Samoa, 8.2% (4/49) of Tujia from Hunan, 6.9% (4/58) of Miao from China, 5.7% (4/70) of Vietnamese, 3.3% (1/30) of a sample from the Moluccas, 3.1% (1/32) of Malaysians, 3.0% (1/33) of a sample from highland Papua New Guinea, 2.6% (1/38) of a sample from Sumatra, 2.3% (2/86) of a sample from Borneo, 2.1% (1/48) of Filipinos, 2.0% (1/51) of She, 1.7% (1/60) of Yao from Guangxi, 1.1% (1/92) of a sample from Lembata, and 0.9% (3/350) of a sample from Sumba.{{cite journal|last1=Karafet|first1=Tatiana|title=Major East–West Division Underlies Y Chromosome Stratification across Indonesia|journal=Molecular Biology and Evolution|volume=27|issue=8|pages=1833–1844|doi=10.1093/molbev/msq063|pmid=20207712|year=2010|doi-access=free}}

In a study published in 2011, O-P203 was observed in 22.2% (37/167) of Han Chinese male volunteers at Fudan University in

Shanghai whose origin may be traced back to East China (Jiangsu, Zhejiang, Shanghai, or Anhui), 12.3% (8/65) of Han Chinese male volunteers whose origin may be traced back to South China, and 1.6% (2/129) of Han Chinese male volunteers whose origin may be traced back to North China.Shi Yan, Chuan-Chao Wang, Hui Li, Shi-Lin Li, Li Jin, and The Genographic Consortium, "An updated tree of Y-chromosome Haplogroup O and revised phylogenetic positions of mutations P164 and PK4." European Journal of Human Genetics (2011) 19, 1013–1015; doi:10.1038/ejhg.2011.64; published online 20 April 2011.

This lineage was observed in one individual from China {{harv|Underhill|2000}} and another from Kota Kinabalu {{harv|Hurles|2005}}.

According to the website of Chinese genetic testing company 23mofang, O-M101 is a subclade of O-M307/P203 (O-M307 > O-F446 > O-F5498 > O-Z23406 > O-M101). Its TMRCA is estimated to be 4,850 years before present, and it is estimated to account for the Y-DNA of approximately 0.21% of all males in present-day China, with its distribution being relatively dense in Hunan, Hubei, Hainan, and Jiangxi.{{cite web | url=https://www.23mofang.com/ancestry/ytree/O-M101/detail | title=O-M101单倍群详情 }} The O-M101 > O-A5863 > O-SK1573 subclade (TMRCA 3,400 ybp) has been estimated to account for the Y-DNA of approximately 0.08% of all males in present-day China, being relatively concentrated in South Central China and Southwest China at present.{{cite web | url=https://www.23mofang.com/ancestry/ytree/O-SK1573/detail | title=O-Sk1573单倍群详情 }} The O-M101 > O-A5863 > O-Y163909 subclade (TMRCA 4,080 ybp) has been observed in 16.7% (3/18) of a sample of Phuan males from Central Thailand.[https://www.theytree.com/tree/O-M119 Phylogenetic tree of haplogroup O-M119 at TheYtree]

This lineage occurs among Austronesian peoples of Taiwan, the Philippines, Indonesia, Melanesia, Micronesia, and Madagascar as well as among some populations of continental Southeast Asia and among Bantu peoples of the Comoros.{{cite journal |ref={{harvid|Msaidie|2010}} |last1=Msaidie |first1=Said |last2=Ducourneau |first2=Axel |last3=Boetsch |first3=Gilles |last4=Longepied |first4=Guy |last5=Papa |first5=Kassim |last6=Allibert |first6=Claude |last7=Yahaya |first7=Ali Ahmed |last8=Chiaroni |first8=Jacques |last9=Mitchell |first9=Michael J |title=Genetic diversity on the Comoros Islands shows early seafaring as major determinant of human biocultural evolution in the Western Indian Ocean |journal=European Journal of Human Genetics |date=January 2011 |volume=19 |issue=1 |pages=89–94 |doi=10.1038/ejhg.2010.128|pmid=20700146 |pmc=3039498 }} It also has been found in a Hawaiian.Swapan Mallick, Heng Li, Mark Lipson, et al., "The Simons Genome Diversity Project: 300 genomes from 142 diverse populations." Nature 538, 201–206 (13 October 2016) doi:10.1038/nature18964

A study published in 2005 found O-M50 in 33.3% (13/39) of a sample of aboriginals in Taiwan, 18.2% (2/11) of a sample of people in Majuro, 17.1% (6/35) of a sample of Malagasy, 9.2% (6/65) of a sample of people in Kota Kinabalu, 9.1% (2/22) of a sample of people in Banjarmasin, 3.6% (1/28) of a sample of people in the Philippines, and 1.9% (1/52) of a sample of people in Vanuatu.{{cite journal|last1=Hurles|first1=Matthew E.|last2=Sykes|first2=Bryan C.|last3=Jobling|first3=Mark A.|last4=Forster|first4=Peter|year=2005|pages=894–901|title=The Dual Origin of the Malagasy in Island Southeast Asia and East Africa: Evidence from Maternal and Paternal Lineages|issue=5|journal=American Journal of Human Genetics|volume=76|pmid=15793703|pmc=1199379|doi=10.1086/430051 }}

Kayser et al. 2008 found O-M110 in 34.1% (14/41) of a sample of Taiwan Aborigines, 17.7% (26/147) of a sample from the Admiralty Islands, 17.3% (9/52) of a sample from the Trobriand Islands, 13.5% (5/37) of a sample from the Philippines, 9.7% (3/31) of a sample from the Nusa Tenggara Islands, 3.8% (2/53) of a sample from Java, 3.0% (1/33) of a sample from the Moluccas, 2.5% (1/40) of a sample from Borneo, 1.0% (1/100) of a sample from Tuvalu, and 0.95% (1/105) of a sample from Fiji.Manfred Kayser, Ying Choi, Mannis van Oven, Stefano Mona, Silke Brauer, Ronald J. Trent, Dagwin Suarkia, Wulf Schiefenhövel, and Mark Stoneking (2008), "The Impact of the Austronesian Expansion: Evidence from mtDNA and Y Chromosome Diversity in the Admiralty Islands of Melanesia." Mol. Biol. Evol. 25(7):1362–1374. doi:10.1093/molbev/msn078

A study published in 2010 found O-M110 in 18.8% (9/48) Taiwanese Aboriginals, 13.3% (8/60) Nias, 8.3% (4/48) Philippines, 7.4% (4/54) Sulawesi, 6.3% (22/350) Sumba, 5.8% (5/86) Borneo, 3.3% (1/30) Moluccas, 2.3% (1/44) Maewo, Vanuatu, 1.6% (1/61) Java, 1.4% (1/74) Mentawai, and 0.8% (5/641) Bali.{{sfn|Karafet|Hallmark|Cox|Sudoyo|2010}}

A study published in 2012 found O-M110 in 4.6% (33/712) of males from the Solomon Islands.Frederick Delfin, Sean Myles, Ying Choi, David Hughes, Robert Illek, Mannis van Oven, Brigitte Pakendorf, Manfred Kayser, and Mark Stoneking, "Bridging Near and Remote Oceania: mtDNA and NRY Variation in the Solomon Islands." Molecular Biology and Evolution 29(2):545–564. 2012 doi:10.1093/molbev/msr186.

{{main|Conversion table for Y chromosome haplogroups}}

Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.

The following research teams per their publications were represented in the creation of the YCC tree.

{{columns-list|colwidth=22em|

• α {{harvnb|Jobling and Tyler-Smith|2000}} and {{harvnb|Kaladjieva|2001}}
• β {{harvnb|Underhill|2000}}
• γ {{harvnb|Hammer|2001}}
• δ {{harvnb|Karafet|2001}}
• ε {{harvnb|Semino|2000}}
• ζ {{harvnb|Su|1999}}
• η {{harvnb|Capelli|2001}}

}}

This phylogenetic tree of haplogroup O subclades is based on the YCC 2008 tree {{harv|Karafet|2008}} and subsequent published research.

• O-M119
• O-M119* China (Sichuan, Guizhou, Hubei, Shanghai, Zhejiang, Guangdong, Henan, Hebei, Liaoning, etc.)
• O-Y14027
• O1a3-Y144065/F1036 China (esp. Zhejiang, Fujian, Jiangsu, and Guangdong)
• O-Y87942/MF38142 China (Zhejiang, Jiangsu, Sichuan, Chongqing, Hubei, etc.)
• O-F1009 China (Shandong, Jiangsu, Hebei, Zhejiang, Beijing, Liaoning, Henan, Shanghai, Guangdong, Anhui, Shanxi, Jilin, Sichuan, Heilongjiang, Tianjin, Hubei, Shaanxi, Hunan, Gansu, etc.), North Korea (Pyeongyang), South Korea
• O1a2-M50/M103/M110/F3288 Austronesia, China (Beijing, Macau), Thailand (Tai Lue,{{cite journal |ref={{harvid|Brunelli|2017}} |last1=Brunelli |first1=Andrea |last2=Kampuansai |first2=Jatupol |last3=Seielstad |first3=Mark |last4=Lomthaisong |first4=Khemika |last5=Kangwanpong |first5=Daoroong |last6=Ghirotto |first6=Silvia |last7=Kutanan |first7=Wibhu |title=Y chromosomal evidence on the origin of northern Thai people |journal=PLOS ONE |date=24 July 2017 |volume=12 |issue=7 |pages=e0181935 |doi=10.1371/journal.pone.0181935|pmid=28742125 |pmc=5524406 |doi-access=free |bibcode=2017PLoSO..1281935B }} Yong, Tai Dam from Loei Province, Iu Mien,Wibhu Kutanan, Rasmi Shoocongdej, Metawee Srikummool, et al. (2020), "Cultural variation impacts paternal and maternal genetic lineages of the Hmong-Mien and Sino-Tibetan groups from Thailand." European Journal of Human Genetics https://doi.org/10.1038/s41431-020-0693-x Isan, Mon, Siamese, Khon Muang), Laos (Lao from Luang Prabang)
• O1a2a (F3288)
• O1a2a1 (B392, Z38606, Z38607, Z38608, Z38609, Z38610, Z38611)
• O1a2a1a (B393, Z38612)
• O1a1-B384/Z23193
• O1a1a (M307.1/P203.1, CTS3422, CTS4351, CTS5059, CTS6864, CTS7015, CTS8229, CTS8875, CTS8934, CTS9321, CTS11688, F31, F54, F89, F303, F333, L83)
• O1a1a1 (F446, CTS4588, F560, FGC15381/K620/Z23389, K613/Z23387, K619/Z23388)
• O1a1a1a (F140, CTS611, CTS3089, CTS3265, CTS3269, CTS11270, F157, F343, F424, F518, F571, FGC15382/Z23466, FGC15391/Y14275/Z23470, FGC15383/K625/Z23474, V68.2, Z23457)
• O1a1a1a* Indonesia
• O1a1a1a1 (F78, CTS4478) China (esp. Shanghai, Jiangsu, Zhejiang[https://www.23mofang.com/ancestry/ytree/O-M119 Phylogenetic Tree of Haplogroup O1a-M119 at 23mofang])
• O-F23879 Philippines (Igorot from Mountain Province, Negros Occidental)
• O-MF14277 China (Hunan, Sichuan, Guizhou, Jiangxi, Guangdong, Hubei, etc.)
• O-MF14175 China (esp. Hunan, Sichuan, and Chongqing)
• O-Y158653/ACT6252 China (Hebei, Jiangxi, Ma'anshan)
• O1a1a1a1a (F81, CTS4910, CTS5709, FGC15392/Y14265/Z23469) China, Philippines, Vietnam (Pathen from Quang Bình District, Tày from Mường Khương District and Cư Jút District, Dao from Hoàng Su Phì District), Thailand (Pray, Isan, Mon,Wibhu Kutanan, Jatupol Kampuansai, Metawee Srikummool, et al. (2019), "Contrasting Paternal and Maternal Genetic Histories of Thai and Lao Populations." Mol. Biol. Evol. Advance Access publication April 12, 2019. doi:10.1093/molbev/msz083 Central Thai, Northern Thai)
• O1a1a1a1a* Guangdong
• O1a1a1a1a1 (CTS2458)
• O1a1a1a1a1a (F533)
• O1a1a1a1a1a1 (F492, CTS2498, CTS2594, CTS4206,CTS5075, CTS11784, F619, K629/Z23478)
• O1a1a1a1a1a1a (F656, Z23481) Hubei, Jiangsu, Hong Kong, Beijing, Xishuangbanna
• O1a1a1a1a1a1a1 (A12440) Jiangsu, Anhui
• O1a1a1a1a1a1a1a (A12439)
• O1a1a1a1a1a1a2 (A14788, A14789)
• O1a1a1a1a1a1a3 (F65, F285, F469)
• O1a1a1a1a1a1a4 (MF1068, MF1069, MF1070)
• O1a1a1a1a1a1a5 (Z23482, Z23484, Z23485, Z23486, Z23487, Z23488, Z23489)
• O1a1a1a1a1a1b (FGC66168, Z23496, Z23505) Guangdong, Guangxi, Jiangsu
• O1a1a1a1a1a1b1 (CTS11553, Z23494, Z23496, Z23503)
• O1a1a1a1a1a1b2 (CTS409, CTS2613, CTS5922) Fujian
• O1a1a1a1a1a1b3 (Y146786, Y146790, Y146817) Hubei, Jiangxi
• O1a1a1a1a1a1c (Y31266, Y31267)
• O1a1a1a1a1a1c* Jiangxi, Fujian
• O1a1a1a1a1a1c1 (Y31261, Y31262, Y31263, Y31264, Y31265, Y31643) Sichuan, Cambodia
• O1a1a1a1a1a1d (A12441, A12442, A12443, Y69066) Henan, Anhui
• O1a1a1a1a1a1e (MF1071, MF1072, MF1073) Jiangsu, Anhui
• O1a1a1a1a1a1e1 (MF1074)
• O1a1a1a1a1a2 (CTS4585, CTS7624, FGC15395/Y14269, FGC15397/Y14272, FGC15398/Y13987, FGC15399/Y13986, FGC15400/Y13989, FGC15401/Y13988, FGC15402/Y13984, FGC15409/Y13985, FGC47328, FGC47336, Y14273) Jiangsu, Beijing
• O1a1a1a1a2 (MF1075, MF1076, MF1077, MF1078, MF1079, MF1080, MF1081, MF1082) Jiangsu, Shanghai
• O1a1a1a2 (YP4610/Z39229)
• O1a1a1a2a (AM00330/AMM480/B386, SK1533, Z39230/YP4600, Z39231/YP4601, Z39232/YP4603, Z39233/YP4605, Z39234/YP4607, Z39235/YP4609, Z39236/YP4611, Z39237/YP4612, Z39238/YP4613, Z39239/YP4614, Z39240/YP4615, Z39241/YP4616, Z39242/YP4617, Z39243/YP4618, Z39244/YP4619, Z39245/YP4620, Z39246/YP4621)
• O1a1a1a2a1 (AM00333/AMM483/B387, Z39247/YP4602) Philippines, Singapore (MalayMonika Karmin, Lauri Saag, Mário Vicente, et al., "A recent bottleneck of Y chromosome diversity coincides with a global change in culture." Genome Research (2015) 25: 459-466. doi: 10.1101/gr.186684.114)
• O1a1a1a2a1a (B388, Z39248/YP4604, Z39249/YP4608) Philippines, Singapore (Malay)
• O1a1a1a2b (SK1555)
• O1a1a1b (SK1568/Z23420, CTS8920, Z23430)
• O1a1a1b1 (M101) China{{cite journal|last1=Underhill|first1=PA|last2=Shen|first2=P|last3=Lin|first3=AA|last4=Jin|first4=L|last5=Passarino|first5=G|last6=Yang|first6=WH|last7=Kauffman|first7=E|last8=Bonné-Tamir|first8=B|last9=Bertranpetit|first9=J|title=Y chromosome sequence variation and the history of human populations|journal=Nature Genetics|volume=26|issue=3|pages=358–61|year=2000|pmid=11062480|doi=10.1038/81685 |s2cid=12893406}} (Guizhou), Thailand (Phuan from Central Thailand), Kota Kinabalu
• O1a1a1b2 (Z23392, Z23404, Z23440/SK1572) Vietnam (Colao from Hoàng Su Phì District), Thailand (Phutai from Sakon Nakhon Province, Siamese from Western Thailand and Eastern Thailand, Mon from Northern Thailand)
• O1a1a1b2* Ho Chi Minh City
• O1a1a1b2a (Z23442, F26575) Xishuangbanna, Hong Kong, Ho Chi Minh City
• O1a1a1b2a1 (SK1571, Z39268)
• O1a1a2 (CTS8423, CTS2915, F4084) China (Jiangsu, Shanghai, Zhejiang, Hubei, Sichuan, etc.)
• O1a1a2a (CTS52, CTS11785, CTS5880) Japan[https://www.familytreedna.com/groups/o-3/dna-results O Y-Haplogroup Project] at Family Tree DNA
• O1a1a2a1 (CTS701, K583, CTS4115, CTS6217) Vietnam (Lachi and Dao from Hoàng Su Phì District, Hanhi and Sila from Mường Tè District, Hmong from Điện Biên Phủ, Tay from Bình Liêu DistrictEnrico Macholdt, Leonardo Arias, Nguyen Thuy Duong, et al., "The paternal and maternal genetic history of Vietnamese populations." European Journal of Human Genetics (2020) 28:636–645. https://doi.org/10.1038/s41431-019-0557-4), Thailand (Tai Lue from Northern Thailand, Shan, Suay from Northeast Thailand, Skaw Karen from Mae Hong Son Province, Isan, Phuan from Central Thailand, Black Tai from Loei Province, Mon, Northern Thai), Laos (Lao from Vientiane)
• O1a1a2a1a (K644/Z23266, Z23269)
• O1a1a2a1a1 (CTS10805, MF2376, CTS4829)
• O-CTS10805* Hunan
• O-Y148878 Guangdong, Sichuan
• O-MF2375
• O-MF2375* Hunan, Beijing
• O-MF6454 Singapore, Indonesia (Greater Jakarta)
• O1a1a2a1a2 (Z23274, Z23275, F24990, F25139)
• O-Z23274* Shandong
• O-YP345 (Z7773, Z23304/Z23304.2) Xishuangbanna Dai
• O-Z23338/Y18196 Xishuangbanna Dai
• O1a1a2a1a3 (CTS9421, CTS3144, CTS9476) China (Fujian Han), Taiwan (Yilan), Singapore, Japan[https://www.familytreedna.com/public/y-dna-haplotree/O;name=O-M119 Y-DNA Haplotree at FamilyTreeDNA]
• O1a1a2a1a4 (Y168496, MF167165/Y192368, Y168556, Y168502) China (Jiangsu)
• O1a1a2a1b (Y157651)
• O-Y156772 Shandong
• O-CTS2118
• O-CTS2118* Singapore
• O-F16334 (CTS11040, CTS1992) Beijing, Guangdong, Liaoning
• O1a1a2a2 (Y89818, MF16617)
• O1a1a2a2a (MF14481, MF17536, MF16199) Zhejiang, Shandong
• O1a1a2a2b (FGC66100, FGC66137, FGC66127) Hubei, Jiangxi, Zhejiang, Hebei, North Korea
• O1a1a2b (F2444, F4243, Y137046) China (Guangdong, Jiangsu, Sichuan, Hubei, Zhejiang, Shandong, Jiangxi, etc.)
• O1a1a2b1-Y137055 China (Found sporadically in Shandong, Hebei, Hong Kong, Hubei, Zhejiang, etc.)
• O-Y137185 China (Guangdong, Sichuan, Jiangsu, Hubei, Shanghai, etc.), Hungary (Győr-Moson-Sopron)
• O1a1a2b2-MF6180 China (Found sporadically in Shaanxi, Henan, Liaoning, Jilin, Heilongjiang, Shanxi), South Korea
• O-F1056 China (Jiangsu, Shanghai, Zhejiang, Shandong, Anhui, etc.)
• O-MF6458 China (esp. Jiangsu, Shanghai, Zhejiang)
• O-MF6284 ''China (Found sporadically in Jiangsu, Shanghai, Henan, Sichuan)
• O1a1a2c-SK1522 China (Jiangsu, Zhejiang, Shanghai, etc.)
• O1a1b (CTS5726, CTS3085, CTS3400) China (Beijing, Fujian), Thailand (Siamese from Western Thailand), the Philippines (Manila, Agta), Singapore (Malay). Also found in "Liangdao Man", an 8,000 year old skeleton found on Liang Island in the Republic of China. {{cite web | url=https://discover.familytreedna.com/y-dna/O-CTS5726/ancient | title=Welcome to FamilyTreeDNA Discover }}

• Austro-Tai languages

{{columns-list|colwidth=22em|

• Genetic genealogy
• Haplogroup
• Haplotype
• Human Y-chromosome DNA haplogroup
• Molecular phylogenetics
• Paragroup
• Subclade
• Y-chromosome haplogroups in populations of the world
• Y-DNA haplogroups by ethnic group
• Y-DNA haplogroups in populations of East and Southeast Asia

}}

{{columns-list|colwidth=22em|

• O-47z
• O-M101
• O-M113
• O-M117
• O-M119
• O-M121
• O-M122
• O-M134
• O-M159
• O-M162
• O-M164
• O-M175
• O-M176
• O-M50
• O-M7
• O-M88
• O-M95
• O-MSY2.2
• O-P31

}}

{{clear}}

{{Y-DNA}}

{{reflist}}

{{refbegin|2}}

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Websites

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O-MSY2.2