Tar Heel/Coachman Formation
{{Infobox rockunit
| name = Tar Heel/Coachman Formation
| image =
| caption =
| type = Formation
| age = Early-Middle Campanian
~{{fossilrange|83.2|77.5}}
| prilithology =Siltstone, sandstone
| otherlithology =
| namedfor =Tar Heel, North Carolina
| namedby =
| region = {{Flag|North Carolina}}
{{Flag|South Carolina}}
| country = {{Flag|United States}}
| coordinates ={{coord|34.6|N|78.5|W|display=inline}}
| unitof = Black Creek Group
| subunits =
| underlies = Bladen Formation
| overlies =Middendorf Formation
| thickness =
| extent =
| area =
| map =
| map_caption =
|period=Campanian|paleocoordinates={{coord|35.0|N|50.2|W|display=inline}}}}
The Tar Heel Formation, also known as the Coachman Formation in South Carolina, is a Late Cretaceous (early to middle Campanian-aged) geologic formation in North Carolina and South Carolina, USA. It preserves fossils, including amber dating back to the Cretaceous period. A locality known as Phoebus Landing, has been dated to 78.5-77.1 Ma, and the formation has been overall dated to the early Campanian based on fossil pollen.{{Cite journal|url=https://www.sciencedirect.com/science/article/pii/S0195667115300471|doi = 10.1016/j.cretres.2015.08.004|title = A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography|year = 2016|last1 = Longrich|first1 = Nicholas R.|journal = Cretaceous Research|volume = 57|pages = 199–207| bibcode=2016CrRes..57..199L }}{{Cite web |title=Geolex — TarHeel publications |url=https://ngmdb.usgs.gov/Geolex/UnitRefs/TarHeelRefs_4067.html |access-date=2024-11-13 |website=ngmdb.usgs.gov}}
Likely deposited in a nearshore coastal environment representing a lower shoreface, it contains a high diversity of vertebrate remains. It has one of the most diverse dinosaur faunas known from the former landmass of Appalachia, the majority of which are known from two sites: Phoebus Landing along the Cape Fear River in Bladen County, North Carolina, as well as Stokes Quarry in Darlington County, South Carolina.{{Cite journal |last=Brownstein |first=Chase D. |date=2018-02-08 |title=The biogeography and ecology of the Cretaceous non-avian dinosaurs of Appalachia |url=https://palaeo-electronica.org/content/2018/2123-appalachia-biogeography |journal=Palaeontologia Electronica |language=en |volume=21 |issue=1 |pages=1–56 |doi=10.26879/801 |issn=1094-8074 |doi-access=free}}{{Cite web |title=PBDB Collection |url=https://paleobiodb.org/classic/basicCollectionSearch?collection_no=57918 |access-date=2024-11-13 |website=paleobiodb.org}}{{Cite journal |last1=Schwimmer |first1=David R. |last2=Sanders |first2=Albert E. |last3=Erickson |first3=Bruce R. |last4=Weems |first4=Robert E. |date=2015 |title=A Late Cretaceous Dinosaur and Reptile Assemblage from South Carolina, Usa |url=https://www.jstor.org/stable/24398224 |journal=Transactions of the American Philosophical Society |volume=105 |issue=2 |pages=i–157 |jstor=24398224 |issn=0065-9746}}
Fossil pollen grains suggest a subtropical to warm, moist temperate climate for the region, with an ecosystem largely dominated by flowering plants.{{Cite thesis|last=Mitra |first=Madhumi |date=2002-07-29 |title=Paleopalynology of the Tar Heel Formation of Atlantic Coastal Plain of North Carolina, United States. |url=https://repository.lib.ncsu.edu/items/00d1cf32-fa52-4e43-9fc0-a13c2886d444 |language=en}} The Tar Heel/Coachman Formation appears to be roughly concurrent with the Marshalltown Formation of New Jersey, which preserves a similar fauna.
Paleobiota
= Cartilaginous fish =
Based on the Paleobiology Database & Robb (1989):{{Cite book |last=Albert J Robb III |url=https://archive.org/details/robb-1989-phoebus-landing-fish |title=The Upper Cretaceous (Campanian, Black Creek Formation) fossil fish fauna of Phoebus Landing, Bladen County, North Carolina |date=1989}}
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colspan="7" align="center" |Cartilaginous fish of the Tar Heel/Coachman Formation |
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Genus
! Species ! Location ! Notes ! Images |
"Asteracanthus"
|A. sp. |Phoebus Landing, NC |A hybodontid shark. | |
Brachyrhizodus
|B. wichitaensis |Phoebus Landing, NC |An eagle ray. |
rowspan="2" |Carcharias
| rowspan="2" |Phoebus Landing, NC | rowspan="2" |A sand shark. | rowspan="2" |frameless |
C. samhammeri |
"Hypolophus"
|H. sp. |Phoebus Landing, NC | |
Meristodonoides
|M. montanensis |Phoebus Landing, NC |A hybodontid shark. |
Rhombodus
|R. laevis |Phoebus Landing, NC |A rhombodontid ray. |
Ischyrhiza
| I. mira |Phoebus Landing, NC | A sawskate. |
Ischyodus
|I. cf. bifurcatus |Phoebus Landing, NC |
Scapanorhynchus
|S. texanus |Phoebus Landing, NC |A goblin shark. |
rowspan="2" |
|S. kaupi | rowspan="2" |Phoebus Landing, NC Stokes Quarry, SC | rowspan="2" | An anacorid shark. Includes a coprolite potentially assignable to S. kaupi, containing the vertebrae of a baby turtle.{{Cite journal |last1=Schwimmer |first1=David R. |last2=Weems |first2=Robert E. |last3=Sanders |first3=Albert E. |date=2015 |title=A Late Cretaceous Shark Coprolite with Baby Freshwater Turtle Vertebrae Inclusions |url=https://www.jstor.org/stable/44708722 |journal=PALAIOS |volume=30 |issue=9/10 |pages=707–713 |doi=10.2110/palo.2015.019 |jstor=44708722 |bibcode=2015Palai..30..707S |issn=0883-1351}} | rowspan="2" |frameless |
S. pristodontus |
Squatina
|Phoebus Landing, NC |An angelshark. |
= Ray-finned fish =
Based on the Paleobiology Database, Robb (1989) & Stringer et al (2018). Some species are known only from otolith remains from Blue Banks & Auger Hole Landings, North Carolina, which are indicated:{{Cite web |last=Czaplewski |first=John J. |title=PBDB Navigator |url=https://paleobiodb.org/navigator/ |access-date=2024-10-10 |website=paleobiodb.org |language=en}}{{Cite journal |last1=Stringer |first1=Gary L. |last2=Clements |first2=Don |last3=Sadorf |first3=Eric |last4=Shannon |first4=Kevin |date=2019 |title=First Description and Significance of Cretaceous Teleostean Otoliths (Tar Heel Formation, Campanian) from North Carolina |url=https://www.eaglehill.us/epalonline/access-pages/001-Stringer-accesspage.shtml |journal=Eastern Paleontologist |volume=4 |pages=1–22 |issn=2475-5117}}
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Genus
! Species ! Location !Material ! Notes ! Images |
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rowspan="3" |Albula
|Blue Banks & Auger Hole Landings, NC |263-378 otoliths | rowspan="3" |A bonefish. | rowspan="3" |frameless |
?A. cf. A. ripleyensis
|Blue Banks Landing, NC |1 otolith |
A. sp.
|Phoebus Landing, NC | |
Anomoeodus
|A. phaseolus |Phoebus Landing, NC | |A pycnodontid pycnodont. |
Ariidae indet.
| |Blue Banks Landing, NC |11 otoliths |A sea catfish of uncertain affinities. | |
Aulopidae indet.
| |Blue Banks Landing, NC |1 otolith |A flagfin of uncertain affinities. | |
rowspan="2" |Beryx
| rowspan="2" |Blue Banks & Auger Hole Landings, NC |55 otoliths | rowspan="2" |A potential relative of alfonsinos. | rowspan="2" |frameless |
?B. zideki
|41 otoliths |
?Congridae indet.
| |Auger Hole Landing, NC |1 otolith |An apparent conger eel of uncertain affinities. | |
Cylindracanthus
|C. ornatus |Phoebus Landing, NC | |A fish of uncertain affinities. |
Enchodus
|E. cf. petrosus |Phoebus Landing, NC | |An enchodontid aulopiform. |
Gonostomatidae indet.
| |Blue Banks & Auger Hole Landings, NC |4 otoliths |A bristlemouth of uncertain affinities. | |
Hoplostethus
|Blue Banks & Auger Hole Landings, NC |4 otoliths |A slimehead. |
Kokenichthys
|K. ensis |Blue Banks Landing, NC |2 otoliths |An elopiform or osteoglossiform.{{Cite journal |last1=Stringer |first1=Gary L. |last2=Sloan |first2=James Carson |date=2023-05-25 |title=First Cretaceous teleostean otolith assemblage (Arkadelphia Formation, upper Maastrichtian) from Arkansas, USA, early Gadiformes, and the Western Interior Seaway |url=https://escholarship.org/uc/item/6zt71586 |journal=PaleoBios |volume=40 |issue=3 |doi=10.5070/P940361192 |issn=2373-8189 |doi-access=free}} | |
Lepisosteidae indet.
| |Phoebus Landing, NC | |A gar. | |
Megalopidae indet.
| |Blue Banks Landing, NC |1 otolith |A tarpon of uncertain affinities. | |
Osmeroides
|O. weileri |Blue Banks Landing, NC |6 otoliths |An osmeroidid elopomorph. |
Paralbula
|P. casei |Phoebus Landing, NC | |A phyllodontid elopomorph. | |
?Pempheris
|Auger Hole Landing, NC |1 otolith |An apparent sweeper. |
Percoidei indet.
| |Blue Banks & Auger Hole Landings, NC |5 otoliths |A percoid. | |
?Polymixia
|Blue Banks Landing, NC | |An apparent beardfish. |
Pterothrissus
|Blue Banks Landing, NC |67 otoliths |A pterothrissine bonefish. |
Saurodon
|S. sp. |Phoebus Landing, NC | |A saurodontid ichthyodectiform. |
Stephanodus
|S. sp. |Phoebus Landing, NC | |A pycnodont. | |
rowspan="2" |Xiphactinus
|X. audax | rowspan="2" |Phoebus Landing, North Carolina, NC | | rowspan="2" |An ichthyodectid ichthyodectiform. | rowspan="2" |frameless |
X. vetus
| |
= Reptiles =
== Dinosaurs ==
=== Ornithischians ===
Based mainly on Brownstein (2018):
An indeterminate hadrosauroid is known from Stokes Quarry, SC.
=== Theropods ===
Indeterminate theropods, ornithomimosaurs, and maniraptorans are known from Stokes Quarry.
== Crocodylomorphs ==
Based on the Paleobiology Database:
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Genus
! Species ! Location ! Notes ! Images |
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Borealosuchus
|B. formidabilis |Phoebus Landing, NC |A eusuchian. |
Bottosaurus
|B. sp. |Stokes Quarry, SC |An early caiman. | |
Deinosuchus
|D. rugosus (=Thecachampsa rugosa, Polydectes biturgidus, Polyptychodon rugosus) |Phoebus Landing & Clifton Farm, NC Stokes Quarry, SC |An alligatoroid, one of the largest known crocodilians. Type locality for genus and species. |
Gavialoidea indet.
| |Stokes Quarry, SC |A gavialoid. | |
== Turtles ==
Based on the Paleobiology Database:
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Genus
! Species ! Location ! Notes ! Images |
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Adocus
|A. beatus |Phoebus Landing, NC Stokes Quarry, SC |An adocid. |
Bothremys
|B. cooki |Phoebus Landing, NC |A bothremydid side-necked turtle. | |
rowspan="2" |Chedighaii
|C. barberi | rowspan="2" |Phoebus Landing, NC | rowspan="2" |A bothremydid side-necked turtle. | rowspan="2" | |
C. hutchinsoni |
Corsochelys
|C. bentlyi |Stokes Quarry, SC |A sea turtle, likely a dermochelyid. Type locality for species. | |
Euclastes
|E. wielandi |Stokes Quarry, SC |A pancheloniid sea turtle. |
Osteopygis
|O. emarginatus |Phoebus Landing, NC |A macrobaenid. |File:The American journal of science (1904) (18152226245).jpg |
?Taphrosphys
|?T. dares |Phoebus Landing, NC |A bothremydid side-necked turtle, taxonomic placement disputed. | |
Toxochelys
|T. sp. |Stokes Quarry, SC |A toxochelyid sea turtle. |
rowspan="2" |"Trionyx"
|Phoebus Landing, NC | rowspan="2" |A stem-softshell turtle, likely not an actual member of the genus Trionyx. Species names are nomina dubia. | rowspan="2" | |
"T." priscus
|Stokes Quarry, SC |
== Plesiosaurs ==
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Genus
! Species ! Location ! Notes ! Images |
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Elasmosauridae indet.
| |Stokes Quarry, South Carolina |An elasmosaurid. | |
== Squamates ==
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Genus
! Species ! Location ! Notes ! Images |
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Halisaurus
|H. sp. |Phoebus Landing, NC |A halisaurine mosasaur. |
Platecarpus
|P. sp. |Phoebus Landing, NC |A plioplatecarpine mosasaur. |
Prognathodon
|P. sp. |Phoebus Landing, NC |A mosasaurine mosasaur. |
Teiidae indet.
| |Stokes Quarry, SC | |
Tylosaurus
|T. sp. |Phoebus Landing, NC Stokes Quarry, SC |A tylosaurine mosasaur. |
= Mammals =
Based on the Paleobiology Database:{{Cite web |title=PBDB Collection |url=https://paleobiodb.org/classic/basicCollectionSearch?collection_no=151919 |access-date=2024-11-15 |website=paleobiodb.org}}
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Genus
! Species ! Location ! Notes ! Images |
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Marsupialia indet.
| |Darlington County, SC |A presumed marsupial relative. | |
Multituberculata indet.
| |Darlington County, SC |A multituberculate. | |