haliotis

{{Short description|Genus of gastropods}}

{{use dmy dates|date=July 2024}}

{{for|the human use of Haliotis molluscs|Abalone}}

{{distinguish|Halitosis}}

{{Merge to|Abalone|date=March 2025}}

{{Automatic taxobox

| fossil_range = {{Fossil range | Late Cretaceous | Recent | ref={{harvnb|Geiger|Groves|1999|p=872}} }}

| image = LivingAbalone.JPG

| image_caption = Living abalone in tank showing epipodium and tentacles, anterior end to the right.

| display_parents = 3

| parent_authority = Rafinesque, 1815

| taxon = Haliotis

| authority = Linnaeus, 1758

| type_species = Haliotis asinina

| type_species_authority = Linnaeus, 1758

| synonyms_ref = {{harvnb|Gofas|Tran|Bouchet|2014}}

| synonyms =

  • Euhaliotis Wenz, 1938
  • Eurotis Habe & Kosuge, 1964
  • Exohaliotis Cotton & Godfrey, 1933
  • Haliotis (Haliotis) Linnaeus, 1758
  • Haliotis (Marinauris) Iredale, 1937
  • Haliotis (Nordotis) Habe & Kosuge, 1964
  • Haliotis (Notohaliotis) Cotton & Godfrey, 1933
  • Haliotis (Padollus) Montfort, 1810
  • Haliotis (Paua) C. Fleming, 1953
  • Haliotis (Sulculus) H. Adams & A. Adams, 1854
  • Marinauris Iredale, 1927
  • Neohaliotis Cotton & Godfrey, 1933
  • Nordotis Habe & Kosuge, 1964
  • Notohaliotis Cotton & Godfrey, 1933
  • Ovinotis Cotton, 1943
  • Padollus Montfort, 1810
  • Paua C. Fleming, 1953
  • Sanhaliotis Iredale, 1929
  • Schismotis Gray, 1856
  • Teinotis H. Adams & A. Adams, 1854
  • Tinotis P. Fischer, 1885 (invalid: unjustified emendation of Teinotis)
  • Usahaliotis Habe & Kosuge, 1964

}}

Haliotis, common name abalone, is the only genus in the family Haliotidae.

This genus once contained six subgenera. These subgenera have become alternate representations of Haliotis.{{harvnb|Gofas|Tran|Bouchet|2014}} The genus consists of small to very large, edible, herbivorous sea snails, marine gastropod molluscs. The number of species recognized worldwide ranges between 30{{harvnb|Dauphin|Cuif|Mutvei|Denis|1989|p=9}} and 130,{{harvnb|Cox|1962|p=8}} with over 230 species-level taxa described. The most comprehensive treatment of the family considers 56 species valid, with 18 additional subspecies.{{Cite journal|last=D.L.|first=Geiger|date=1999|title=Distribution and biogeography of the recent Haliotidae (Gastropoda: Vetigastropoda) world-wide|url=http://www.marinespecies.org/imis.php?module=ref&refid=999&basketaction=add|journal=Bollettino Malacologico|language=en}}

Other common names are ear shells, sea ears, and, rarely, muttonfish or muttonshells in parts of Australia, ormer in the UK, perlemoen in South Africa, and the Māori name for three species in New Zealand is pāua.{{harvnb|Beesley|Ross|Wells|1998}}{{page needed|date=August 2014}}

Description

Image:AbaloneInside.jpg inside surface of a red abalone shell from Northern California: The coin is about one inch in diameter.]]

The shells of abalone have a low, open, spiral structure, and are characterized by having several open respiratory pores in a row near the shell's outer edge. The thick inner layer of the shell is composed of nacre, which in many species of abalone is highly iridescent, giving rise to a range of strong, changeable colors, which make the shells attractive to humans as decorative objects, in jewelry, and as a source of colorful mother-of-pearl.

The shell of abalone is convex, rounded to oval shape, and may be highly arched or very flattened. The shell of the majority of species is ear-shaped, presenting a small, flat spire and two to three whorls. The last whorl, known as the body whorl, is auriform, meaning that the shell resembles an ear, giving rise to the common name "ear shell". Haliotis asinina has a somewhat different shape, as it is more elongated and distended. The shell of Haliotis cracherodii cracherodii is also unusual as it has an ovate form, is imperforate, shows an exserted spire, and has prickly ribs.

A mantle cleft in the shell impresses a groove in the shell, in which are the row of holes (known as tremata), characteristic of the genus. These holes are respiratory apertures for venting water from the gills and for releasing sperm and eggs into the water column. They make up what is known as the selenizone which forms as the shell grows. This series of eight to 38 holes is near the anterior margin. Only a small number are generally open. The older holes are gradually sealed up as the shell grows and new holes form. Therefore, the number of tremata is not characteristic for the species. Each species has a number of open holes, between four and 10, in the selenizone. This number is not fixed and can vary within a species and between populations. Abalone have no operculum. The aperture of the shell is very wide and nacreous.

The exterior of the shell is striated and dull. The color of the shell is very variable from species to species, which may reflect the animal's diet. The iridescent nacre that lines the inside of the shell varies in color from silvery white, to pink, red and green-red, to deep blue, green to purple.

The animal shows fimbriated head-lobes. The side-lobes are also fimbriated and cirrated. The rounded foot is very large. The radula has small median teeth, and the lateral teeth are single and beam-like. About 70 uncini are present, with denticulated hooks, the first four very large. The soft body is coiled around the columellar muscle, and its insertion, instead of being on the columella, is on the middle of the inner wall of the shell. The gills are symmetrical and both well developed.{{harvnb|Tryon|1880|p=41}}

These snails cling solidly with their broad muscular foot to rocky surfaces at sublittoral depths, although some species such as Haliotis cracherodii used to be common in the intertidal zone. Abalone reach maturity at a relatively small size. Their fecundity is high and increases with their size (from 10,000 to 11 million eggs at a time). The spermatozoa are filiform and pointed at one end, and the anterior end is a rounded head.{{harvnb|Tryon|1880|p=46}}

The larvae are lecithotrophic. The adults are herbivorous and feed with their rhipidoglossan radula on macroalgae, preferring red or brown algae. Sizes vary from {{cvt|20|mm|in|frac=32}} (Haliotis pulcherrima) to {{cvt|200|mm|in|frac=8}}, while Haliotis rufescens is the largest of the genus at {{cvt|12|in|cm|order=flip}}.{{harvnb|Hoiberg|1993|p=7}}

By weight, about one-third of the animal is edible meat, one-third is offal, and one-third is shell.{{citation needed|date=August 2014}}

Structure and properties of the shell

The shell of the abalone is exceptionally strong and is made of microscopic calcium carbonate tiles stacked like bricks. Between the layers of tiles is a clingy protein substance. When the abalone shell is struck, the tiles slide instead of shattering and the protein stretches to absorb the energy of the blow. Material scientists around the world are studying this tiled structure for insight into stronger ceramic products such as body armor.{{harvnb|Lin|Meyers|2005|p=27 & 38}} The dust created by grinding and cutting abalone shell is dangerous; appropriate safeguards must be taken to protect people from inhaling these particles. There is much discussion of this topic online.{{Citation needed|date=January 2019}}Abalone Toxicity https://orchid.ganoksin.com/t/abalone-toxicity/9357/2Shell Dust Dangers https://www.banjohangout.org/archive/187285

Species

The number of species that are recognized within the genus Haliotis has fluctuated over time, and depends on the source that is consulted. The number of recognized species ranges from 30 to 130. This list finds a compromise using the "WoRMS" database, plus some species that have been added, for a total of 57.{{harvnb|Gofas|Tran|Bouchet|2014}}{{harvnb|Abbott|Dance|2000}} The majority of abalone have not been evaluated for conservation status. Those that have been reviewed tend to show that the abalone in general is declining in numbers, and will need protection throughout the globe.

= Extant species =

class="wikitable"

|+ Species of abalone

! Species !! Range !! Conservation status

Haliotis alfredensis Bartsch, 1915{{refn|group=nb|This species, depending on the source is its own species{{harvnb|Tran|Bouchet|2009}} or is a synonym of Haliotis speciosa.{{harvnb|EoL|2014}}}}South Africa| {{IUCN status|DD|78748407|1}}
Haliotis arabiensis Owen, Regter & Van Laethem, 2016Off Yemen and Oman| {{IUCN status|NT|157247863|1}}
Haliotis asinina Linnaeus, 1758Philippines; Indonesia; Australia; Japan; Thailand; Vietnam| {{IUCN status|LC|78749198|1}}
Haliotis australis Gmelin, 1791New Zealand| {{IUCN status|LC|78763353|1}}
Haliotis brazieri Angas, 1869Eastern Australia| {{IUCN status|NT|78763607|1}}
Haliotis clathrata Reeve, 1846Seychelles; Comores; Madagascar; Mauritius; Kenya| {{IUCN status|LC|78763614|1}}
Haliotis coccoradiata Reeve, 1846Eastern Australia| {{IUCN status|LC|78763643|1}}
Haliotis corrugata Wood, 1828California, USA; Baja California, Mexico| {{IUCN status|CR|78763727|1}}, Species of Concern National Marine Fisheries Service;{{harvnb|Neuman|2007}} Vulnerable (global) and imperiled (California) California Department of Fish and Wildlife{{harvnb|State of California|2011}}
Haliotis cracherodii Leach, 1814California, USA; Baja California, Mexico| {{IUCN status|CR|41880|1}}, Vulnerable (Global, Nation: US, State: California) California Department of Fish and Wildlife;{{harvnb|Anon|2014f}} Listed endangered National Marine Fisheries Service{{harvnb|Anon|2009}}
Haliotis cyclobates Péron & Lesueur, 1816Southern Australia| {{IUCN status|LC|78763977|1}}
Haliotis dalli Henderson, 1915Galapagos Islands, western Colombia| {{IUCN status|DD|78764148|1}}
Haliotis discus Reeve, 1846Japan; South Korea| {{IUCN status|EN|78764186|1}}
Haliotis dissona (Iredale, 1929)Australia; New Caledonia| {{IUCN status|LC|78764274|1}}
Haliotis diversicolor Reeve, 1846Japan; Australia; Southeast Asia| {{IUCN status|DD|78764384|1}}
Haliotis drogini Owen & Reitz, 2012Cocos Island| {{IUCN status|VU|78764677|1}}
Haliotis elegans Koch & Philippi, 1844Western Australia| {{IUCN status|LC|78764711|1}}
Haliotis exigua Dunker, R.W., 1877 (synonym of H. diversicolor)Japan| Not evaluated
Haliotis fatui Geiger, 1999Tonga Mariana Islands| {{IUCN status|DD|78764797|1}}
Haliotis fulgens Philippi, 1845California, USA; Baja California, Mexico| {{IUCN status|CR|78768961|1}}, Vulnerable (Global, State: California California Department of Fish and Wildlife); Species of Concern NMFS{{harvnb|Neuman|2009}}
Haliotis geigeri Owen, 2014São Tomé and Príncipe Islands| {{IUCN status|VU|78768971|1}}
Haliotis gigantea Gmelin, 1791Japan| {{IUCN status|EN|78768975|1}}
Haliotis glabra Gmelin, 1791Philippines; Vietnam| {{IUCN status|LC|78768980|1}}
Haliotis iris Gmelin, 1791New Zealand| {{IUCN status|LC|78769001|1}}
Haliotis jacnensis Reeve, 1846Japan; Nicobar Islands; Ryukyu Islands; Pacific Islands;| {{IUCN status|LC|78769011|1}}
Haliotis kamtschatkana Jonas, 1845Western North America| {{IUCN status|EN|61743|1}}, Imperiled (Alaska, British Columbia), Vulnerable (global, US), critically imperiled (California);{{harvnb|Anon|2014c}} Species of Concern NMFS{{harvnb|Gustafson|Rumsey|2007}}
Haliotis laevigata Donovan, 1808South Australia; Tasmania| {{IUCN status|VU|78769674|1}}
Haliotis madaka (Habe, 1977)Japan; South Korea| {{IUCN status|EN|78769692|1}}
Haliotis mariae Wood, 1828Oman; Yemen| {{IUCN status|EN|78769871|1}}
Haliotis marmorata Linnaeus, 1758Liberia; Ivory Coast; Ghana| {{IUCN status|LC|78769875|1}}
Haliotis melculus (Iredale, 1927)Australia (New South Wales, Queensland)| {{IUCN status|VU|78771080|1}}
Haliotis midae Linnaeus, 1758South Africa| {{IUCN status|EN|78771094|1}}
Haliotis mykonosensis Owen, Hanavan & Hall, 2001Greece; Turkey; Tunisia| {{IUCN status|LC|78771101|1}}
Haliotis ovina Gmelin, 1791Thailand; Vietnam; southern part of the Pacific Ocean; Andaman Islands; Maldives; Ryukyu Islands| {{IUCN status|LC|78771163|1}}
Haliotis papulata Reeve, 1846

|Australia; Papua New Guinea; Philippines; Sri Lanka; Thailand

|{{IUCN status|LC|161334052|1}}

Haliotis parva Linnaeus, 1758South Africa; Angola| {{IUCN status|DD|78771325|1}}
Haliotis pirimoana Walton, Marshall, Rawlence & Spencer, 2024Manawatāwhi / Three Kings Islands, New Zealand{{cite journal |doi=10.1080/13235818.2024.2390476 |title=Haliotis virginea Gmelin, 1791 and a new abalone from Aotearoa New Zealand (Mollusca: Gastropoda: Haliotidae) |year=2024 |last1=Walton|first1=Kerry |last2=Marshall |first2=Bruce A. |last3=Rawlence|first3=Nicolas J. |last4=Spencer|first4=Hamish G. |journal=Molluscan Research |volume= 44|issue= 4|pages= 305–315|doi-access=free |bibcode=2024MollR..44..305W }}{{Cite news|url=https://theconversation.com/the-paua-that-clings-to-the-sea-a-new-species-of-abalone-found-only-in-waters-off-a-remote-nz-island-chain-236568 |title='The pāua that clings to the sea': a new species of abalone found only in waters off a remote NZ island chain |last1=Walton|first1=Kerry |last2=Spencer|first2=Hamish |last3=Rawlence|first3=Nic |date=3 September 2024 |work=The Conversation|access-date=17 September 2024}}| Not evaluated
Haliotis planata G. B. Sowerby II, 1882Ryukyu Islands; Sri Lanka; Indonesia; Fiji; Andaman Sea| {{IUCN status|LC|78771329|1}}
Haliotis pourtalesii Dall, 1881Eastern USA; Gulf of Mexico; Eastern South America; northern Colombia| {{IUCN status|DD|78771388|1}}
Haliotis pulcherrima Gmelin, 1791Polynesia| {{IUCN status|DD|78771440|1}}
Haliotis queketti E.A. Smith, 1910Eastern Africa| {{IUCN status|DD|78771516|1}}
Haliotis roei Gray, 1826Australia| {{IUCN status|NT|78771528|1}}
Haliotis rubiginosa Reeve, 1846Lord Howe Island| {{IUCN status|CR|78771575|1}}
Haliotis rubra Leach, 1814Southern and Eastern Australia| {{IUCN status|VU|78771579|1}}
Haliotis rufescens Swainson, 1822Western North America| {{IUCN status|CR|78771583|1}}, apparently secure (global, US); critically imperiled (Canada){{harvnb|Anon|2014d}}
Haliotis rugosa Lamarck, 1822South Africa; Madagascar; Mauritius; Red Sea| {{IUCN status|LC|78771608|1}}
Haliotis scalaris (Leach, 1814)Southern and Western Australia| {{IUCN status|LC|78771632|1}}
Haliotis semiplicata Menke, 1843Western Australia| {{IUCN status|LC|78771692|1}}
Haliotis sorenseni Bartsch, 1940California, USA; Baja California, Mexico| {{IUCN status|CR|78771696|1}}, critically imperiled (global, US, California);{{harvnb|Anon|2014e}} Endangered NMFS{{harvnb|Anon|2001}}
Haliotis spadicea Donovan, 1808South Africa| {{IUCN status|LC|78771700|1}}
Haliotis speciosa Reeve, 1846 (synonym of H. tuberculata)

| Eastern South Africa

| Not evaluated
Haliotis squamosa Gray, 1826Southern Madagascar| {{IUCN status|DD|78771710|1}}
Haliotis stomatiaeformis Reeve, 1846Malta; Sicily| {{IUCN status|VU|78772043|1}}
Haliotis supertexta Lischke, 1870 (synonym of H. diversicolor)Japan; Sao Tome| Not evaluated
Haliotis thailandis Dekker & Patamakanthin, 2001 (synonym of H. papulata)Andaman Sea| Not evaluated
Haliotis tuberculata Linnaeus, 1758Ireland (introduced); Channel Islands; Azores; Canary Islands; Madeira ; Brittany; Great Britain| {{IUCN status|VU|78772221|1}}
Haliotis unilateralis Lamarck, 1822Gulf of Aqaba; East Africa; Seychelles;| {{IUCN status|LC|78772250|1}}
Haliotis varia Linnaeus, 1758Mascarene Basin; Red Sea; Sri Lanka; Western Pacific;| {{IUCN status|LC|78772274|1}}
Haliotis virginea Gmelin, 1791New Zealand; Chatham Islands; Auckland Islands; Campbell Island| {{IUCN status|LC|78772279|1}}
Haliotis walallensis Stearns, 1899Western North America{{IUCN status|CR|78772302|1}}

File:Ass’s ear abalone (Haliotis asinina) S01.jpg|A dorsal view of a live ass's ear abalone, Haliotis asinina

File:Pinkabalone 300.jpg|the pink abalone, Haliotis corrugata

File:Haliotis cracherodii.JPG|The black abalone, Haliotis cracherodii

File:Blacklip abalone.jpg|Dorsal (left) and ventral (right) views of the blacklip abalone, Haliotis rubra

File:Whiteabalone 300.jpg|The white abalone, Haliotis sorenseni

File:Haliotis varia f. dohrniana 001.jpg|A shell of Haliotis varia form dohrniana

{{clear}}

Image:Haliotis asinina 01.JPG|Haliotis asinina

Image:Haliotis australis 001.jpg|Haliotis australis

Image:Haliotis clathrata 01.JPG|Haliotis clathrata

Image:Haliotis coccoradiata.jpg|Haliotis coccoradiata

Image:Haliotis corrugata.jpg|Haliotis corrugata

Image:Haliotis cracherodii.jpg|Haliotis cracherodii

Image:Haliotis cyclobates.jpg|Haliotis cyclobates

Image:Haliotis discus discus 01.JPG|Haliotis discus discus

Image:Haliotis diversicolor 01.jpg|Haliotis diversicolor

Image:Haliotis elegans 001.jpg|Haliotis elegans

Image:Haliotis fatui 001.jpg|Haliotis fatui

Image:Haliotis fulgens fulgens 01.JPG|Haliotis fulgens fulgens

Image:Haliotis gigantea f. sieboldii 01.JPG|Haliotis gigantea f. sieboldii

Image:Haliotis glabra 01.JPG|Haliotis glabra

Image:Haliotis iris 01.JPG|Haliotis iris

Image:Haliotis jacnensis 002.jpg|Haliotis jacnensis

Image:Haliotis kamtschatkana assimilis 01.JPG|Haliotis kamtschatkana assimilis (South California).

Image:Haliotis laevigata 01.JPG|Haliotis laevigata (South Australia).

Image:Haliotis madaka 001.jpg|Haliotis madaka

Image:Haliotis mariae 002.jpg|Haliotis mariae

Image:Haliotis marmorata 002.jpg|Haliotis marmorata

Image:Haliotis midae 01.jpg|Haliotis midae

Image:Haliotis ovina ovina 01.JPG|Haliotis ovina

Image:Haliotis parva 001.jpg|Haliotis parva

Image:Haliotis planata 01.JPG|Haliotis planata

Image:Haliotis pulcherrima 001.jpg|Haliotis pulcherrima

Image:Haliotis queketti 001.jpg|Haliotis queketti

Image:Haliotis roei 002.jpg|Haliotis roei

Image:Haliotis rubra P2164176.JPG|Haliotis rubra

Image:Haliotis rufescens 22a.jpg|Haliotis rufescens

Image:Haliotis rugosa 2.jpg|Haliotis rugosa

Image:Haliotis scalaris (staircase abalone) (Quinn's Rocks, Western Australia) 1 (23565526643).jpg|Haliotis scalaris

Image:Haliotis semùiplicata 001.jpg|Haliotis semiplicata

Image:Haliotis sorenseni 001.jpg|Haliotis sorenseni

Image:Haliotis spadicea (blood-spotted abalone) (South Africa) 3 (24139744761).jpg|Haliotis spadicea

Image:Haliotis squamosa 001.jpg|Haliotis squamosa

Image:Haliotis stomatiaeformis 002.jpg|Haliotis stomatiaeformis

Image:Haliotis supertexta 001.jpg|Haliotis supertexta

Image:Haliotis thailandis 002.jpg|Haliotis thailandis

Image:Haliotis tuberculata tuberculata 01.JPG|Haliotis tuberculata (Europe)

Image:Haliotis unilateralis 002.jpg|Haliotis unilateralis

Image:Haliotis varia 01.JPG|Haliotis varia

Image:Haliotis virginea shell 2.jpg|Haliotis virginea

Image:Haliotis walallensis.jpg|Haliotis walallensis

= Fossil species =

Conservation

Over half of the modern Haliotis species with sufficient data are considered threatened to some extent on the IUCN Red List, with all but one species from the Pacific coast of North America being critically endangered as a consequence of massive historical overharvesting, withering abalone syndrome, and recent marine heatwaves which have caused collapses of both abalone and their habitat. Haliotis species from elsewhere are also threatened by overexploitation and climate change. In addition, abalone as a whole are considered highly vulnerable to ocean acidification due to their accretion of aragonite and dependence on susceptible coralline algae for development, and thus may eventually go extinct unless the rate of ocean acidification is arrested.{{cite journal|last1=Peters|first1=H.|last2=Rogers-Bennett|first2=L.|last3=De Shields|first3=R. M.|date=2021-05-21|url=https://www.iucnredlist.org/species/78771583/78772573|title=Haliotis rufescens|website=The IUCN Red List of Threatened Species |id=2021: e.T78771583A78772573|publisher=IUCN|doi=10.2305/IUCN.UK.2021-3.RLTS.T78771583A78772573.en|access-date=2022-12-24|s2cid=245151062|doi-access=free}}{{Cite web |last=Kerlin |first=Katherine E. |date=2022-12-12 |title=All West Coast Abalones at Risk of Extinction on the IUCN Red List |url=https://www.ucdavis.edu/climate/news/all-west-coast-abalone-added-endangered-iucns-red-list |access-date=2022-12-24 |website=UC Davis |language=en}}{{Cite web |title=First-ever global Red List assessment of abalone underlines urgency of combatting illegal trade - Wildlife Trade News from TRAFFIC |url=https://www.traffic.org/news/first-ever-global-red-list-assessment-of-abalone-underlines-urgency-of-combatting-illegal-trade/ |access-date=2022-12-24 |website=www.traffic.org |language=en}}{{Cite journal |last1=Byrne |first1=Maria |last2=Ho |first2=Melanie |last3=Wong |first3=Eunice |last4=Soars |first4=Natalie A. |last5=Selvakumaraswamy |first5=Paulina |last6=Shepard-Brennand |first6=Hannah |last7=Dworjanyn |first7=Symon A. |last8=Davis |first8=Andrew R. |date=2010-12-22 |title=Unshelled abalone and corrupted urchins: development of marine calcifiers in a changing ocean |url=http://dx.doi.org/10.1098/rspb.2010.2404 |journal=Proceedings of the Royal Society B: Biological Sciences |volume=278 |issue=1716 |pages=2376–2383 |doi=10.1098/rspb.2010.2404 |pmid=21177689 |pmc=3119014 |issn=0962-8452}}

Synonyms

{{Main|List of abalone synonyms}}

See also

References

{{Reflist|2}}

Notes

{{Reflist|group=nb}}

Sources

  • {{cite book | last1 = Abbott | first1 = R. Tucker | last2 = Dance | first2 = S. Peter | title = Compendium of Seashells | url = https://archive.org/details/Compendium_of_Seashells_by_S._Peter_Dance | year = 2000 | publisher = Odyssey Publishing | isbn = 978-0-9661720-0-3 | pages = [https://archive.org/details/Compendium_of_Seashells_by_S._Peter_Dance/page/n29 19]–23 | edition = 4th }}
  • {{cite web | author = Anon | year = 2014c | url = http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=113951&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | access-date = 22 August 2014 | title = Haliotis kamtschatkana – Jonas, 1845 | website = NatureServe Explorer | archive-url = https://web.archive.org/web/20140826120301/http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=113951&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | archive-date = 26 August 2014 | url-status = dead }}
  • {{cite web | author = Anon | year = 2014d | url = http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=107385&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | access-date = 22 August 2014 | title = Haliotis refescens – Swainson, 1822 | website = NatureServe Explorer | archive-url = https://web.archive.org/web/20140826160903/http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=107385&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | archive-date = 26 August 2014 | url-status = dead }}
  • {{cite web | author = Anon | year = 2014e | url = http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=116467&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | access-date = 22 August 2014 | title = Haliotis sorenseni – Bartsch, 1940 | website = NatureServe Explorer | archive-url = https://web.archive.org/web/20140826115417/http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=116467&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | archive-date = 26 August 2014 | url-status = dead }}
  • {{cite web | author = Anon | year = 2014f | url = http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=120086&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | access-date = 22 August 2014 | title = Haliotis cracherodii – Leach, 1814 | website = NatureServe Explorer | archive-url = https://web.archive.org/web/20140826120205/http://explorer.natureserve.org/servlet/NatureServe?sourceTemplate=tabular_report.wmt&loadTemplate=species_RptComprehensive.wmt&selectedReport=RptComprehensive.wmt&summaryView=tabular_report.wmt&elKey=120086&paging=home&save=true&startIndex=1&nextStartIndex=1&reset=false&offPageSelectedElKey=814921&offPageSelectedElType=species&offPageYesNo=true&post_processes=&radiobutton=radiobutton&selectedIndexes=814921&selectedIndexes=120086&selectedIndexes=814920&selectedIndexes=113951&selectedIndexes=107385&selectedIndexes=116467 | archive-date = 26 August 2014 | url-status = dead }}
  • {{cite web |author=Anon |date=14 January 2009 |website=Federal Register: The Daily Journal of the United States |title=Endangered and Threatened Species; Endangered Status for Black Abalone |url=https://www.federalregister.gov/articles/2009/01/14/E9-635/endangered-and-threatened-wildlife-and-plants-endangered-status-for-black-abalone |access-date=22 August 2014 |archive-date=26 August 2014 |url-status=live |archive-url=https://web.archive.org/web/20140826114139/https://www.federalregister.gov/articles/2009/01/14/E9-635/endangered-and-threatened-wildlife-and-plants-endangered-status-for-black-abalone }}
  • {{cite web |author=Anon |date=29 May 2001 |website=Federal Register: The Daily Journal of the United States |title=Endangered and Threatened Species; Endangered Status for White Abalone |url=https://www.federalregister.gov/articles/2001/05/29/01-13430/endangered-and-threatened-species-endangered-status-for-white-abalone |access-date=22 August 2014 |archive-date=29 July 2014 |url-status=live |archive-url=https://web.archive.org/web/20140729124601/https://www.federalregister.gov/articles/2001/05/29/01-13430/endangered-and-threatened-species-endangered-status-for-white-abalone }}
  • {{cite book | last1 = Beesley | first1 = P. L. | last2 = Ross | first2 = G. J. B. | last3 = Wells | first3 = A. | title = Mollusca: The Southern Synthesis: An Essential Reference | year = 1998 | publisher = CSIRO Publishing | isbn = 978-0-643-05756-2 | pages = 667–669 | location = Melbourne, Australia }}
  • {{cite journal | last = Cox | first = Keith W. | year = 1962 | title = California abalone, family Haliotidae | journal = The Resources Agency of California Department of Fish and Game: Fish Bulletin | volume = 118 | issn = 6306-2593 }}
  • {{cite journal | last1 = Dauphin | first1 = Y. | last2 = Cuif | first2 = J. P. | last3 = Mutvei | first3 = H. | last4 = Denis | first4 = A. | year = 1989 | title = Mineralogy, Chemistry and Ultrastructure of the External Shell-layer in Ten Species of Haliotis With Reference to Haliotis tuberculata (Mollusca, Archaeogastropoda) | journal = Bulletin of the Geological Institutions of the University of Uppsala | volume = 15 | pages = 7–38 | issn = 0302-2749 }}
  • {{cite web |author=EoL |year=2014 |url=http://eol.org/pages/3048993/overview |website=Encyclopedia of Life |access-date=21 August 2014 |archive-date=21 August 2014 |archive-url=https://web.archive.org/web/20140821215843/http://eol.org/pages/3048993/overview |title=Haliotis speciosa: Splendid Abalone |url-status=live }}
  • {{cite journal | last1 = Geiger | first1 = Daniel L. | last2 = Groves | first2 = Lindsey T. | title = Review of Fossil Abalone (Gastropoda, Vetigastropoda, Haliotidae) with Comparison to Recent Species | journal = Journal of Paleontology | volume = 73 | issue = 5 | pages = 872–885 | doi = 10.1017/S0022336000040713| date = September 1999 | bibcode = 1999JPal...73..872G | s2cid = 87537607 | issn = 0022-3360 }}
  • {{cite web |last1=Gofas |first1=Serge |last2=Tran |first2=Bastien |last3=Bouchet |first3=Phillippe |year=2014 |title=WoRms Taxon Details: Haliotis Linnaeus, 1758 |website=WoRMS (World Register of Marine Species) |access-date=16 August 2014 |archive-date=10 October 2014 |url=http://www.marinespecies.org/aphia.php?p=taxdetails&id=138050 |archive-url=https://web.archive.org/web/20141010200912/http://www.marinespecies.org/aphia.php?p=taxdetails&id=138050 |url-status=live }}
  • {{cite web |last1=Gustafson |first1=Rick |last2=Rumsey |first2=Scott |date=1 November 2007 |access-date=22 August 2014 |archive-date=7 October 2012 |url=http://www.nmfs.noaa.gov/pr/pdfs/species/pintoabalone_detailed.pdf |archive-url=https://web.archive.org/web/20121007180626/http://www.nmfs.noaa.gov/pr/pdfs/species/pintoabalone_detailed.pdf |title=Species of Concern: Pinto Abalone Haliotis Kamtschatkana |website=NOAA: National Marine Fisheries Service |url-status=live }}
  • {{cite encyclopedia | editor-last = Hoiberg | editor-first = Dale H. | encyclopedia = Encyclopædia Britannica| edition = 15th | year = 1993 | publisher = Encyclopædia Britannica, Inc. | volume = 1: A-ak Bayes | location = Chicago, IL | isbn = 978-0-85229-961-6 | lccn = 2002113989 | title = The New Encyclopaedia Britannica }}
  • {{cite journal | last1 = Lin | first1 = Albert | last2 = Meyers | first2 = Marc André | doi = 10.1016/j.msea.2004.06.072 | title = Growth and Structure in Abalone Shell | date = 15 January 2005 | journal = Materials Science and Engineering: A | volume = 390 | issue = 1–2 | issn = 0921-5093 | pages = 27–41 }}
  • {{cite web |last=Neuman |first=Melissa |date=10 June 2009 |access-date=22 August 2014 |archive-date=4 March 2016 |url=http://www.nmfs.noaa.gov/pr/pdfs/species/greenabalone_detailed.pdf |archive-url=https://web.archive.org/web/20160304111523/http://www.nmfs.noaa.gov/pr/pdfs/species/greenabalone_detailed.pdf |title=Species of Concern: Green Abalone Haliotis fulgens |website=NOAA: National Marine Fisheries Service |url-status=live }}
  • {{cite web |last=Neuman |first=Melissa |date=2 November 2007 |access-date=22 August 2014 |archive-date=4 March 2016 |url=http://www.nmfs.noaa.gov/pr/pdfs/species/pinkabalone_detailed.pdf |archive-url=https://web.archive.org/web/20160304050111/http://www.nmfs.noaa.gov/pr/pdfs/species/pinkabalone_detailed.pdf |title=Species of Concern: Pink Abalone Haliotis corrugata |website=NOAA: National Marine Fisheries Service |url-status=live }}
  • {{cite web |author=State of California |date=January 2011 |url=http://www.dfg.ca.gov/biogeodata/cnddb/pdfs/spanimals.pdf |title=Special Animals (898 Taxa) |website=State of California: Division of Fish and Game |access-date=22 August 2014 |archive-date=24 January 2014 |archive-url=https://web.archive.org/web/20140124164328/http://www.dfg.ca.gov/biogeodata/cnddb/pdfs/spanimals.pdf |url-status=dead }}
  • {{cite web |last1=Tran |first1=Bastien |last2=Bouchet |first2=Phillippe |year=2009 |url=http://www.marinespecies.org/aphia.php?p=taxdetails&id=445309 |website=WoRMS: World Register of Marine Species |title=WoRMS Taxon Details: Haliotis corrugata oweni Talmadge, 1966 |access-date=19 August 2014 |archive-date=17 October 2014 |archive-url=https://web.archive.org/web/20141017054534/http://www.marinespecies.org/aphia.php?p=taxdetails&id=445309 |url-status=live }}
  • Linnaeus, C. 1758. Systema Naturae per Regna tria Naturae, secundem Classes, Ordines, Genera, Species, cum Characteribus, Differentis, Synonymis, Locis. Tom.1 Editio decima, reformata. Holmiae : Laurentii Salvii 824 pp.
  • Iredale, T. 1927. Caloundra Shells. The Australian Zoologist 4: 331–336, pl. 46
  • Iredale, T. 1929. Queensland molluscan notes, No. 1. Memoirs of the Queensland Museum 9(3): 261–297, pls 30-31
  • Cotton, B.C. & Godfrey, F.K. 1933. South Australian Shells. Part 9. South Australian Naturalist 15(1): 14-24
  • Cotton, B.C. 1943. Australian Shells of the Family Haliotidae. Transactions of the Royal Society of South Australia 67(1): 175-180
  • Moore, R.C. (ed.) 1960. Treatise on Invertebrate Paleontology. Part I. Mollusca 1. Boulder, Colorado & Lawrence, Kansas : Geological Society of America & University of Kansas Press xxiii + 351 pp.
  • Wilson, B. 1993. Australian Marine Shells. Prosobranch Gastropods. Kallaroo, Western Australia : Odyssey Publishing Vol. 1 408 pp.
  • Geiger, D.L. & Poppe, G.T. 2000. A Conchological Iconography. The family Haliotidae. Germany : ConchBooks 135 pp.
  • Bouchet, P.; Gofas, S. (2014). Haliotis Linnaeus, 1758. Accessed through: [http://www.marinespecies.org/aphia.php?p=taxdetails&id=138050 World Register of Marine Species]
  • {{cite book | last = Tryon | first = George W. Jr.| author-link = George Washington Tryon | year = 1880 | title = Manual of Conchology; Structural and Systematic With Illustrations of the Species | volume = II: Muricinæ, Purpurinæ | publisher = Academy of Natural Sciences | location = Philadelphia, PA | url = https://archive.org/details/manualofconcholo2tryo | format = PDF }}