User:Wikirictor/sandbox
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scope="row" style="width: 23%; font-size:100%; background:#fcfebe; color:#000000;"|Title | scope="row" style="width: 17%; font-size:100%; background:#fcfebe; color:#000000;"|Director | scope="row" style="width: 15%; font-size:100%; background:#fcfebe; color:#000000;"|Format | scope="row" style="width: 41%; font-size:100%; background:#fcfebe; color:#000000;"|Info |
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Looney Tunes Mad as a Mars Hare | Chuck Jones | Cartoon Series | Bugs Bunny is turned into a "Neanderthal Rabbit" after getting hit by a ray from a time-projector gun by Marvin the Martian. |
Quest for Fire | Jean-Jacques Annaud | 1981 film | features Neanderthals and a Cro-Magnon attempting to carry a vessel containing fire to the Neanderthal's tribe. |
Clan of the Cave Bear | Michael Chapman | 1986 film | novel by Jean M. Auel about prehistoric times. It is the first book in the Earth's Children book series |
Iceman | Fred Schepisi | 1984 film | from a screenplay written by John Drimmer, depicts a frozen Neanderthal coming to life again in the 1980s at an arctic research station. |
Ghost Light | Andrew Cartmel | 1989 film | an episode in the television series Doctor Who, a Neanderthal called "Nimrod" is the butler of a Victorian era household. |
Night at the Museum | Shawn Levy | 2006 film | four Neanderthals were put on display in the American Museum of Natural History. An ancient Egyptian tablet, the Tablet of Akhemrah, causes everything on display in the museum to come to life at night. The Neanderthals showed an interest in fire after it was shown to them by the night guard, Larry Daley. |
The Real Adventures of Jonny Quest | Sherry Gunther | TV series | the mythical yetis are stated to be a relict population of Neanderthals. |
Dinosaurs | Brian Henson | TV series | Generic "cavemen" have appeared in multiple episodes notably season 3 episodes, "The Discovery," and "Charlene and Her Amazing Humans." |
You Can't Do That on Television | Geoffrey Darby | TV program | A Neanderthal-like family was a frequent recurring sketch in the children's show, In keeping with the theme of that particular episode, the sketch often parodied modern issues with coarse, overbearing parents outside of a pre-historic cave setting.[YCDTOTV.com FAQ "31. Which sets were used for YCDTOTV sketches?" - see "The cave" under Miscellaneous sets. Note: do not correct url formatting as per Wikipedia's Blacklist, June 2010] |
GEICO Cavemen | Joe Lawson | advertisement | trademarked characters in a series of television advertisements for the auto insurance company GEICO that have aired from 2004 to present, featuring Neanderthal-like cavemen in a modern setting |
The Croods | Chris Sanders Kirk DeMicco | 2013 animated film | features the titular family as they embark on a journey to find a new home along with a Cro-Magnon boy who has mastered fire and other "technologies" they had never previously encountered. |
Walking with Beasts | Tim Haines | Documentary | One is charged by a woolly rhinoceros, but escapes, in part because of his stocky constitution. The climax of the episode is when the clan of Neanderthals attack the herd of mammoth as they turn back to the north. |
Ao: The Last Hunter Ao, le dernier Néandertal | Jacques Malaterre | prehistoric film | Ao is the protagonist in a 2010 French prehistoric film{{cite web |url=http://www.ugcdistribution.fr/ao/le-film/ |title= AO, le dernier Néandertal - site officiel du film | publisher= UGC YM |date= |accessdate=July 24, 2016}} |
The contemporary perception of Neanderthals and their stereotypical portrayal has its origins in 19th century Europe. Naturalists and anthropologists were confronted with an increasing number of fossilized bones that that did not match any known taxon. Carl Linnaeuss 1758 Systema Naturae in which he had Homo sapiens introduced as a species without diagnosis and description, was the authoritative encyclopedia of the time. The notion of extinct species was unheard of and if so, would have contradicted the paradigm of the immutability of species and the physical world, which was the infallible product of a single and deliberate act of a creator god.
Most scholars simply declared the early Neanderthal fossils to be representatives of early "races" of modern man. Thomas Henry Huxley, a future supporter of Darwin's theory of evolution, saw in the Engis 2 fossil a "man of low degree of civilization". The discovery in the Neandertal he interpreted as to be within the range of variation of modern humans.Thomas Henry Huxley: On some fossil remains of man. Kapitel 3 in: Evidence as to man's place in nature. D. Appleton and Company, New York 1863
In mid 19th century Germany biological sciences were dominated by Rudolf Virchow, who described the bones as a "remarkable individual phenomenon" and as "plausible individual deformation". This statement is the reason why the characteristics of the Neanderthals were perceived as a form of pathological skeleton change of modern man in German-speaking countries for many years to come.
August Franz Josef Karl Mayer, an associate of Virchow emphasizes disease, prolonged pain and struggle on comparison with modern human features.Martin Kuckenberg: Lag Eden im Neandertal? Auf der Suche nach dem frühen Menschen. Econ Verlag, Düsseldorf 1997, S. 51, {{ISBN|3-430-15773-0}} "He confirmed the Neanderthal's rachitic changes in bone development[...]. Mayer argued among other things, that the thigh - and pelvic bones of Neanderthal man were shaped like those of someone who had spent all his life on horseback. The broken right arm of the individual had only healed very badly and the resulting permanent worry lines about the pain were the reason for the distinguished brow ridges. The skeleton was, he speculated, that of a mounted Russian Cossack, who had roamed the region in 1813/14 during the turmoils of the wars of liberation from Napoleon." Friedemann Schrenk, Stephanie Müller: Die Neandertaler, S. 16
Arthur Keith of Britain and Marcellin Boule of France, were both senior members of their respective national paleontological institutes and among the most eminent paleoanthropologists of the early 20th century. Both men argued that this "primitive" Neanderthal could not be a direct ancestor of modern man. As a result the museum's copy of the almost complete Neanderthal fossil of La Chapelle-aux-Saints was inaccurately mounted in an exaggerated crooked pose with a deformed and heavily curved spine and legs buckled.{{cite web | url=https://archive.org/stream/lhommefossiledel00boul#page/n1/mode/2up | title =L'homme fossile de La Chapelle-aux-Saints - full text - Volume VI (p. 11–172), Volume VII (p. 21–56), Volume VIII (p. 1–70), 1911–1913 | publisher =Royal College of Surgeons of England | date= |accessdate=July 26, 2016 }}{{cite web | url=https://www.britannica.com/biography/Marcellin-Boule | title =Marcellin Boule - French geologist | publisher =Encyclopædia Britannica | date= |accessdate=July 26, 2016 }}{{cite web | url=https://www.therai.org.uk/search?q=Arthur+Keith&Search= | title =Arthur Keith | publisher =Royal Anthropological Institute | date= |accessdate=July 26, 2016 }}{{cite web | url=http://humanorigins.si.edu/evidence/human-fossils/fossils/la-chapelle-aux-saints | title =La Chapelle-Aux-Saints - The old man of La Chapelle - The original reconstruction of the 'Old Man of La Chapelle' by scientist Pierre Marcellin Boule led to the reason why popular culture stereotyped Neanderthals as dim-witted brutes for so many years. | publisher =Smithsonian Institution | date= |accessdate=July 26, 2016 }}
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{{Speciesbox
| taxon = Homo neanderthalensis
| name = Neanderthal
| fossil_range = Middle–Late Pleistocene
| image= Homo sapiens neanderthalensis.jpg
| image_caption = Neanderthal skull at La Chapelle-aux-Saints
| image2 =
| image2_alt =
| image2_caption =
| binomial = Homo neanderthalensis
| binomial_authority = King, 1864
| synonyms =Homo mousteriensis hauseri{{cite book|url=https://books.google.com/books?id=440TmWXToLAC&dq=Homo+mousteriensis+hauseri&pg=PT132 |title=Wiley-Blackwell Encyclopedia of Human Evolution, 2 Volume Set - edited by Bernard Wood |date= 31 March 2011|isbn=9781444342475 |accessdate=September 26, 2016|last1=Wood |first1=Bernard }}
Homo sapiens neanderthalensis{{cite web|url=http://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?id=63221 |title=Homo sapiens neanderthalensis |publisher=National Center for Biotechnology Information, U.S. National Library of Medicine |date= |accessdate=September 26, 2016}}
Palaeoanthropus neanderthalensis[https://books.google.com/books?id=mR_cxcxO8h8C&pg=PA556&lpg=PA556&dq=Palaeoanthropus+neanderthalensis&source=bl&ots=wsgpR8oudS&sig=3TDe6QN_tlrnqVz10HK2s8Ls9Ho&hl=en&sa=X&ei=cnTtUrWtOqWMyQHWg4DgCg&ved=0CHwQ6AEwCw#v=onepage&q=Palaeoanthropus%20neanderthalensis&f=false Bibliography of Fossil Vertebrates 1954-1958 - C.L. Camp, H.J. Allison, and R.H. Nichols - Google Books]. Books.google.ca. Retrieved on 2014-05-24.
| range_map = Range of Homo neanderthalensis.png
| range_map_width = 220px
| range_map_caption = Range of Homo neanderthalensis, that may extend to include the Okladnikov Cave/Altai Mountains and Mamotnaia/Ural Mountains
}}
Neanderthals or Neandertals {{IPAc-en|UK|n|i|ˈ|æ|n|d|ər|ˌ|t|ɑː|l}}, American English also {{IPAc-en|n|eɪ|}}-, -{{IPAc-en|ˈ|ɑː|n|d|ə|r|}}-, -{{IPAc-en|ˌ|t|ɔː|l}}, -{{IPAc-en|ˌ|θ|ɔː|l}}){{cite web|title=Neanderthal in ODE|url=http://www.oxforddictionaries.com/definition/english/Neanderthal|website=Oxford Dictionaries}}{{cite web|title="Neanderthal" in Random House Dictionary (US) & Collins Dictionary (UK)|url=http://dictionary.reference.com/browse/neanderthal|website=Dictionary.com}} are an extinct species or subspecies of the genus Homo, that was a close relative of modern humans. Both species occupied a common habitat in Europe for several thousand years. Research has so far no universally accepted conclusive explanation as to what caused the Neanderthal's extinction between 40.000 and 28.000 years ago.{{cite web|url=http://www.nytimes.com/2006/09/13/science/14neanderthal.html?ex=1315800000&en=ca90a9bfe57071f2&ei=5089&_r=0 |title=Neanderthals' Last Stand Is Traced |work= The New York Times |date=September 13, 2006 |accessdate=September 26, 2016}}
Neanderthal classification has been subject to debate since the discovery of the fdo to this day.{{cite web|url=http://insider.si.edu/2015/08/why-did-neanderthals-go-extinct/ |title=WHY DID NEANDERTHALS GO EXTINCT? - ...debate whether Homo neanderthalensis should be classified as a subspecies of Homo sapiens... |publisher= Smithsonian Insider |date=August 11, 2015 |accessdate=September 26, 2016}} Currently two formal species names reflect the ongoing controversy - Homo neanderthalensis the binominal name of a distinct species and Homo sapiens neanderthalensis, that classifies Neanderthals as a subspecies of Homo sapiens.{{cite journal |doi=10.1073/pnas.0904119106 |title=The origin of Neandertals |year=2009 |last1=Hublin |first1=J. J. |journal=Proceedings of the National Academy of Sciences |volume=106 |issue=38 |pages=16022–7 |pmid=19805257 |jstor=40485013 |bibcode=2009PNAS..10616022H |pmc=2752594|doi-access=free }}{{Cite journal |title=Neanderthal taxonomy reconsidered: implications of 3D primate models of intra- and interspecific differences |pmid=14745010 |last1=Harvati |first1=K. |last2=Frost |first2=S.R. |last3=McNulty |first3=K.P. |date=2004 |doi=10.1073/pnas.0308085100 |pmc=337021 |volume=101 |journal=Proc. Natl. Acad. Sci. U.S.A. |issue=5 |pages=1147–52|doi-access=free }}{{cite web |url=http://www.sci-news.com/othersciences/anthropology/science-neanderthal-genes-modern-human-dna-01734.html |title= Scientists Identify Neanderthal Genes in Modern Human DNA | publisher= Sci-News.com |date=January 30, 2014 |accessdate=September 26, 2016}}
Sharing a common ancestor of the genus Homo with Homo sapiens, who evolved independently in Africa,{{cite journal | last1 = Liu | first1 = Hua | display-authors = etal | year = 2006 | title = A Geographically Explicit Genetic Model of Worldwide Human-Settlement History | doi = 10.1086/505436 | journal = The American Journal of Human Genetics | volume = 79 | issue = 2| pages = 230–237 | quote = Currently available genetic and archaeological evidence is generally interpreted as supportive of a recent single origin of modern humans in East Africa. However, this is where the near consensus on human settlement history ends, and considerable uncertainty clouds any more detailed aspect of human colonization history. | pmid=16826514 | pmc=1559480}} the specie's specific morphology developed in Eurasia during the last 350.000 years, reaching its final, "classical Neanderthal" form approximately 130.000 years ago.{{cite journal|title=Modern Humans Did Not Admix with Neanderthals during Their Range Expansion into Europe - ...morphology developed in Europe during the last 350,000 y under the effect of selection and genetic drift... |date=November 30, 2004 |pmc=532389 |last1=Currat |first1=M. |last2=Excoffier |first2=L. |journal=PLOS Biology |volume=2 |issue=12 |pages=e421 |doi=10.1371/journal.pbio.0020421 |pmid=15562317 }}
Neanderthals left a rich fossil record in the limestone caves of Eurasia, from Western Europe to the Altai Mountains in Central Asia and the Ural Mountains in the North to the Levant in the South.{{cite web|url=https://www.britannica.com/topic/Neanderthal |title=Neanderthal ANTHROPOLOGY - ...Neanderthals inhabited Eurasia from the Atlantic regions... |publisher= Encyclopaedia Britannica |date=January 29, 2015 |accessdate=September 26, 2016}} Discoveries include cultural assemblages and Neanderthals are associated with the Mousterian culture, which first appeared approximately 160.000 years ago.{{cite book|editor1-last=Shaw|editor1-first=Ian|editor2-last=Jameson|editor2-first=Robert|title=A Dictionary of Archaeology|date=1999|publisher=Blackwell|isbn=0-631-17423-0|page=408|url=https://books.google.com/books?id=8HKDtlPuM2oC&dq=mousterian+40%2C000&pg=PA408|accessdate=1 August 2016}}{{cite web|url=http://humanorigins.si.edu/evidence/human-fossils/species/homo-neanderthalensis |title=Homo neanderthalensis - ...The Mousterian stone tool industry of Neanderthals is characterized by... |publisher= Smithsonian Institution |date=September 22, 2016 |accessdate=September 26, 2016}}{{Cite journal|doi=10.1038/nature05195 |date=October 2006 |author1=Finlayson, C |author2=Pacheco, FG |author3=Rodríguez-Vidal, J |author4=Fa, DA |author5=Gutierrez López, JM |author6=Santiago Pérez, A |author7=Finlayson, G |author8=Allue, E |author9=Baena Preysler, J |author10=Cáceres, I |author11=Carrión, JS |author12=Fernández Jalvo, Y |author13=Gleed-Owen, CP |author14=Jimenez Espejo, FJ |author15=López, P |author16=López Sáez, JA |author17=Riquelme Cantal, JA |author18=Sánchez Marco, A |author19=Guzman, FG |author20=Brown, K |author21=Fuentes, N |author22=Valarino, CA |author23=Villalpando, A |author24=Stringer, CB |author25=Martinez Ruiz, F |author26=Sakamoto, T |title=Late survival of Neanderthals at the southernmost extreme of Europe |volume=443 |issue=7113 |pages=850–3 |issn=0028-0836 |pmid=16971951 |journal=Nature|bibcode = 2006Natur.443..850F |s2cid=4411186 }}
Generally small and widely dispersed fossil sites suggest, that Neanderthals lived in less numerous and socially more isolated groups than contemporary Homo sapiens. Tools such as Mousterian flint stone flakes and Levallois points are remarkably sophisticated from the outset, yet they have a slow rate of variability and general technological inertia is noticeable during the entire fossil period. Artefacts are of utilitarian nature, symbolic behavioral traits are undocumented before the arrival of modern humans in Europe around 40.000 to 35.000 years ago.{{cite web|url=http://eol.org/pages/4454114/details |title=Homo neanderthalensis Brief Summary |publisher=EOL |date= |accessdate=September 26, 2016}}{{cite journal | url=https://www.ncbi.nlm.nih.gov/pubmed/25020018 | title =Symbolic or utilitarian? Juggling interpretations of Neanderthal behavior: new inferences from the study of engraved stone surfaces. | year =2014 | pmid =25020018 |accessdate=July 26, 2016 | last1 =Peresani | first1 =M. | last2 =Dallatorre | first2 =S. | last3 =Astuti | first3 =P. | last4 =Dal Colle | first4 =M. | last5 =Ziggiotti | first5 =S. | last6 =Peretto | first6 =C. | journal =Journal of Anthropological Sciences = Rivista di Antropologia : Jass | volume =92 | issue =92 | pages =233–255 | doi =10.4436/JASS.92007 }}
The Neanderthal genome sequence was successfully generated in 2010 as the central assignment for the Neanderthal genome project.{{cite web|url=http://www.eva.mpg.de/neandertal/index.html |title=A high-quality Neandertal genome sequence |publisher= Max Planck Institute for Evolutionary Anthropology |date= |accessdate=September 26, 2016}} The DNA structure proved to be 99.7% identical with anatomically modern humans.[https://www.sciencedaily.com/releases/2010/05/100506141555.htm Complete Neanderthal genome sequenced: DNA signatures found in present-day Europeans and Asians, but not in Africans], ...showed that Neanderthal DNA is 99.7 percent identical to present-day human DNA ScienceDaily Subsequently the genetic relationship is being investigated{{cite web | url=http://kas.berkeley.edu/documents/Issue_102-103/9_Baillie.pdf | title =Neandertals: Unique from Humans, or Uniquely Human? | author =Colin P.T. Baillie | author2 =University of California, Berkeley | publisher =berkeley.edu}}{{cite web | url=http://humanorigins.si.edu/evidence/genetics/ancient-dna-and-neanderthals/neanderthal-mitochondrial-dna | title =Ancient DNA and Neanderthals | author =Smithsonian Institution National Museum of American History | publisher =si.edu}} and ongoing research has yet helped to interpret and verify kinship lines and interbreeding theories with modern humans and other species of the genus Homo and resulting evolutionary implications.{{cite journal |date= October 17, 2012|title= North African Populations Carry the Signature of Admixture with Neandertals|journal= PLOS ONE|doi= 10.1371/journal.pone.0047765|volume=7|pages=e47765|doi-access= free|last1= Sánchez-Quinto|first1= Federico|last2= Botigué|first2= Laura R.|last3= Civit|first3= Sergi|last4= Arenas|first4= Conxita|last5= Ávila-Arcos|first5= María C.|last6= Bustamante|first6= Carlos D.|last7= Comas|first7= David|last8= Lalueza-Fox|first8= Carles|issue= 10|pmid= 23082212|pmc= 3474783}}{{cite journal |date= 23 October 2014|title= Genome sequence of a 45,000-year-old modern human from western Siberia|journal= Nature|volume= 514|issue= 7523|pages= 445–449|doi= 10.1038/nature13810|pmid=25341783|last1= Fu|first1= Qiaomei|last2= Li|first2= Heng|last3= Moorjani|first3= Priya|last4= Jay|first4= Flora|last5= Slepchenko|first5= Sergey M.|last6= Bondarev|first6= Aleksei A.|last7= Johnson|first7= Philip L. F.|last8= Aximu-Petri|first8= Ayinuer|last9= Prüfer|first9= Kay|last10= De Filippo|first10= Cesare|last11= Meyer|first11= Matthias|last12= Zwyns|first12= Nicolas|last13= Salazar-García|first13= Domingo C.|last14= Kuzmin|first14= Yaroslav V.|last15= Keates|first15= Susan G.|last16= Kosintsev|first16= Pavel A.|last17= Razhev|first17= Dmitry I.|last18= Richards|first18= Michael P.|last19= Peristov|first19= Nikolai V.|last20= Lachmann|first20= Michael|last21= Douka|first21= Katerina|last22= Higham|first22= Thomas F. G.|last23= Slatkin|first23= Montgomery|last24= Hublin|first24= Jean-Jacques|last25= Reich|first25= David|last26= Kelso|first26= Janet|last27= Viola|first27= T. Bence|last28= Pääbo|first28= Svante|pmc= 4753769|hdl= 10550/42071}}Brahic, Catherine. [http://www.newscientist.com/article/dn24988-humanitys-forgotten-return-to-africa-revealed-in-dna.html#.VZy0CvkmNOI "Humanity's forgotten return to Africa revealed in DNA"], The New Scientist (February 3, 2014).
In 2013 researchers announced that the assemblages of at least 40 widely scattered archaeological sites suggest that Neanderthals intentionally buried their dead.{{cite news |last=Wilford |first=John Noble |title=Neanderthals and the Dead|url=http://www.nytimes.com/2013/12/17/science/neanderthals-and-the-dead.html |date=December 16, 2013 |work=New York Times |accessdate=December 17, 2013 }} In 2016 a team of researchers found ring structures made of stalagmites fragments inside the Bruniquel Cave, south-western France around 176.000 years old, which were attributed to early Neanderthals. These discoveries might revive the debate on the degree of advancement of Neanderthal's social structure.{{cite journal|doi=10.1038/nature18291|title=Early Neanderthal Constructions deep in Bruniquel Cave in Southwestern France|author1=Jaubert, Jacques|author2=Verheyden, Sophie|author3=Genty, Dominique|author4=Soulier, Michel|author5=Cheng, Hai|author6=Blamart, Dominique|author7=Burlet, Christian|author8=Camus, Hubert|author9=Delaby, Serge|author10=Deldicque, Damien|author11=Edwards, R. Lawrence|author12=Ferrier, Catherine|author13=Lacrampe-Cuyaubère, François|author14=Lévêque, François|author15=Maksud, Frédéric|author16=Mora, Pascal|author17=Muth, Xavier|author18=Régnier, Édouard|author19=Rouzaud, Jean-Noël|author20=Santos, Frédéric|journal=Nature |volume=534|number=7605|date= 2 June 2016|pages=111–114 |pmid=27251286 |orig-year= online 25 May 2016|issn=0028-0836|url=http://www.nature.com/nature/journal/v534/n7605/full/nature18291.html}}
Etymology
The species was named by geologist William King in his 1864 species description The Reputed Fossil Man of the Neanderthal after having introduced the term Homo Neanderthalensis King one year earlier giving a lecture at the British Association for the Advancement of Science.{{cite web|url=http://www.boneandstone.com/articles_classics/king_1864.pdf |title=The Reputed Fossil Man of the Neanderthal. by Professor Wiiliam King |publisher= Bone and Stone |date= |accessdate=September 26, 2016}}William King: On the Neanderthal Skull, or Reasons for believing it to belong to the Clydian Period and to a species different from that represented by Man. In: British Association for the Advancement of Science, Notices and Abstracts for 1863, Part II, London, 1864, p. 81 f.William King: The Neanderthal Skull. In: The Anthropological Review. Vol. 1, Nr. 3, 1863, S. 393–394, [https://www.jstor.org/stable/pdf/3024846.pdf?_=1464700412077 full text (PDF)] The type specimen Neanderthal 1 was only discovered in 1856 in a limestone grotto in the Neandertal (Neander-valley, spelled Neanderthal before the 1901 German spelling reform), {{convert|12|km|mi|abbr=on}} east of Düsseldorf, Germany.Kunzig, Robert. [http://discovermagazine.com/1998/jan/theyearinscience1346 "The Year in Science: Human Origins 1997"], Discover (magazine) (January 1, 1998) reprinted in [https://books.google.com/books?id=6B37S4LW1MIC Contemporary Readings in Physical Anthropology], p. 145 (Alan Almquist ed., Prentice Hall, 2000)
Discovery
{{multiple image
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| image1 = Neanderthal-Fundstelle in Erkrath 2008.jpg
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| image2 = Neanderthal position.png
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| footer = Neanderthal excavation site and Neander Valley in North Rhine-Westphalia, Germany
}}
The first recorded Neanderthal fossil of a partial skull and some associated fragments, later dubbed Engis 2 was discovered in 1829 by Dutch physician and naturalist Philippe-Charles Schmerling in the Awirs Cave, just north of the Belgian municipality Engis{{cite web |url=http://humanorigins.si.edu/evidence/3d-collection/engis-2 |title= Engis 2 | publisher= Smithsonian Institution |date= |accessdate=September 27, 2015}} and professionally described in 1833.Philippe-Charles Schmerling: Recherches sur les ossements fossiles découverts dans les cavernes de la Province de Liège. P.-J. Collardin, Liège 1833, pp. 1–66{{cite web |url=https://www.nespos.org/display/PublicNesposSpace/Engis+3+-+individuum+Engis+2 |title= Engis 3 - individuum Engis 2 - Public NESPOS Space | publisher= NESPOS Society |date= |accessdate=September 27, 2015}} The second fossil Gibraltar 1, was discovered by Captain Edmund Flint in the Forbes' Quarry of Gibraltar in 1848. Both findings received only little attention and were misunderstood as to represent early forms or pathologically developed, abnormal human individuals.Thomas Henry Huxley: On some fossil remains of man. chapter 3 in: Evidence as to man's place in nature. D. Appleton and Company, New York 1863 A universal definition of species-specific characteristics of species in the genus Homo did not exist in the early 19th century. Methodical recognition of distinct and extinct species only began after Charles Darwin's publication of On the Origin of Species in 1859.Thomas Henry Huxley: On some fossil remains of man. Kapitel 3 in: Evidence as to man's place in nature. D. Appleton and Company, New York 1863
The 1856 in a limestone quarry - the Kleine Feldhofer Grotte in the Neandertal discovered type specimen consisted of a skull cap, two femora, three bones from the right arm, two from the left arm, part of the left ilium, fragments of a scapula, and ribs. These quickly came into possession of local teacher and amateur naturalist Johann Carl Fuhlrott, who studied the fossil in co-operation with anatomist Hermann Schaaffhausen. The jointly made publication in 1857 represents the beginning of paleoanthropology as a scientific discipline.{{cite web |url=http://archaeologyinfo.com/homo-neanderthalensis/ |title=Homo neanderthalensis |publisher=Patrick Johnson | date= |accessdate=September 26, 2016}}{{cite web |url=http://www.pnas.org/content/96/13/7117.full |title= Hominids and hybrids: The place of Neanderthals in human evolution by Ian Tattersall and Jeffrey H. Schwartz | publisher= National Academy of Sciences |date= |accessdate=September 26, 2016}} The scholarly community remained divided for another 30 years until the discovery of two well preserved and almost complete Neanderthal skeletons in Spy/Belgium in 1886 lead to general recognition of the species.Ian Tattersall: Neandertaler. Der Streit um unsere Ahnen. Birkhäuser Verlag, Basel 1999, S. 81, {{ISBN|3-7643-6051-8}}.
Origin
Altamura Man (between 128.000 and 187.000 years old) is the oldest fossil of the Neanderthal lineage, that has yielded a reliable DNA sequence.{{Cite web|url = http://www.ancient-origins.net/news-evolution-human-origins/calcified-altamura-man-oldest-neanderthal-dna-sample-020285 | title= Calcified remains of Altamura Man yield oldest ever Neanderthal DNA sample | publisher=Ancient Origins | date=April 4, 2015 | accessdate=September 3, 2016 }} Older specimen are until recently interpreted and classified based on morphology and metric analyses. 430.000 years old skull fossils recovered from the Sima de los Huesos cave have proto-Neanderthal features, such as "the beginning of a prominent brow ridge, a distinctive jaw shape and patterns of cusps on the teeth, [but] lack other traits that define the species — notably a large cranium".{{cite web|url=http://www.nature.com/news/pit-of-bones-catches-neanderthal-evolution-in-the-act-1.15430 |title='Pit of bones' catches Neanderthal evolution in the act |publisher=Nature |date=June 19, 2014 |accessdate=September 27, 2016}}{{cite web |url=http://www.scientificamerican.com/article/oldest-ancient-human-dna-details-dawn-of-neandertals/ |title= Oldest Ancient-Human DNA Details Dawn of Neandertals | publisher= SCIENTIFIC AMERICAN |date=March 14, 2016 |accessdate=September 26, 2016}}{{cite web |url=http://humanorigins.si.edu/evidence/human-fossils/species/homo-heidelbergensis |title= Homo heidelbergensis - Comparison of Neanderthal and modern human DNA suggests that the two lineages diverged from a common ancestor, most likely Homo heidelbergensis | publisher= Smithsonian Institution |date= |accessdate=September 26, 2016}} DNA tests confirm that these skulls represent early Neanderthals.{{cite web|url=http://www.nature.com/news/oldest-ancient-human-dna-details-dawn-of-neanderthals-1.19557 |title=Oldest ancient-human DNA details dawn of Neanderthals |publisher=Nature |date=March 14, 2016 |accessdate=September 27, 2016}} Anthropologists debate over whether the species evolved from Homo heidelbergensis, or from what some researchers call "a controversial species" - Homo antecessor.{{cite web|url=http://www.smithsonianmag.com/science-nature/homo-antecessor-common-ancestor-of-humans-and-neanderthals-143357767/?no-ist |title=Homo antecessor: Common Ancestor of Humans and Neanderthals? |publisher=Smithsonian |date=November 26, 2012 |accessdate=September 27, 2016}}
"Not quite Neanderthals, the Sima de los Huesos humans probably represent one of many small, isolated groups that dotted the European continent at the time, notes Jean-Jacques Hublin, a palaeoanthropologistat at the Max Planck Institute for Evolutionary Anthropology in Leipzig, Germany. Such isolation is ideal for anatomical features to be established by chance, and this genetic drift could explain many of the attributes that would later define Neanderthals."
Fossils found in Tourville-la-Rivière,{{cite web|url=http://www.lemonde.fr/archeologie/article/2014/10/09/le-bras-d-un-pre-neandertalien-trouve-en-bord-de-seine_4503785_1650751.html |title=Le bras d'un pré-Néandertalien trouvé au bord de la Seine] » |publisher=Le Monde |date=October 9, 2014 |accessdate=September 28, 2016}} the 400.000 year old Tautavel Man, the 600.000 year old mandible Mauer 1 and the Petralona skull have all been attributed to the Neanderthal lineage, although only based on morphological and metric analyses.{{cite journal|title=Middle Pleistocene Human Remains from Tourville-la-Rivière (Normandy, France) and Their Archaeological Context |journal=PLOS ONE |date=October 8, 2014 |doi=10.1371/journal.pone.0104111 |doi-access=free |last1=Faivre |first1=Jean-Philippe |last2=Maureille |first2=Bruno |last3=Bayle |first3=Priscilla |last4=Crevecoeur |first4=Isabelle |last5=Duval |first5=Mathieu |last6=Grün |first6=Rainer |last7=Bemilli |first7=Céline |last8=Bonilauri |first8=Stéphanie |last9=Coutard |first9=Sylvie |last10=Bessou |first10=Maryelle |last11=Limondin-Lozouet |first11=Nicole |last12=Cottard |first12=Antoine |last13=Deshayes |first13=Thierry |last14=Douillard |first14=Aurélie |last15=Henaff |first15=Xavier |last16=Pautret-Homerville |first16=Caroline |last17=Kinsley |first17=Les |last18=Trinkaus |first18=Erik |volume=9 |issue=10 |pages=e104111 |pmid=25295956 |pmc=4189787 }}
Classification
Scientists to this day debate over whether Neanderthals should be classified as a distinct species - Homo neanderthalensis - or as Homo sapiens neanderthalensis, the latter placing Neanderthals as a subspecies of H. sapiens.{{cite journal |doi=10.1073/pnas.96.13.7117 |title=Hominids and hybrids: The place of Neanderthals in human evolution |year=1999 |last1=Tattersall |first1=Ian |last2=Schwartz |first2=Jeffrey H. |journal=Proceedings of the National Academy of Sciences |volume=96 |issue=13 |pages=7117–9 |jstor=48019 |bibcode=1999PNAS...96.7117T |pmid=10377375 |pmc=33580|doi-access=free }}{{cite journal |doi=10.1073/pnas.96.13.7604 |title=The early Upper Paleolithic human skeleton from the Abrigo do Lagar Velho (Portugal) and modern human emergence in Iberia |year=1999 |last1=Duarte |first1=Cidália |last2=Mauricio |first2=João |last3=Pettitt |first3=Paul B. |last4=Souto |first4=Pedro |last5=Trinkaus |first5=Erik |last6=Van Der Plicht |first6=Hans |last7=Zilhao |first7=João |journal=Proceedings of the National Academy of Sciences |volume=96 |issue=13 |pages=7604–9 |jstor=48106 |bibcode=1999PNAS...96.7604D |pmid=10377462 |pmc=22133|doi-access=free }} During the early 20th century the prevailing view has been heavily influenced by Arthur Keith and Marcellin Boule, who wrote the first scientific description of a nearly complete Neanderthal skeleton. {{cite web | url=https://archive.org/stream/lhommefossiledel00boul#page/n1/mode/2up | title =L'homme fossile de La Chapelle-aux-Saints - full text - Volume VI (p. 11–172), Volume VII (p. 21–56), Volume VIII (p. 1–70), 1911–1913 | publisher =Royal College of Surgeons of England | date= |accessdate=July 26, 2016 }} Senior members of their respective national paleontological institutes and among the most eminent paleoanthropologists of their time,{{cite web | url=https://www.britannica.com/biography/Marcellin-Boule | title =Marcellin Boule - French geologist | publisher =Encyclopædia Britannica | date= |accessdate=July 26, 2016 }}{{cite web | url=https://www.therai.org.uk/search?q=Arthur+Keith&Search= | title =Arthur Keith | publisher =Royal Anthropological Institute | date= |accessdate=July 26, 2016 }} both men argued that this "primitive" Neanderthal could not be a direct ancestor of modern man. This idea was reflected in an erroneous and inaccurate reconstruction of the Neanderthal findings of La Chapelle-aux-Saints, mounted in a crooked pose with a deformed and heavily curved spine and legs buckled.{{cite web | url=http://humanorigins.si.edu/evidence/human-fossils/fossils/la-chapelle-aux-saints | title =La Chapelle-Aux-Saints - The old man of La Chapelle - The original reconstruction of the 'Old Man of La Chapelle' by scientist Pierre Marcellin Boule led to the reason why popular culture stereotyped Neanderthals as dim-witted brutes for so many years. | publisher =Smithsonian Institution | date= |accessdate=July 26, 2016 }}
During the 1930s scholars Ernst Mayr, George Gaylord Simpson and Theodosius Dobzhansky reinterpreted the existing fossil record and came to different conclusions.{{cite web | url=http://www.scientificamerican.com/article/our-neandertal-brethren/ | title =Our Neandertal Brethren: Why They Were Not a Separate Species | publisher =SCIENTIFIC AMERICAN | date=August 1, 2010 |accessdate=July 26, 2016 }} Neanderthal man was classified as Homo sapiens neanderthalensis - an early subspecies of what was now called Homo sapiens sapiens. The obviously unbroken succession of fossil sites of both subspecies in Europe was considered evidence that there was a slow and gradual evolutionary transition from Neanderthals to modern humans. Contextual interpretations of similar excavation sites in Asia lead to the hypothesis of multiregional origin of modern man in the 1980s.{{cite journal | last1=Wolpoff | first1=MH | last2=Hawks |first2=J |authorlink2=John D. Hawks |last3=Caspari |first3=R | year=2000 | title=Multiregional, not multiple origins | journal=American Journal of Physical Anthropology | volume=112 | issue=1 | pages=129–36 | url=http://www-personal.umich.edu/~wolpoff/Papers/Multiregional.PDF |doi=10.1002/(SICI)1096-8644(200005)112:1<129::AID-AJPA11>3.0.CO;2-K |pmid=10766948| hdl=2027.42/34270 }}
=Contemporary views=
Current scientific ideas hold that both species evolved from a common African ancestor - Homo erectus. Fossil and tool finds support the idea that during the first migration wave of around 2 million years ago small groups of Homo erectus left Africa via the Levant, the Black Sea area into Georgia (fossils of Dmanisi) and possibly via North-western Africa towards southern Spain. The 1.2 million years old fragmented finds in Spain are referred to as Homo antecessor by their discoverers and considered to be the ancestors of Neanderthal. This interpretation, however, is controversial.
Some 600,000 years ago a second propagation wave of African Homo erectus took place. Skulls from that period found in Spain for example, suggest a brain volume of between 1100 ccm to 1450 ccm. The brain volume of fossils from the first propagation wave is however estimated only to be slightly above 1000 ccm. Homo erectus that arrived in Europe in the second wave of colonization developed into Neanderthals through the intermediate Homo heidelbergensis. Since about 200,000 years ago in Africa the so-called early anatomically modern humans descended from the local Homo erectus through the intermediate Homo rhodesiensis. Evidence from mitochondrial DNA studies has been interpreted as evidence Neanderthals were not a subspecies of H. sapiens.{{Cite journal|author=Hedges SB |title=Human evolution. A start for population genomics |journal=Nature |volume=408 |issue=6813 |pages=652–3 |date=December 2000|pmid=11130051 |doi=10.1038/35047193 |s2cid=3240904 |url= }} Others, for example University of Cambridge Professor Paul Mellars, say "no evidence has been found of cultural interaction".{{Cite news|url=http://www.abc.net.au/news/newsitems/200509/s1450949.htm |title=Modern humans, Neanderthals shared earth for 1,000 years |date=1 September 2005 |accessdate=19 September 2006 |publisher=ABC News (Australia) |url-status=dead |archiveurl=https://web.archive.org/web/20060115225948/http://www.abc.net.au/news/newsitems/200509/s1450949.htm |archivedate=January 15, 2006 }}
Debated remains the period when the Neanderthal lineage separated from the lineage that lead to modern man. In 2010 a period between 440,000 - 270,000 years B.P. was calculated by the molecular clock method. However, the reliability of molecular clocks is questionable as dating determined by stratigraphic methods varies considerably from those determined using the molecular clock. In 2012 recalculation of the mutation rates revealed evidence of a much earlier separation between 800,000 and 400,000 years ago.
Timeline of discovery and research
Neanderthals are after H. sapiens the best documented hominin species as they habitually occupied limestone caves and rock shelters, locations with a high fossilization and preservation ratio for bones, artefacts and fire places. The bones of over 400 specimen have been found since the early 19th century.{{cite web|url=http://www.sciencedaily.com/releases/2007/09/070912154630.htm | title= New Evidence On The Role Of Climate In Neanderthal Extinction - The bones of more than 400 Neanderthals have been found since... | publisher= Science Daily |date=September 13, 2007 |accessdate=October 12, 2016}}
class="wikitable" style="text-align:center; margin: auto; width:99%"
! colspan="33" style="text-align:center; font-size:130%; background:#fcfebe; color:#000000;"|Timeline and locations of discovery, specimen age, associated research and data |
style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Year
!style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Discovery !style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Research !style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Fossil age !style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Location !style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Notability !style="font-size:90%; text-align:center; background:#bee5fe; color:#000000;"|Image |
---|
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1829
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Engis 2 by Philippe-Charles Schmerling !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| described in 1833 but not specified !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Awirs Cave, Belgium !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| first fossil discovery !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Engis 2 Schaedel 1.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1848
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Gibraltar 1 Skull by Captain Edmund Flint !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Forbes' Quarry Gibraltar !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neanderthal skull from Forbes' Quarry.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1856
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Neanderthal 1 by Johann Karl Fuhlrott !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| described as a separate species and published in 1857, but not universally recognized !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 40.000 BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Neandertal, Germany !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| first paleo-anthropologic publication, !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neander1.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1864
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| William King introduces the species name Homo neanderthalensis !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:William King.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1880
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Šipka cave mandible by Karel Jaroslav Maška !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|first discovery of Neanderthal remains in their cultural context !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Šipka, Czech Republic !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| first Mousterian tools discovered !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Sipka Cave Stramberk CZ 02.JPG |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1886
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Spy Skeletons by Marcel de Puydt and Max Lohest !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| discovery lead to international scientific recognition of the existence of more than one human species !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Spy Cave, Belgium !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| cave site was also occupied by H. sapiens, who might have lived alongside Neanderthal !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Spy Skull.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1899
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Krapina Fossils by Dragutin Gorjanović-Kramberger !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| over eight hundred fossil remains discovered, first evidence for morphological variations !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Hušnjakovo Hill, Krapina, Croatia !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| fossils prove Neanderthal evolution !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Krapina 3. Homo neanderthalensis.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1908
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| La Chapelle-aux-Saints 1 by Jean and Amédée Bouyssonie, L. Bardon{{cite web|url=http://eol.org/pages/4454114/details |title= Neanderthal - Homo neanderthalensis - Details - Key Fossils - La Chapelle-aux-Saints | publisher=Encyclopedia of Life |date=|accessdate=July 18, 2016 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| discovery of over 1,000 pieces of stone (flint) and bone tools, Mousterian assemblages !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 60,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| La Chapelle-aux-Saints, France !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| first recognized burial of a Neanderthal !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neandertal skull from la chapelle aux saints.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1909
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| La Ferrassie Skull by Denis Peyrony and Louis Capitan !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Eight buried Neanderthals have been found in La Ferrassie, including infants and two fetuses in a cave flanked by two rock shelters !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 70–50,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Savignac-de-Miremont, Dordogne, France. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The La Ferrassie 1 skull is the largest and most complete Neanderthal skull ever found !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Ferrassie skull.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1909
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Le Moustier skull with a large nasal cavity and a somewhat less developed brow ridge and occipital bun, as might be expected in a juvenile. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The site are two rock shelters near a village in the Dordogne, first excavated from 1863 by the Englishman Henry Christy and the Frenchman Édouard Lartet. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 45,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Peyzac-le-Moustier, Dordogne, France. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Mousterian tool culture is named after this site !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Museum für Vor- und Frühgeschichte Berlin 069.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1926
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Skull fragments of Gibraltar 2, a four-year-old Neanderthal girl, discovered by Dorothy Garrod !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Devil's Tower cave, Gibraltar !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| File:Homo neanderthalensis face (University of Zurich).JPG |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1953–1957
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Ralph Solecki uncovered nine Neanderthal skeletons in Shanidar Cave, Kurdistan !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| A Mousterian layer which contained hundreds of stone tools including points, side-scrapers, flakes and bones, good condition of fossils yields data and information on individual specimen. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 35,000 to 65,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Shanidar, Irak !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Shanidar 1, survived severe injuries attributed to care from other members of his band, Shanidar 4, lay beside a flower that might be explained as evidence of burial rituals - or animal action. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Shanidar skull.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1961-1964
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Amud 1 - the skeleton of an adult male about 25 years old by University of Tokyo expeditions !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Amud 1 has a cranial capacity of about 1,740 cubic cm (106 cubic inches), which is significantly larger than the average capacity of modern humans. Nonetheless, the skull has Neanderthal features, with browridges and a receding forehead. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 50,000 to 70,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Nahal Amud Upper Galilee !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| evidence of Neanderthal habitation and Mousterian toolmaking were revealed, including flaked blades and points as well as deer, cattle, horse, pig, and fox remains.{{cite web|url=https://www.britannica.com/place/Amud-anthropological-and-archaeological-site-Israel#ref891416 |title=Amud |publisher=encyclopaedia britannica |date=|accessdate=October 13, 2016}} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Amud 1. Homo neanderthalensis.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1975
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Erik Trinkaus' study of Neanderthal feet confirmed they walked like modern humans. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neanderthal Foot Print.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1981
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Teeth and part of a jawbone at the Bontnewydd Palaeolithic site by Dr Stephen Aldhouse Green !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The teeth show evidence of taurodontism, enlarged pulp cavities and short roots, which is characteristic of Neanderthals, and although it is not unique to them it is one of the reasons that the species was identified as Neanderthal !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 230,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|near St Asaph, Denbighshire, Wales !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| the most north-western fossil site in Eurasia. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neanderthaler Fund.png |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1982-1987
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| "Moshe" skeleton discovered in 1982 by Professor Ofer Bar-Yosef !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| In 1987 Thermo-luminescence results date Neanderthals at Kebara ("Moshe") to 60,000 BP. Electron spin resonance (ESR) date Es Skhul to 80,000 BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 60,000 to 80,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|Kebara Cave, Es Skhul, Israel !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Neanderthals present in the region from 200,000 to 45,000 BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neanderthal-burial.gif |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1991
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Tabun Cave contains a Neanderthal-type female !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| ESR dates showed the Tabun Neanderthal was contemporaneous with modern humans from Skhul and Qafzeh. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 120,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Nahal Me'arot Nature Reserve, Israel !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The cave was occupied intermittently during the Lower and Middle Paleolithic ages (500,000 to around 40,000 years ago). !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Panel2-tabun 1.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1993
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Scladina Neanderthal child. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The specimen includes a partial cranium and jaw bone, yielding a 127,000 years DNA sample extracted from a tooth molar. He or she is thought to be 8 years and 17 days old at the time of death !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 100–127,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Scladina, Andenne, Belgium !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Two Neanderthal occupations were identified: one dating from 130,000 years, and the other 40 000 years. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Scladina01.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1994
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Sidrón Cave Neanderthal discovery !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| remains of least 12 individuals: three men, three adolescent boys, three women, and three infants inadvertently uncovered. In 2009 Neanderthal ancient mtDNA was partially sequenced in HVR region for three distinct Neanderthals there. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Piloña municipality, Asturias, northwestern Spain !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The first sequencing of the Neanderthal Y chromosome was successfully completed from a specimen from Sidrón Cave[http://www.upf.edu/cexs/actualitat/neandarthals.htm "Universitat Pompeu Fabra, Barcelona, A paleogenetical study determines the blood group of Neanderthal man"] {{webarchive |url=https://web.archive.org/web/20121122123322/http://www.upf.edu/cexs/actualitat/neandarthals.htm |date=November 22, 2012 }}{{cite journal | url = http://www.sciencedirect.com/science/article/pii/S0940960211000641 | doi=10.1016/j.aanat.2011.01.014 | volume=194 | title=Palaeogenetic research at the El Sidrón Neanderthal site | journal=Annals of Anatomy - Anatomischer Anzeiger | year=2012 | pages=133–137| last1=Lalueza-Fox | first1=Carles | last2=Rosas | first2=Antonio | last3=Rasilla | first3=Marco de la | issue=1 | pmid=21482084 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Sidron 2.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1997
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Matthias Krings et al. are the first to amplify Neanderthal mitochondrial DNA (mtDNA) using a specimen from the Feldhofer grotto in the Neander valley{{Cite journal|pmid=9230299 | year=1997 |author1=Krings, M |author2=Stone, A |author3=Schmitz, RW |author4=Krainitzki, H |author5=Stoneking, M |author6=Pääbo, S |title=Neandertal DNA sequences and the origin of modern humans |volume=90 |issue=1 |pages=19–30 |issn=0092-8674 |journal=Cell |doi=10.1016/S0092-8674(00)80310-4| s2cid=13581775 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Germany !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1997–2000
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| During new excavations in the Neandertal, additional bone fragments are found, some of which fit the fragments found in 1856, thus pinpointing the exact location of the original find. The exact location had previously been unknown, as the site of the find (the „Kleine Feldhofer Grotte“) was destroyed by limestone mining.{{ cite journal | last1=Schmitz | first1=Ralf W | display-authors=etal | year=2002 | title=The Neandertal type site revisited: Interdisciplinary investigations of skeletal remains from the Neander Valley, Germany | journal=Proceedings of the National Academy of Sciences | volume=99| issue=20 | pages= 13342–13347 | doi=10.1073/pnas.192464099 | pmid=12232049 | pmc=130635| doi-access=free }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Germany !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Neanderthal-Fundstelle in Erkrath 2008.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 1998
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The Lapedo child by archeologist João Zilhão !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|an early Upper Paleolithic human burial in which provided evidence of early modern humans from the west of the Iberian Peninsula. The remains, a largely complete skeleton of an approximately 4-year-old child, buried with pierced shell and red ochre{{cite journal |doi=10.1073/pnas.96.13.7604 |display-authors=et al. |last1=Duarte |first1=C |year=1999 |title=The early Upper Paleolithic human skeleton from the Abrigo do Lagar Velho (Portugal) and modern human emergence in the Iberian Peninsula |publisher=PNAS |volume=96 |issue=13 |pages=7604–7609 |pmid=10377462 |last2=Maurício |first2=J |last3=Pettitt |first3=PB |last4=Souto |first4=P |last5=Trinkaus |first5=E |last6=Van Der Plicht |first6=H |last7=Zilhão |first7=J |pmc=22133 |journal=Proceedings of the National Academy of Sciences of the United States of America|bibcode = 1999PNAS...96.7604D |doi-access=free }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 24,500 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Abrigo do Lagar Velho, Portugal !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The cranium, mandible, dentition, and postcrania present a mosaic of European early modern human and Neanderthal features. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Abrigodolagarvelho.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2000
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Neanderthal infant of Mezmaiskaya Cave by Igor Ovchinnikov et al. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|Ancient DNA was recovered for a mtDNA sequence showing 3.48% divergence from that of Neanderthal 1, some {{convert|2500|km|mi|0|abbr=on}} to the west. Phylogenetic analysis places the two in a clade distinct from modern humans, suggesting that their mtDNA types have not contributed to the modern human mtDNA pool.{{Cite journal | last1=Ovchinnikov | first1=IV | last2=Götherström | first2=A | last3=Romanova | first3=GP | last4=Kharitonov | first4=VM | last5=Lidén | first5=K | last6=Goodwin | first6=W | year=2000 | title=Molecular analysis of Neanderthal DNA from the northern Caucasus | journal=Nature |volume=404 |issue=6777 |pages=490–3 | pmid=10761915 | doi=10.1038/35006625 | s2cid=3101375 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 29,195 ± 965 years B.P.. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Russian Republic of Adygea, Caucasus Mountains !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Вид на посёлок Мезмай.jpg |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2005
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|The Max Planck Institute for Evolutionary Anthropology launched the Neanderthal genome project{{cite web|url=http://www.nature.com/nature/journal/v444/n7117/full/nature05336.html |title= Analysis of one million base pairs of Neanderthal DNA | publisher=Nature |date=November 16, 2006|accessdate=July 18, 2016 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| In co-operation with Connecticut-based 454 Life Sciences and the Institute of Quaternary Paleontology and Geology in Zagreb, Croatia. In 2009, the Max Planck Institute announced the "first draft" of a complete Neanderthal genome is completed.{{Cite news|first=James |last=Morgan |date=12 February 2009 |title=Neanderthals 'distinct from us' |url=http://news.bbc.co.uk/2/hi/science/nature/7886477.stm |publisher=BBC News |accessdate=22 May 2009}} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Leipzig MPI-EVA.JPG |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2010
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Comparison of Neanderthal genome with modern humans from Africa and Eurasia shows that 1–4% of modern non-African human genome might come from the Neanderthals. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Human-Neandertal mtDNA-HE.png |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2010
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Discovery of Neanderthal tools far away from the influence of H. sapiens indicate that the species might have been able to create and evolve tools on its own, and therefore be more intelligent than previously thought. Furthermore, it was proposed that the Neanderthals might be more closely related to Homo sapiens than previously thought and that may in fact be a subspecies of it. Evidence has more recently emerged that these artifacts are probably of H. sapiens sapiens origin.{{cite web|url=http://news.discovery.com/archaeology/neanderthals-more-intelligent-than-thought.html |title=Neanderthals More Intelligent Than Thought|date=September 22, 2010|accessdate=September 27, 2010 |url-status=dead |archiveurl=https://web.archive.org/web/20100925191858/http://news.discovery.com/archaeology/neanderthals-more-intelligent-than-thought.html |archivedate=September 25, 2010 }}{{Cite journal | last1 = Benazzi | first1 = S. | last2 = Douka | first2 = K. | last3 = Fornai | first3 = C. | last4 = Bauer | first4 = C. C. | last5 = Kullmer | first5 = O. | last6 = Svoboda | first6 = J. Í. | last7 = Pap | first7 = I. | last8 = Mallegni | first8 = F. | last9 = Bayle | first9 = P. | last10 = Coquerelle | doi = 10.1038/Nature10617 | first10 = M. | last11 = Condemi | first11 = S. | last12 = Ronchitelli | first12 = A. | last13 = Harvati | first13 = K. | last14 = Weber | first14 = G. W. | title = Early dispersal of modern humans in Europe and implications for Neanderthal behaviour | journal = Nature | volume = 479 | issue = 7374 | pages = 525–528 | year = 2011 | pmid = 22048311| pmc = | s2cid = 205226924 }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File: |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2012
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Charcoal found next to six paintings of seals in the Nerja caves, Malaga, Spain, has been dated to be between 42,300 and 43,500 years old. The paintings themselves have not been dated until 2014 and if their pigment matches the date of the charcoal, they would be the oldest known cave paintings. José Luis Sanchidrián at the University of Cordoba, Spain believes the paintings are more likely to have been painted by Neanderthals than early modern humans.{{cite web|url=http://www.newscientist.com/article/dn21458-first-neanderthal-cave-paintings-discovered-in-spain.html|title=First Neanderthal cave paintings discovered in Spain |author=Fergal MacErlean|date=10 February 2012|accessdate=10 February 2012|work=New Scientist}}{{cite web|url=https://paleolithicpersonhood.wordpress.com/category/cave-art/ |title=Our Artistic Relatives: Did Neanderthals Have the Cognitive Capacity to Produce Parietal Art? The Case of El Castillo Cave… |publisher=Global Multimedia Protocols Group|date=March 29, 2014|accessdate=October 13, 2016}} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File:Cuevas de Nerja.JPG |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2013
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|An individual’s jaw, which was unearthed at the Riparo di Mezzena rock-shelter{{cite web|url=http://news.discovery.com/human/evolution/neanderthal-skeleton-provides-evidence-of-interbreeding-with-humans-130327.htm|title=First Love Child of Human, Neanderthal Found|first=Jennifer|last= Viegas|date=27 March 2013|work=Discovery News|accessdate=11 April 2013}} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The jawbone has features intermediate between Neanderthals and Homo sapiens suggesting it could be a hybrid. The mitochondrial DNA is Neanderthal. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|40,000-30,000 years BP. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|Riparo di Mezzena, Monti Lessini region, Italy. !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2014
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|Researchers at the University of Colorado Museum in Boulder !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| Neanderthals were not less intelligent than modern humans and "single-factor explanations for the disappearance of the Neandertals are not warranted any more."{{cite web|title=Neanderthals were not less intelligent than modern humans, scientists find|website=TheGuardian.com |url=http://www.theguardian.com/science/2014/apr/30/neanderthals-not-less-intelligent-humans-scientists|date=April 30, 2014|accessdate=April 30, 2014}} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File: |
style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| 2014
!style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|Prof Thomas Higham of the University of Oxford !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| The most comprehensive dating of Neanderthal bones and tools ever carried out, which demonstrated that Neanderthals died out in Europe between 41,000 and 39,000 years ago.{{cite book |url=https://books.google.com/books?id=5TRHOmTUTP4C&dq=Engis+1&pg=PA338 |title= The Human Lineage by Matt Cartmill, Fred H. Smith |date= 30 March 2009|isbn= 9780471214915 |accessdate=September 27, 2015|last1= Cartmill |first1= Matt |last2= Smith |first2= Fred H. }} !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"| !style="font-size:80%; text-align:center; background:#e4f5ff; color:#000000;"|File: |
=Chronology=
Behavior
The behavioral patterns of the Neanderthals are inferred from their anatomy in combination with the fossil record, the remains and debris they left behind at caves, rock shelters and a number of open air settlements. On the contrary behavior has become the key factor in explaining the variability of the archaeological record.
Neanderthals shared a number of important characteristics with modern humans, such as large brains, manual dexterity and bipedal walking ability, tool use and production, which requires a certain degree of social sophistication. However, Neanderthals lived in relatively small and territorial more isolated groups. Generally small sites and the considerable depth of bone and debris pits suggests that they frequently migrated but retured to the same locations often.{{cite web|url=https://www.britannica.com/topic/Neanderthal/Neanderthal-behaviour |title=NEANDERTHAL BEHAVIOUR |publisher=encyclopaedia britannica |date=|accessdate=October 13, 2016}}
The materials left behind show only minor variations among sites and their foraging systems were considerably less efficient than those of modern human hunter-gatherers, without planned differential use of the landscape necessitating more-muscular limbs and greater endurance.{{cite web|url=http://www.nature.com/scitable/knowledge/library/neanderthal-behavior-59267999 |title=Neanderthal Behavior |publisher=Nature |date=|accessdate=October 13, 2016}}{{cite web|url=http://www.livescience.com/28036-neanderthals-facts-about-our-extinct-human-relatives.html |title=Neanderthals: Facts About Our Extinct Human Relatives |publisher=Live Science |date=April 13, 2016|accessdate=October 13, 2016}}{{cite web|url=http://www.smithsonianmag.com/science-nature/rethinking-neanderthals-83341003/?no-ist=&page=2 |title=Rethinking Neanderthals |publisher=Smithsonian |date=April 13, 2016|accessdate=October 13, 2016}}
=Regional differences=
Habitat and culture of "classical Neandertals" since about 125,000 years ago stretched from western Europe all the way to the Levant in the Middle East and beyond the Crimean Peninsula towards Siberia.{{cite book|url=https://books.google.com/books?id=oA7iMFhbvBYC&dq=Neanderthal+behavior&pg=PA1 |title=High Resolution Archaeology and Neanderthal Behavior: Time and Space in ...edited by Eudald Carbonell i Roura |date=13 March 2012|isbn=9789400739222 |accessdate=October 13, 2016|last1=Roura |first1=Eudald Carbonell I. }} Behavior was influenced by climate, terrain, seasonal access to drinking water, the presence of game and other food-resources. Migrations patterns were influenced by the search for raw materials for stone tools.
Elmenteitan
{{
Infobox archaeological culture
|name = Elmenteitan
|map = Lake Elmenteita satellite image.png
|mapalt = Lake Elmenteita satellite image
|altnames = Elmenteitan Culture
|horizon =
|period = Mesolithic, Neolithic
|dates = around 3,300 to 1,300 BP
|typesite = Gamble's Cave
|majorsites = Gamble's Cave, Gogo Falls, Njoro River Cave
|extra =
|precededby = Kenya Stillbay, Highland Savanna PN
|followedby = Kenya Wilton
}}
The Elmenteitan culture was a prehistoric lithic industry and pottery tradition with a distinct pattern of land use, hunting and pastoral practice that appeared on the western plains of Kenya, East Africa during the Mesolithic.{{cite journal |url=https://www.jstor.org/stable/124525 |title= The Elmenteitan: An Early Food-Producing Culture in East Africa | publisher= Jstor |jstor= 124525 |accessdate=December 6, 2016|last1= Robertshaw |first1= Peter |journal= World Archaeology |year= 1988 |volume= 20 |issue= 1 |pages= 57–69 |doi= 10.1080/00438243.1988.9980056 }}
Since the late Neolithic continuous food-production, animal domestication and morturary rituals are documented. Progress and growth and continuity is related to wetter periods of increased precipitation, caused by pleistocene fluctuations and retreaing ice sheets.{{cite web |url=http://alexandriaarchive.org/bonecommons/archive/files/giffordgonzalez_icaz2010_0d1a567ef4.pdf |title= East African Pastoralism: Routes, Outcomes, Questions | publisher= Alexandria Archive |date= |accessdate=December 6, 2016}}{{cite web |url=http://alexandriaarchive.org/bonecommons/archive/files/giffordgonzalez_icaz2010_0d1a567ef4.pdf |title= East African Pastoralism: Routes, Outcomes, Questions | publisher= Alexandria Archive |date= |accessdate=December 6, 2016}}
It was named by Louis Leakey after Lake Elmenteita (also Elementaita), a soda lake, located in the Great Rift Valley, about {{convert|120|km|mi|0|abbr=on}} northwest of Nairobi.{{cite web |url=https://www.merriam-webster.com/dictionary/Elmenteitan |title= Definition of Elmenteitan | publisher= Merriam Webster Dictionary |date= |accessdate=December 6, 2016}}
Typical object and artifact assemblages include large double-edged obsidian microliths and blades, ceramic bowls and shallow stone vessels. Cattle and small stock was domesticated and and herded in combination with hunting, fishing and foraging. Patterns and degree of subsistence economy varied depending on the location and the local climate.{{cite web |url=http://www.archaeologywordsmith.com/lookup.php?terms=Elmenteitan |title= Elmenteitan | publisher= Archaeology Wordsmith |date= |accessdate=December 6, 2016}}{{cite web |url=http://erepository.uonbi.ac.ke/handle/11295/23464 |title= A contribution to the study of subsistence patterns of elmenteitan populations with reference to animal bones from Gogo falls in Sourh Nyanza District, Kenya | publisher= University of Nairobi Digital Archive |date= |accessdate=December 8, 2016}} Unlike earlier and contemporary cultures, such as the Highland Savanna PN, cremation of the dead took place in caves (Egerton Cave, Keringet Caves). The Njoro River Cave, first excavated in 1938 by Mary Leakey served as a mass-burial site. Associated finds include beads, blades, stone bowls, palettes and pottery vessels, that are interpreted as evidence for burial rituals.{{cite web |url=http://www.actforlibraries.org/archaeological-sites-njoro-river-cave/ |title= Archaeological Sites Njoro River Cave | publisher= Act For Libraries |date= |accessdate=December 6, 2016}}
The Elmenteitan was first described in 1931 by Louis Leakey who excavated at the Gamble's Cave and in 1938 at the Njoro River Cave. Leakey had noticed a locally distinct cluster of the lithic industry and a universal pottery tradition in a restricted area on the plains west of the central Great Rift Valley and at the Mau Escarpment.{{cite book |url=https://books.google.com/books?id=kJMFMpoHuVgC&dq=elmenteitan&pg=PA127 |title= The Archaeological and Linguistic Reconstruction of African History by Christopher Ehret, Merrick Posnansky |date= January 1982|isbn= 9780520045934 |accessdate=December 8, 2016|last1= Ehret |first1= Christopher |last2= Posnansky |first2= Merrick }} Many tool types are present that were common during the Aurignacian, but the Elmenteitan is distinguished by the high percentage of long symmetrical two-edged obsidian blades which were used unmodified and also served as blanks for a great variety of smaller tools.{{cite web |url=https://books.google.com/books?id=mtOhAwAAQBAJ&dq=elmenteitan&pg=PA369 |title= The Archaeology of Africa: Food, Metals and Towns y Bassey Andah, Alex Okpoko, Thurstan Shaw |date= |accessdate=December 8, 2016}}
All Elmenteitan excavation sites are associated with potsherds where pottery was fully developed.
Ceramic vessels are mainly undecorated. Several rare, but very distinctive ornamental designs such as irregular punctuations and rim millings were found. Occationally small bowls with outturned rims, handles with holes or horizontal lugs were discovered.
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style="background:#c5dfe1; color:#000;font-size:120%;"|Canadian Arctic Tundra |
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File:Auyuittuq National Park, Baffin Island, Nunavut -b.jpg Auyuittuq National Park, Baffin Island |
Area: {{convert|1420000|km2|sqmi|abbr=on}} |
climate : ET |
biome: Tundra |
elevation: max. {{convert|2500|m|ft|abbr=on}} |
coordinates: N {{coord|83|4|N|74|10|W|display=inline|type:landmark}}, S {{coord|62|22|N|79|38|W|display=inline|type:landmark}}, E {{coord|66|37|N|61|17|W|display=inline|type:landmark}}, W {{coord|72|2|N|125|37|W|display=inline|type:landmark}} |
Geology: |
style="text-align:center; "|File:Canada Köppen.svg Canadian tundra (light grey area) |
Country: Canada |
Protection: |
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The Canadian Arctic Tundra occupies the immediate regions north of the treeline in Canada's Arctic Lands. These territories encompass {{convert|2600000|km2|sqmi|abbr=on}}, 26 % of the country's landmass, including the Arctic Coastal Plains, Arctic Lowlands and the Innuitian Region of the High Arctic . Tundra accounts for approximately {{convert|1420000|km2|sqmi|abbr=on}} at the Yukon, the [[Northwest
Territories]], Nunavut, northeastern Manitoba, northern Ontario,
northern Quebec, northern Labrador and on 36,500 islands, of which Baffin Island with {{convert|422000|km2|sqmi|abbr=on}} is the largest.{{cite web |url=http://www.thecanadianencyclopedia.ca/en/article/physiographic-regions/ |title= Physiographic Regions | publisher= The Canadian Encyclopedia |date= |accessdate=December 9, 2016}}
Being part of the global tundra belt of the northern hemisphere Canada's tundra is characterized by extreme conditions with year round frozen grounds, long, cold winters, a very short growing season and precipitation lower than most deserts. The fact of low precipitation requires the classification of the tundra as a (cold) desert, yet permafrost causes even lower degrees of drainage and evaporation and as a result the ground, lakes and glaciers hold large quantities of fresh water.
Harsh environmental and weather conditions, slow growth, poor soils, lacking resources and nutrients, barly any variants of land forms support to highly adapted and migratory life forms only. Biodiversity is one dimensional and only sustainable for brief periods of appropriate conditions. Hundreds of species of flowering plants reproduce by budding and division rather than by interaction with insects.
{{cite web |url=http://www.ucmp.berkeley.edu/exhibits/biomes/tundra.php#arctic |title= The tundra biome | publisher= California Academy of Sciences |date= |accessdate=December 9, 2016}}{{cite web |url=http://wwf.panda.org/about_our_earth/ecoregions/canadian_lowarctic_tundra.cfm |title= Canadian Low Arctic Tundra - About the Area | publisher= World Wide Fund For Nature |date= |accessdate=December 9, 2016}}
Prolonged periods without sun light, frozen soils and strong winds prevent substantial tree growth.{{cite web |url=http://study.com/academy/lesson/tundra-biome-climate-locations-plants-animals.html |title= Tundra Biome: Climate, Locations, Plants & Animals
| publisher= Study.com |date= |accessdate=December 9, 2016}} Lichen has developed a unique and alternative adaptation strategy, that does not depend on soils for growth. It is one of the most notable, widespread and enduring organisms of the tundra. This symbiotic life form has algae or cyanobacteria live among the filaments of a fungus and benefit from moisture, elementary nutrients and are protected from the environment. The photobiont's (algae) photosynthetic processes in turn generates organic carbon sugars for the fungus.
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Western imperialism in Asia
{{New Imperialism}}
Western imperialism in Asia has its origins in European traders seeking unrestricted and independent trade with India and the Southeast Asian spice islands or the East Indies after the Ottoman empire had completely shut the Eurasian trade routes in the eastern Mediterranean by the late 15th century. This the Age of Discovery, and the introduction of early modern warfare into what was then called the Far East. By the early 16th century the Age of Sail greatly expanded Western European influence and development of the Spice Trade under colonialism. There has been a presence of Western European colonial empires and imperialism in Asia throughout six centuries of colonialism, formally ending with the independence of the Portuguese Empire's last colony East Timor in 2002. The empires introduced Western concepts of nation and the multinational state. This article attempts to outline the consequent development of the Western concept of the nation state.
The thrust of European political power, commerce, and culture in Asia gave rise to growing trade in commodities—a key development in the rise of today's modern world free market economy. In the 16th century, the Portuguese broke the (overland) monopoly of the Arabs and Italians of trade between Asia and Europe by the discovery of the sea route to India around the Cape of Good Hope.M. Weisner-Hanks, Early Modern Europe 1450–1789 (Cambridge, 2006) With the ensuing rise of the rival Dutch East India Company, Portuguese influence in Asia was gradually eclipsed.{{#tag:ref| For fifty or sixty years, the Portuguese enjoyed the exclusive trade to China and Japan. In 1717, and again in 1732, the Chinese government offered to make Macao the emporium for all foreign trade, and to receive all duties on imports; but, by a strange infatuation, the Portuguese government refused, and its decline is dated from that period.{{cite book |last=Roberts|first=Edmund |authorlink=Edmund Roberts (diplomat) |title=Embassy to the Eastern courts of Cochin-China, Siam, and Muscat : in the U. S. sloop-of-war Peacock ... during the years 1832-3-4 |url=https://books.google.com/books?id=aSgPAAAAYAAJ |date=Digitized 12 October 2007 |origyear=First published in 1837 |publisher=Harper & brothers |oclc=12212199 |at=image 173, p.166}}|group="nb"}} Dutch forces first established independent bases in the East (most significantly Batavia, the heavily fortified headquarters of the Dutch East India Company) and then between 1640 and 1660 wrestled Malacca, Ceylon, some southern Indian ports, and the lucrative Japan trade from the Portuguese. Later, the English and the French established settlements in India and established a trade with China and their own acquisitions would gradually surpass those of the Dutch. Following the end of the Seven Years' War in 1763, the British eliminated French influence in India and established the British East India Company as the most important political force on the Indian Subcontinent.
Before the Industrial Revolution in the mid-to-late 19th century, demand for oriental goods such as (porcelain, silk, spices and tea) remained the driving force behind European imperialism, and (with the important exception of British East India Company rule in India) the Western European stake in Asia remained confined largely to trading stations and strategic outposts necessary to protect trade. Industrialisation, however, dramatically increased European demand for Asian raw materials; and the severe Long Depression of the 1870s provoked a scramble for new markets for European industrial products and financial services in Africa, the Americas, Eastern Europe, and especially in Asia. This scramble coincided with a new era in global colonial expansion known as "the New Imperialism", which saw a shift in focus from trade and indirect rule to formal colonial control of vast overseas territories ruled as political extensions of their mother countries. Between the 1870s and the beginning of World War I in 1914, the United Kingdom, France, and the Netherlands—the established colonial powers in Asia—added to their empires vast expanses of territory in the Middle East, the Indian Subcontinent, and South East Asia. In the same period, the Empire of Japan, following the Meiji Restoration; the German Empire, following the end of the Franco-Prussian War in 1871; Tsarist Russia; and the United States, following the Spanish–American War in 1898, quickly emerged as new imperial powers in East Asia and in the Pacific Ocean area.
In Asia, World War I and World War II were played out as struggles among several key imperial powers—conflicts involving the European powers along with Russia and the rising American and Japanese powers. None of the colonial powers, however, possessed the resources to withstand the strains of both world wars and maintain their direct rule in Asia. Although nationalist movements throughout the colonial world led to the political independence of nearly all of the Asia's remaining colonies, decolonisation was intercepted by the Cold War; and South East Asia, South Asia, the Middle East, and East Asia remained embedded in a world economic, financial, and military system in which the great powers compete to extend their influence. However, the rapid post-war economic development of the East Asian Tigers, India, the People's Republic of China, along with the collapse of the Soviet Union, have loosened European and American influence in Asia, generating speculation today about emergence of modern India and China as potential superpowers.
References
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