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2021 in paleobotany

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{{Year nav topic20 |2021|paleobotany |paleontology |arthropod paleontology |paleoentomology |paleomalacology |reptile paleontology |archosaur paleontology |mammal paleontology |paleoichthyology }}

This article records new taxa of fossil plants described during the year 2021, as well as other significant discoveries and events related to paleobotany that occurred in 2021.

Ferns and fern allies

class="wikitable sortable" align="center" width="80%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Arthropitys buritiranensis{{Cite journal|last1=Neregato |first1=R. |last2=Rößler |first2=R. |last3=Noll |first3=R. |last4=Rohn |first4=R. |year=2021 |title=New petrified calamitaleans from the Permian of the Parnaíba Basin, central-north Brazil, part III, with some concerns regarding anatomical features of Paleozoic petrified sphenophytes |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104499 |doi=10.1016/j.revpalbo.2021.104499 |bibcode=2021RPaPa.29304499N |doi-access=free }}

|

Sp. nov

|

In press

|

Neregato et al.

|

Permian

|

Motuca Formation

|

{{Flag|Brazil}}

|

A member of Calamitales.

|

Azolla andreisii{{Cite journal|last1=De Benedetti |first1=F. |last2=Zamaloa |first2=M. C. |last3=Gandolfo |first3=M. A. |last4=Cúneo |first4=N. R. |year=2021 |title=Water fern spores (Salviniales) from the Late Cretaceous of Patagonia, Argentina |journal=Review of Palaeobotany and Palynology |volume=290 |pages=Article 104428 |doi=10.1016/j.revpalbo.2021.104428 |bibcode=2021RPaPa.29004428D |s2cid=234815591 |doi-access=free }}

|

Sp. nov

|

In press

|

De Benedetti et al.

|

Late Cretaceous (Maastrichtian)

|

La Colonia Formation

|

{{Flag|Argentina}}

|

A species of Azolla.

|

Caulopteris ellipticus{{Cite journal|last1=Wang |first1=S.-J. |last2=Wang |first2=J. |last3=Liu |first3=L. |last4=Hilton |first4=J. |year=2021 |title=Stem diversity of the marattialean tree fern family Psaroniaceae from the earliest Permian Wuda Tuff Flora |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104378 |doi=10.1016/j.revpalbo.2021.104378 |bibcode=2021RPaPa.29404378W |s2cid=234276315 }}

|

Sp. nov

|

In press

|

Wang et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A marattialean tree fern belonging to the family Psaroniaceae.

|

Caulopteris neimengensis

|

Sp. nov

|

In press

|

Wang et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A marattialean tree fern belonging to the family Psaroniaceae.

|

Caulopteris obovatus

|

Sp. nov

|

In press

|

Wang et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A marattialean tree fern belonging to the family Psaroniaceae.

|

Cicatricosisporites pseudograndiosus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the genus Ceratopteris.

|

Cladarastega{{Cite journal|last=Poinar |first=G. |title=A new fern, Cladarastega burmanica gen. et sp. nov. (Dennstaedtiaceae: Polypodiales) in mid-Cretaceous Burmese amber |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=153–160 |doi=10.18476/pale.v14.a7 |doi-access=free }}

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Gen. et sp. nov

|

Valid

|

Poinar

|

Late Cretaceous (Cenomanian)

|

Burmese amber

|

{{Flag|Myanmar}}

|

A fern belonging to the family Dennstaedtiaceae. Genus includes new species C. burmanica.

|

Claytosmunda zhangii{{Cite journal|last1=Tian |first1=N. |last2=Wang |first2=Y.-D. |last3=Jiang |first3=Z.-K. |year=2021 |title=A new permineralized osmundaceous rhizome with fungal remains from the Jurassic of western Liaoning, NE China |journal=Review of Palaeobotany and Palynology |volume=290 |pages=Article 104414 |doi=10.1016/j.revpalbo.2021.104414 |bibcode=2021RPaPa.29004414T | issn=0034-6667 |s2cid=233549860 }}

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Sp. nov

|

In press

|

Tian, Wang & Jiang

|

Late Jurassic

|

Tiaojishan Formation

|

{{Flag|China}}

|

A fern, a species of Claytosmunda.

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Dennstaedtia christophelii{{Cite journal|last1=Pigg |first1=K. B. |last2=DeVore |first2=M. L. |last3=Greenwood |first3=D. R. |last4=Sundue |first4=M. A. |last5=Schwartsburd |first5=P. |last6=Basinger |first6=J. F. |year=2021 |title=Fossil Dennstaedtiaceae and Hymenophyllaceae from the Early Eocene of the Pacific Northwest |journal=International Journal of Plant Sciences |volume=182 |issue=9 |pages=793–807 |doi=10.1086/715633 |s2cid=239036762 }}

|

Sp. nov

|

Valid

|

Pigg et al.

|

Early Eocene

|

Klondike Mountain Formation

|

{{Flag|United States}}
({{Flag|Washington}})

|

A fern, a species of Dennstaedtia.

|

File:Dennstaedtia christophelii holotype SRIC SR 13-004-001 A img1.tif]]

Echinosporis densiechinatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Marattiaceae.

|

Eoangiopteris congestus{{Cite journal|last1=Sun |first1=W. |last2=Li |first2=D. |last3=Zhou |first3=W. |last4=Bek |first4=J. |last5=Liu |first5=L. |last6=Wang |first6=J. |year=2021 |title=Eoangiopteris congestus sp. nov., a marattialean fern from the Lower Permian Taiyuan Formation of Wuda Coalfield, Inner Mongolia |journal=Acta Palaeontologica Sinica |volume=60 |issue=2 |pages=224–242 |doi=10.19800/j.cnki.aps.2020012 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210203 }}

|

Sp. nov

|

Valid

|

Sun et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A fern belonging to the group Marattiales.

|

Hymenophyllum axsmithii

|

Sp. nov

|

Valid

|

Pigg et al.

|

Early Eocene

|

Klondike Mountain Formation

|

{{Flag|United States}}
({{Flag|Washington}})

|

A fern, a species of Hymenophyllum.

|

Iberisetum{{Cite journal|last1=Correia |first1=P. |last2=Šimůnek |first2=Z. |last3=Sá |first3=A. A. |year=2021 |title=The equisetalean Iberisetum wegeneri gen. nov., sp. nov. from the Upper Pennsylvanian of Portugal |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3495–3505 |doi=10.1080/08912963.2021.1874373 |s2cid=234064743 }}

|

Gen. et sp. nov

|

In press

|

Correia, Šimůnek & Sá

|

Carboniferous (Gzhelian)

|

Douro Basin

|

{{Flag|Portugal}}

|

A member of Equisetales. Genus includes new species I. wegeneri.

|

Laevigatosporites cultellus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Polypodiaceae.

|

Marsileaceaephyllum ciliatum{{Cite journal|last1=Wang |first1=S. |last2=Long |first2=X. |last3=Zhang |first3=H. |last4=Cai |first4=H. |last5=Engel |first5=M. S. |last6=Shi |first6=C. |year=2021 |title=A semi-aquatic fern (Marsileaceae) from the mid-Cretaceous amber of northern Myanmar |journal=Cretaceous Research |volume=133 |pages=Article 105119 |doi=10.1016/j.cretres.2021.105119 |s2cid=245298780 }}

|

Sp. nov

|

In press

|

Wang et al.

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A member of the family Marsileaceae.

|

Nemejcopteris haiwangii{{Cite journal|last1=Pšenička |first1=J. |last2=Wang |first2=J. |last3=Bek |first3=J. |last4=Pfefferkorn |first4=H. W. |last5=Opluštil |first5=S. |last6=Zhou |first6=W. |last7=Frojdová |first7=J. |last8=Libertín |first8=M. |year=2021 |title=A zygopterid fern with fertile and vegetative parts in anatomical and compression preservation from the earliest Permian of Inner Mongolia, China |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104382 |doi=10.1016/j.revpalbo.2021.104382 |bibcode=2021RPaPa.29404382P |s2cid=234092362 }}

|

Sp. nov

|

In press

|

Pšenička et al.

|

Permian (Asselian)

|

Taiyuan Formation

|

{{Flag|China}}

|

A zygopterid fern.

|

Neocalamites iranensis{{Cite journal |last1=Mazaheri-Johari |first1=M. |last2=Kustatscher |first2=E. |last3=Roghi |first3=G. |last4=Ghasemi-Nejad |first4=E. |last5=Gianolla |first5=P. |year=2021 |title=A monotypic stand of Neocalamites iranensis n. sp. from the Carnian Pluvial Episode (Late Triassic) of the Aghdarband area, NE Iran (Turan Plate) |journal=Rivista Italiana di Paleontologia e Stratigrafia |volume=127 |issue=2 |pages=189–209 |doi=10.13130/2039-4942/15646 }}

|

Sp. nov

|

Valid

|

Kustatscher, Mazaheri-Johari & Roghi in Mazaheri-Johari et al.

|

Late Triassic (Carnian)

|

Miakuhi Formation

|

{{Flag|Iran}}

|

A member of the family Equisetaceae.

|

Odontosoria marekgaltieri{{Cite journal|last1=Pšenička |first1=J. |last2=Sakala |first2=J. |last3=Dašková |first3=J. |year=2021 |title=Odontosoria marekgaltieri sp. nov. (Lindsaeaceae), a new fern from the early Miocene of the Czech Republic: First evidence of the genus in the fossil record |journal=Review of Palaeobotany and Palynology |volume=297 |pages=Article 104580 |doi=10.1016/j.revpalbo.2021.104580 }}

|

Sp. nov

|

In press

|

Pšenička, Sakala & Dašková

|

Early Miocene

|

Most Basin

|

{{Flag|Czech Republic}}

|

A species of Odontosoria.

|

Oligosporangiopteris{{Cite journal|last1=Votočková Frojdová |first1=J. |last2=Wang |first2=J. |last3=Pšenička |first3=J. |last4=Bek |first4=J. |last5=Opluštil |first5=S. |last6=Libertín |first6=M. |year=2021 |title=A new leptosporangiate fern Oligosporangiopteris zhongxiangii gen. and sp. nov. from the lowermost Permian of Inner Mongolia, China – morphology, anatomy and reproductive organs |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104479 |doi=10.1016/j.revpalbo.2021.104479 |bibcode=2021RPaPa.29404479V |s2cid=237931032 }}

|

Gen. et sp. nov

|

In press

|

Votočková Frojdová et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A leptosporangiate fern. Genus includes new species O. zhongxiangii.

|

Osmunda zhangpuensis{{Cite journal|last1=Wang |first1=Z. |last2=Shi |first2=G. |last3=Sun |first3=B. |last4=Dong |first4=C. |last5=Yin |first5=S. |last6=Wu |first6=X. |year=2021 |title=A new species of Osmunda L. (Osmundaceae) from the middle Miocene of Fujian, Southeast China |journal=Acta Palaeontologica Sinica |volume=60 |issue=3 |pages=Article 2021045 |doi=10.19800/j.cnki.aps.2021045 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210306 }}

|

Sp. nov

|

|

Wang & Sun in Wang et al.

|

Miocene

|

Fotan Group

|

{{Flag|China}}

|

A fern, a species of Osmunda.

|

Patagoniapteris{{Cite journal|last1=Gnaedinger |first1=S. C. |last2=Zavattieri |first2=A. M. |title=A new Late Triassic dipteridacean fern from the Paso Flores Formation, Neuquén Basin, Argentina |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=4 |pages=885–900 |doi=10.4202/app.00864.2020 |doi-access=free |hdl=11336/171135 |hdl-access=free }}

|

Gen. et sp. nov

|

Valid

|

Gnaedinger & Zavattieri

|

Late Triassic (Norian–Rhaetian)

|

Paso Flores Formation

|

{{Flag|Argentina}}

|

A member of the family Dipteridaceae. Genus includes new species P. artabeae.

|

Pectinangium xuanweiense{{Cite journal|last1=Zhou |first1=Y. |last2=Guo |first2=Y. |last3=Pšenička |first3=J. |last4=Bek |first4=J. |last5=Yang |first5=S.-L. |last6=Feng |first6=Z. |year=2021 |title=A new marattialean fern, Pectinangium xuanweiense sp. nov., from the Lopingian of Southwest China |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104500 |doi=10.1016/j.revpalbo.2021.104500 |bibcode=2021RPaPa.29504500Z |doi-access=free }}

|

Sp. nov

|

In press

|

Zhou et al.

|

Permian (Lopingian)

|

|

{{Flag|China}}

|

A fern belonging to the group Marattiales.

|

Polypodiisporites densus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Polypodiaceae.

|

Polypodiisporites fossulatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Lomariopsidaceae.

|

Psilatriletes marginatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the genus Cyathea.

|

Qasimia yunnanica{{Cite journal|last1=Guo |first1=Y. |last2=Zhou |first2=Y. |last3=Bek |first3=J. |last4=Yang |first4=S.-L. |last5=Feng |first5=Z. |year=2021 |title=Qasimia yunnanica sp. nov., a marattialean fern with bivalvate synangia from the Lopingian of Southwest China |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104497 |doi=10.1016/j.revpalbo.2021.104497 |bibcode=2021RPaPa.29304497G |doi-access=free }}

|

Sp. nov

|

In press

|

Guo et al.

|

Permian (Lopingian)

|

Xuanwei Formation

|

{{Flag|China}}

|

A fern belonging to the group Marattiales.

|

Reticulosporis diversus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Polypodiaceae.

|

Rothwellopteris sanjiaoshuensis{{Cite journal|last1=He |first1=X.-Y. |last2=Hilton |first2=J. |last3=Wang |first3=S.-J. |last4=Cheng |first4=X.-S. |year=2021 |title=Exploring the stem to crown group transition in Marattiales: A new species of frond from the late Permian of China with features of the Psaroniaceae and Marattiaceae |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104506 |doi=10.1016/j.revpalbo.2021.104506 |bibcode=2021RPaPa.29504506H }}

|

Sp. nov

|

In press

|

He et al.

|

Late Permian

|

Xuanwei Formation

|

{{Flag|China}}

|

A fern belonging to the group Marattiales.

|

Sphenophyllum fanwanense{{Cite journal|last1=Huang |first1=P. |last2=Liu |first2=L. |last3=Liu |first3=L. |last4=Wang |first4=J.-S. |last5=Xue |first5=J.-Z. |title=Sphenophyllum Brongniart (Sphenopsida) from the Upper Devonian of South China |journal=Palaeoworld |year=2022 |volume=31 |issue=3 |pages=402–418 |doi=10.1016/j.palwor.2021.09.007 |s2cid=242991171 }}

|

Sp. nov

|

In press

|

Huang et al.

|

Late Devonian

|

|

{{Flag|China}}

|

A sphenophyllalean equisetid

|

Sphenophyllum parvifolium{{Cite journal|last1=Libertín |first1=M. |last2=Bek |first2=J. |last3=Wang |first3=J. |last4=Opluštil |first4=S. |last5=Pšenička |first5=J. |last6=Votočková Frojdová |first6=J. |year=2021 |title=New data about three sphenophylls and their spores from the volcanic tuff of Wuda, Taiyuan Formation, earliest Permian, China |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104484 |doi=10.1016/j.revpalbo.2021.104484 |bibcode=2021RPaPa.29404484L }}

|

Sp. nov

|

In press

|

Libertín et al.

|

Early Permian

|

Taiyuan Formation

|

{{Flag|China}}

|

A sphenophyllalean equisetid

|

Tapelrayen{{Cite journal|last1=Machado |first1=M. A. |last2=Vera |first2=E. I. |last3=Passalia |first3=M. G. |last4=Ponce |first4=M. M. |year=2021 |title=Eupolypod ferns with dryopteroid/thelypteroid traits from Arroyo Chacay (Huitrera Formation, Eocene), Río Negro Province, Argentina |journal=Review of Palaeobotany and Palynology |volume=287 |pages=Article 104381 |doi=10.1016/j.revpalbo.2021.104381 |bibcode=2021RPaPa.28704381M |s2cid=233532654 |url=http://rid.unrn.edu.ar/handle/20.500.12049/6970 }}

|

Gen. et sp. nov

|

In press

|

Machado et al.

|

Eocene

|

Huitrera Formation

|

{{Flag|Argentina}}

|

Fertile remains of a fern comparable with Thelypteridaceae and Dryopteridaceae. Genus includes new species T. helgae.

|

Thyrsopteris cyathindusia{{Cite journal|last1=Zhang |first1=H.-R. |last2=Shi |first2=C. |last3=Long |first3=X.-X. |last4=Feng |first4=Q. |last5=Cai |first5=H.-H. |last6=Lü |first6=Y.-T. |last7=Wang |first7=S. |title=A new fossil record of Thyrsopteridaceae (Cyatheales) from the mid-Cretaceous amber of Myanmar |journal=Palaeoworld |year=2022 |volume=31 |issue=3 |pages=478–484 |doi=10.1016/j.palwor.2021.09.002 |s2cid=240519538 }}

|

Sp. nov

|

In press

|

Zhang et al.

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A tree fern, a species of Thyrsopteris.

|

Woodwardia changchangensis{{Cite journal|last1=Song |first1=H.-Z. |last2=Naugolnykh |first2=S. V. |last3=Wu |first3=X.-K. |last4=Liu |first4=X.-Y. |last5=Jin |first5=J.-H. |title=Fertile Woodwardia from the middle Eocene of South China and its implications for palaeogeography and palaeoclimate |journal=Plant Diversity |year=2022 |volume=44 |issue=6 |pages=565–576 |doi=10.1016/j.pld.2021.09.003 |pmid=36540713 |pmc=9751083 |issn=2468-2659 |doi-access=free }}

|

Sp. nov

|

In press

|

Naugolnykh & Song in Song et al.

|

Middle Eocene

|

Changchang Formation

|

{{Flag|China}}

|

A fern, a species of Woodwardia.

|

Bennettitales

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Nilssoniopteris jogiana{{Cite journal|last1=Blomenkemper |first1=P. |last2=Bäumer |first2=R. |last3=Backer |first3=M. |last4=Abu Hamad |first4=A. |last5=Wang |first5=J. |last6=Kerp |first6=H. |last7=Bomfleur |first7=B. |year=2021 |title=Bennettitalean Leaves From the Permian of Equatorial Pangea—The Early Radiation of an Iconic Mesozoic Gymnosperm Group |journal=Frontiers in Earth Science |volume=9 |pages=Article 652699 |doi=10.3389/feart.2021.652699 |bibcode=2021FrEaS...9..162B |doi-access=free }}

|

Sp. nov

|

|

Blomenkemper & Abu Hamad in Blomenkemper et al.

|

Permian (Changhsingian)

|

Umm Irna Formation

|

{{Flag|Jordan}}

|

A member of Bennettitales.

|

Nilssoniopteris shanxiensis

|

Sp. nov

|

|

Bäumer, Backer & Wang in Blomenkemper et al.

|

Permian (Cisuralian)

|

Upper Shihhotse Formation

|

{{Flag|China}}

|

A member of Bennettitales.

|

Pterophyllum pottii

|

Sp. nov

|

|

Bomfleur & Kerp in Blomenkemper et al.

|

Permian (Changhsingian)

|

Umm Irna Formation

|

{{Flag|Jordan}}

|

A member of Bennettitales.

|

Weltrichia magna{{Cite journal|last1=Guzmán-Madrid |first1=D. S. |last2=Velasco de León |first2=M. P. |year=2021 |title=Weltrichia magna sp. nov., a new record for the Middle Jurassic of Oaxaca, México |journal=Acta Palaeobotanica |volume=61 |issue=1 |pages=95–106 |doi=10.35535/acpa-2021-0005 |s2cid=237760410 |doi-access=free }}

|

Sp. nov

|

Valid

|

Guzmán-Madrid & Velasco de León

|

Middle Jurassic (Bajocian)

|

Zorrillo Formation

|

{{Flag|Mexico}}

|

|

Weltrichia xochitetlii{{Cite journal|last1=Lozano-Carmona |first1=D. E. |last2=Corro-Ortiz |first2=M. G. |last3=Morales |first3=R. L. |last4=Velasco-de León |first4=M. P. |year=2021 |title=Weltrichia xochitetlii sp. nov. (Bennettitales) from the Middle Jurassic of northwestern Oaxaca, Mexico: First paleobotanical evidence from the Tecomazúchil formation |journal=Journal of South American Earth Sciences |volume=108 |pages=Article 103230 |doi=10.1016/j.jsames.2021.103230|issn=0895-9811 |bibcode=2021JSAES.10803230L |s2cid=234085434 }}

|

Sp. nov

|

In press

|

Lozano-Carmona et al.

|

Middle Jurassic (Callovian)

|

Tecomazuchil Formation

|

{{Flag|Mexico}}

|

A member of Bennettitales.

|

Williamsonia sanjuanensis{{Cite journal|last1=Lozano-Carmona |first1=D. E. |last2=Velasco-de León |first2=M. P. |year=2021 |title=Bennettitales from the Middle Jurassic of northwestern Oaxaca, Mexico: Diversity, sedimentary environments, and phytogeography |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103404 |doi=10.1016/j.jsames.2021.103404| issn=0895-9811 |bibcode=2021JSAES.11003404L }}

|

Sp. nov

|

In press

|

Lozano-Carmona & Velasco-de León

|

Middle Jurassic

|

|

{{Flag|Mexico}}

|

|

Cycadales

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Becklesia franconica{{Cite journal|last1=Van Konijnenburg-van Cittert |first1=J. H. A. |last2=Pott |first2=C. |last3=Schmeißner |first3=S. |last4=Dütsch |first4=G. |last5=Kustatscher |first5=E. |year=2021 |title=The Rhaetian flora of Wüstenwelsberg, Bavaria, Germany: Description of selected gymnosperms (Ginkgoales, Cycadales, Coniferales) together with an ecological assessment of the locally prevailing vegetation |journal=Review of Palaeobotany and Palynology |volume=288 |pages=Article 104398 |doi=10.1016/j.revpalbo.2021.104398 |bibcode=2021RPaPa.28804398V |s2cid=233918337 |doi-access=free }}

|

Sp. nov

|

In press

|

Van Konijnenburg-van Cittert et al.

|

Late Triassic (Rhaetian)

|

Exter Formation

|

{{Flag|Germany}}

|

A member of Cycadales of uncertain phylogenetic placement.

|

Iratinia{{Cite journal|last1=Spiekermann |first1=R. |last2=Jasper |first2=A. |last3=Siegloch |first3=A. M. |last4=Guerra-Sommer |first4=M. |last5=Uhl |first5=D. |year=2021 |title=Not a lycopsid but a cycad-like plant: Iratinia australis gen. nov. et sp. nov. from the Irati Formation, Kungurian of the Paraná Basin, Brazil |journal=Review of Palaeobotany and Palynology |volume=289 |pages=Article 104415 |doi=10.1016/j.revpalbo.2021.104415 |bibcode=2021RPaPa.28904415S |s2cid=233860955 }}

|

Gen. et sp. nov

|

In press

|

Spiekermann et al.

|

Permian (Kungurian)

|

Irati Formation

|

{{Flag|Brazil}}

|

A cycad-like plant. Genus includes new species I. australis.

|

Ginkgoales

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Eretmophyllum hamiensis{{Cite journal|last1=Tang |first1=D.-L. |last2=Wang |first2=Z.-E. |last3=Huang |first3=Y.-T. |last4=Ding |first4=H. |last5=Ding |first5=S.-T. |last6=Wu |first6=J.-Y. |title=A new species of Eretmophyllum (Ginkgoales) from the Middle Jurassic of Turpan-Hami Basin, Xinjiang, China |journal=Palaeoworld |year=2022 |volume=31 |issue=4 |pages=646–657 |doi=10.1016/j.palwor.2021.12.001 |s2cid=245302175 }}

|

Sp. nov

|

In press

|

Tang et al.

|

Middle Jurassic

|

Xishanyao Formation

|

{{Flag|China}}

|

|

Ginkgoites villardeseoanii{{Cite journal|last1=Andruchow-Colombo |first1=A. |last2=Gandolfo |first2=M. A. |last3=Cúneo |first3=N. R. |last4=Escapa |first4=I. H. |year=2021 |title=Ginkgoites villardeseoanii sp. nov., a ginkgophyte with insect damage from the Upper Cretaceous (Maastrichtian) Lefipán Formation (Chubut, Patagonia, Argentina) |journal=Cretaceous Research |volume=133 |pages=Article 105124 |doi=10.1016/j.cretres.2021.105124 |s2cid=245458671 |doi-access=free }}

|

Sp. nov

|

In press

|

Andruchow-Colombo et al.

|

Late Cretaceous (Maastrichtian)

|

Lefipán Formation

|

{{Flag|Argentina}}

|

|

Ginkgoxylon arcticum{{Cite journal|last1=Afonin |first1=M. |last2=Gromyko |first2=D. |title=First record of Ginkgoxylon (Ginkgoales) fossil wood in the Lower Cretaceous of the Arctic region |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104868 |doi=10.1016/j.cretres.2021.104868 |bibcode=2021CrRes.12504868A }}

|

Sp. nov

|

In press

|

Afonin & Gromyko

|

Early Cretaceous

|

|

{{Flag|Russia}}
({{Flag|Arkhangelsk Oblast}})

|

A member of Ginkgoales described on the basis of fossil wood.

|

Karkenia irkutensis{{Cite journal|last1=Nosova |first1=N. |last2=Crane |first2=P. R. |last3=Shi |first3=G. |year=2021 |title=Ovule-bearing structures of Karkenia Archangelsky, associated dispersed seeds and Sphenobaiera leaves from the Middle Jurassic of East Siberia, Russia |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104522 |doi=10.1016/j.revpalbo.2021.104522 |bibcode=2021RPaPa.29504522N |s2cid=239696262 }}

|

Sp. nov

|

In press

|

Nosova, Crane & Shi

|

Middle Jurassic (Aalenian)

|

Prisayan Formation

|

{{Flag|Russia}}

|

|

Vladimariales

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Pseudotorellia doludenkoae{{Cite journal|last1=Nosova |first1=N. V. |last2=Kostina |first2=E. I. |last3=Bugdaeva |first3=E. V. |year=2021 |title=Pseudotorellia Florin from the Upper Jurassic–Lower Cretaceous of the Bureya Basin, Russian Far East |journal=Stratigraphy and Geological Correlation |volume=29 |issue=4 |pages=434–449 |doi=10.1134/S0869593821040031 |bibcode=2021SGC....29..434N |doi-access=free }}

|

Sp. nov

|

Valid

|

Nosova, Kostina & Bugdaeva

|

Late Jurassic–Early Cretaceous (Oxfordian–Berriasian)

|

Dublikan Formation

Talyndzhan Formation

|

{{Flag|Russia}}
({{Flag|Khabarovsk Krai}})

|

|

Pseudotorellia irkutensis{{Cite journal|last=Nosova|first=N.|year=2021|title=Female reproductive structures of Umaltolepis Krassilov and associated leaves of Pseudotorellia Florin from the Middle Jurassic of East Siberia, Russia|journal=Review of Palaeobotany and Palynology|volume=289|pages=Article 104412|doi=10.1016/j.revpalbo.2021.104412|bibcode=2021RPaPa.28904412N |s2cid=233790779}}

|

Sp. nov

|

In press

|

Nosova

|

Middle Jurassic (Aalenian–Bajocian)

|

Prisayan Formation

|

{{Flag|Russia}}

|

A Vladimariales foliage species

|

Umaltolepis irkutensis

|

Sp. nov

|

In press

|

Nosova

|

Middle Jurassic (Aalenian–Bajocian)

|

Prisayan Formation

|

{{Flag|Russia}}

|

A Vladimariales reproducive structure species

|

Conifers

=Araucariaceae=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Agathoxylon mendezii{{Cite journal|last1=Del Fueyo |first1=G. M. |last2=Carrizo |first2=A. |last3=Poiré |first3=D. G. |last4=Lafuente Diaz |first4=M. A. |title=Recurrent volcanic activity recorded in araucarian wood from the Lower Cretaceous Springhill Formation, Patagonia, Argentina: Palaeoenvironmental interpretations |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=1 |pages=231–253 |doi=10.4202/app.00783.2020 |doi-access=free |hdl=11336/137848 |hdl-access=free }}

|

Sp. nov

|

Valid

|

Del Fueyo et al.

|

Early Cretaceous (Berriasian–Valanginian)

|

Springhill Formation

|

{{Flag|Argentina}}

|

An Araucariaceae fossil wood.

|

Agathoxylon santanensis{{Cite journal|last1=dos Santos |first1=Â. C. S. |last2=Siegloch |first2=A. M. |last3=Guerra-Sommer |first3=M. |last4=Degani-Schmidt |first4=I. |last5=Carvalho |first5=I. S. |year=2021 |title=Agathoxylon santanensis sp. nov. from the Aptian Crato fossil Lagerstätte, Santana Formation, Araripe Basin, Brazil |journal=Journal of South American Earth Sciences |volume=112, Part 2 |pages=Article 103633 |doi=10.1016/j.jsames.2021.103633 |bibcode=2021JSAES.11203633S |s2cid=244110901 }}

|

Sp. nov

|

In press

|

Dos Santos et al.

|

Early Cretaceous (Aptian)

|

Crato Formation

|

{{Flag|Brazil}}

|

|

Araucaria violetae{{Cite journal|last1=Batista |first1=M. E. P. |last2=Loiola |first2=M. I. B. |last3=Soares |first3=A. A. |last4=Mastroberti |first4=A. A. |last5=Sá |first5=A. A. |last6=Nascimento Jr. |first6=D. R. |last7=Silva Filho |first7=W. F. |last8=Kunzmann |first8=L. |year=2021 |title=New Insights into the Evolution of Mucilage Cells in Araucariaceae: Araucaria violetae sp. nov. from the Early Cretaceous Araripe Basin (Northeast Brazil) |journal=International Journal of Plant Sciences |volume=183 |issue=1 |pages=43–60 |doi=10.1086/717104 |s2cid=239548378 }}

|

Sp. nov

|

In press

|

Batista et al.

|

Early Cretaceous

|

|

{{Flag|Brazil}}

|

A species of Araucaria.

|

=Cheirolepidiaceae=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Brachyoxylon lalongense{{Cite journal|last1=Yang |first1=X.-J. |last2=Li |first2=J.-G. |title=A petrified wood Brachyoxylon from the Lower Cretaceous of Bangoin, Tibet (Xizang), Southwest China |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105064 |doi=10.1016/j.cretres.2021.105064 |s2cid=240429516 }}

|

Sp. nov

|

In press

|

Yang & Li

|

Early Cretaceous (Berriasian–Barremian)

|

Duoni Formation

|

{{Flag|China}}

|

|

Brachyoxylon patagonicum{{Cite journal|last1=Rombola |first1=C. F. |last2=Greppi |first2=C. D. |last3=Pujana |first3=R. R. |last4=García Massini |first4=J. L. |last5=Bellosi |first5=E. S. |last6=Marenssi |first6=S. A. |title=Brachyoxylon fossil woods with traumatic resin canals from the Upper Cretaceous Cerro Fortaleza Formation, southern Patagonia (Santa Cruz Province, Argentina) |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105065 |doi=10.1016/j.cretres.2021.105065 |hdl=11336/148807 |s2cid=239597933 |url=https://www.sciencedirect.com/science/article/abs/pii/S019566712100313X |hdl-access=free }}

|

Sp. nov

|

In press

|

Rombola et al.

|

Late Cretaceous

|

Cerro Fortaleza Formation

|

{{Flag|Argentina}}

|

Fossil wood of a member of the family Cheirolepidiaceae.

|

Watsoniocladus cunhae{{Cite journal|last1=Kvaček |first1=J. |last2=Mendes |first2=M. M. |year=2021 |title=A new Cheirolepidiaceae conifer Watsoniocladus cunhae sp. nov. from the Early Cretaceous (late Aptian–early Albian) of western Portugal |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104519 |doi=10.1016/j.revpalbo.2021.104519 |bibcode=2021RPaPa.29504519K }}

|

Sp. nov

|

In press

|

Kvaček & Mendes

|

Early Cretaceous (Aptian–Albian)

|

Almargem Formation

|

{{Flag|Portugal}}

|

A member of the family Cheirolepidiaceae.

|

=Cupressaceae=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Cupressinoxylon widdringtonioides{{Cite journal|last1=de Wit |first1=M. |last2=Bamford |first2=M. |title=Fossil wood from the Upper Cretaceous crater sediments of the Salpeterkop volcano, North West Province, South Africa |year=2021 |journal=South African Journal of Geology |volume=124 |issue=3 |pages=751–760 |doi=10.25131/sajg.124.0028 |bibcode=2021SAJG..124..751D }}

|

Sp. nov

|

Valid

|

De Wit & Bamford

|

Late Cretaceous

|

|

{{Flag|South Africa}}

|

Fossil wood of a member or a relative of the family Cupressaceae.

|

Fokienia tianpingensis{{Cite journal|last1=Wu |first1=X. |last2=Zhang |first2=H. |last3=Kodrul |first3=T. M. |last4=Maslova |first4=N. P. |last5=Jiang |first5=S. |last6=Yin |first6=Q. |last7=Quan |first7=C. |last8=Jin |first8=J. |title=First Fossil Fokienia (Cupressaceae) in South China and Its Palaeogeographic and Palaeoecological Implications |year=2021 |journal=Frontiers in Earth Science |volume=9 |pages=Article 709663 |doi=10.3389/feart.2021.709663 |bibcode=2021FrEaS...9..555W |doi-access=free }}

|

Sp. nov

|

Valid

|

Wu & Jin in Wu et al.

|

Miocene

|

Erzitang Formation

|

{{Flag|China}}

|

A species of Fokienia.

|

Nishidastrobus{{Cite journal|last1=Atkinson |first1=B. A. |last2=Contreras |first2=D. L. |last3=Stockey |first3=R. A. |last4=Rothwell |first4=G. W. |title=Ancient diversity and turnover of cunninghamioid conifers (Cupressaceae): two new genera from the Upper Cretaceous of Hokkaido, Japan |year=2021 |journal=Botany |volume=99 |issue=8 |pages=457–473 |doi=10.1139/cjb-2021-0005 |s2cid=237705866 }}

|

Gen. et sp. nov

|

Valid

|

Atkinson et al.

|

Late Cretaceous

|

|

{{Flag|Japan}}

|

A member of the family Cupressaceae belonging to the subfamily Cunninghamioideae. Genus includes new species N. japonicum.

|

Ohanastrobus

|

Gen. et sp. nov

|

Valid

|

Atkinson et al.

|

Late Cretaceous

|

|

{{Flag|Japan}}

|

A member of the family Cupressaceae belonging to the subfamily Cunninghamioideae. Genus includes new species O. hokkaidoensis.

|

Protaxodioxylon metangulense{{Cite journal|last1=Nhamutole |first1=N. |last2=Bamford |first2=M. |last3=Araújo |first3=R. |year=2021 |title=New species of Protaxodioxylon (conifer wood) from the Middle Permian of the Metangula Graben (Niassa Province, Mozambique) and their implications |journal=Journal of African Earth Sciences |volume=183 |pages=Article 104323 |doi=10.1016/j.jafrearsci.2021.104323 |bibcode=2021JAfES.18304323N }}

|

Sp. nov

|

In press

|

Nhamutole, Bamford & Araújo

|

Permian (late Capitanian)

|

K5 Formation

|

{{Flag|Mozambique}}

|

A member of the family Cupressaceae.

|

Protaxodioxylon verniersii

|

Sp. nov

|

In press

|

Nhamutole & Bamford in Nhamutole, Bamford & Araújo

|

Permian (late Capitanian)

|

K5 Formation

|

{{Flag|Mozambique}}

|

A member of the family Cupressaceae.

|

Thujopsoxylon

|

Gen. et sp. nov

|

Valid

|

Dolezych, LePage & Williams

|

Oligocene (Chattian)

|

Korlikov Formation

|

{{Flag|Russia}}
({{Flag|Tomsk Oblast}})

|

Genus includes new species T. schneiderianum.

|

=Pinaceae=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Lepidocasus{{Cite journal|last1=Herrera |first1=F. |last2=Shi |first2=G. |last3=Bickner |first3=M. A. |last4=Ichinnorov |first4=N. |last5=Leslie |first5=A. B. |last6=Crane |first6=P. R. |last7=Herendeen |first7=P. S. |year=2021 |title=Early Cretaceous abietoid Pinaceae from Mongolia and the history of seed scale shedding |journal=American Journal of Botany |volume=108 |issue=8 |pages=1483–1499 |doi=10.1002/ajb2.1713|issn=0002-9122 |pmid=34458982 |s2cid=239667187 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Herrera et al.

|

Early Cretaceous (Aptian–Albian)

|

|

{{Flag|Mongolia}}

|

A member of the family Pinaceae. Genus includes new species L. mellonae.

|

Nothotsuga sinogaia{{Cite journal|last1=Ding |first1=S.-T. |last2=Chen |first2=S.-Y. |last3=Ruan |first3=S.-C. |last4=Yang |first4=M. |last5=Han |first5=Y. |last6=Wang |first6=X.-H. |last7=Zhang |first7=T.-H. |last8=Sun |first8=B.-N. |year=2021 |title=First fossil record of Nothotsuga (Pinaceae) in China: implications for palaeobiogeography and palaeoecology |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3617–3624 |doi=10.1080/08912963.2021.1881781 |s2cid=233975517 }}

|

Sp. nov

|

In press

|

Ding et al.

|

Late Miocene

|

|

{{Flag|China}}

|

A species of Nothotsuga

|

Piceoxylon nikitinii{{Cite journal|last1=Dolezych |first1=M. |last2=LePage |first2=B. A. |last3=Williams |first3=C. J. |year=2021 |title=A Chattian-Aquitanian wood flora from the West Siberian Plain: Implications for regional palaeobiogeography |journal=Palaeontographica Abteilung B |volume=302 |issue=1–6 |pages=37–169 |doi=10.1127/palb/2021/0074 |bibcode=2021PalAB.302...37D |s2cid=237651777 }}

|

Sp. nov

|

Valid

|

Dolezych, LePage & Williams

|

Oligocene (Chattian)

|

Korlikov Formation

|

{{Flag|Russia}}
({{Flag|Tomsk Oblast}})

|

A Picea relative wood.

|

Pinus leptokrempfii{{Cite journal|last1=Zhang |first1=J.-W. |last2=Wang |first2=L. |last3=D'Rozario |first3=A. |last4=Liang |first4=X.-Q. |last5=Huang |first5=J. |last6=Zhou |first6=Z.-K. |year=2021 |title=Pinus leptokrempfii, an Oligocene Relative of the Flat-Needled Pine P. krempfii (Pinaceae) from China: Implications for Paleogeographic Origin |journal=International Journal of Plant Sciences |volume=182 |issue=5 |pages=389–400 |doi=10.1086/713957 |s2cid=233888471 }}

|

Sp. nov

|

Valid

|

Zhang et al.

|

Early Oligocene

|

|

{{Flag|China}}

|

A pine.

|

Pinus nongyaplongensis{{Cite journal|last1=Grote |first1=P. J. |last2=Srisuk |first2=P. |year=2021 |title=Fossil Pinus from the Cenozoic of Thailand |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104501 |doi=10.1016/j.revpalbo.2021.104501 |bibcode=2021RPaPa.29504501G |issn=0034-6667 }}

|

Sp. nov

|

In press

|

Grote in Grote & Srisuk

|

Oligocene-early Miocene

|

|

{{Flag|Thailand}}

|

A pine.

|

Pinus weichangensis{{Cite journal|last1=Li |first1=Y. |last2=Yi |first2=T.-M. |last3=Grote |first3=P. J. |last4=An |first4=P.-C. |last5=Zhu |first5=Y.-B. |last6=Zhang |first6=Z.-Y. |last7=Li |first7=C.-S. |year=2021 |title=A new species of Pinus (Pinaceae) from the Miocene of Weichang, Hebei Province, China and its evolutionary significance |journal=Historical Biology: An International Journal of Paleobiology |volume=34 |issue=5 |pages=885–896 |doi=10.1080/08912963.2021.1952197 |s2cid=237692568 }}

|

Sp. nov

|

In press

|

Li et al.

|

Early Miocene

|

|

{{Flag|China}}

|

A pine.

|

Schizolepidopsis ediae{{Cite journal|last1=Matsunaga |first1=K. K. S. |last2=Herendeen |first2=P. S. |last3=Herrera |first3=F. |last4=Ichinnorov |first4=N. |last5=Crane |first5=P. R. |last6=Shi |first6=G. |year=2021 |title=Ovulate Cones of Schizolepidopsis ediae sp. nov. Provide Insights into the Evolution of Pinaceae |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=490–507 |doi=10.1086/714281 |s2cid=235426888 |doi-access=free }}

|

Sp. nov

|

Valid

|

Matsunaga et al.

|

Early Cretaceous

|

Huolinhe Formation

Tevshiin Govi Formation

|

{{Flag|China}}

{{Flag|Mongolia}}

|

A member or a close relative of the family Pinaceae.

|

=Podocarpaceae=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Podocarpus yunnanensis{{Cite journal|last1=Wu |first1=J. |last2=Chen |first2=H. |last3=Ruan |first3=S. |last4=Yang |first4=M. |last5=Mo |first5=L. |last6=Ji |first6=B. |last7=Zhang |first7=J. |last8=Ding |first8=S. |year=2021 |title=Fossil leaves of Podocarpus subgenus Foliolatus (Podocarpaceae) from the Pliocene of southwestern China and biogeographic history of Podocarpus |journal=Review of Palaeobotany and Palynology |volume=287 |pages=Article 104380 |doi=10.1016/j.revpalbo.2021.104380 |bibcode=2021RPaPa.28704380W |s2cid=234282697 }}

|

Sp. nov

|

In press

|

Wu et al.

|

Early Pliocene

|

|

{{Flag|China}}

|

A species of Podocarpus.

|

Protophyllocladoxylon hilarioense{{Cite journal|last1=Vallejos Leiz |first1=L. |last2=Crisafulli |first2=A. |last3=Gnaedinger |first3=S. |title=New records of Late Triassic wood from Argentina and their biostratigraphic, paleoclimatic, and paleoecological implications |journal=Acta Palaeontologica Polonica |volume=67 |issue=2 |year=2022 |pages=329–340 |doi=10.4202/app.00939.2021 |doi-access=free }}

|

Sp. nov

|

In press

|

Vallejos Leiz, Crisafulli & Gnaedinger

|

Late Triassic (Norian–Rhaetian)

|

Hilario Formation

|

{{Flag|Argentina}}

|

A member of the family Podocarpaceae.

|

Protophyllocladoxylon yiwuense{{Cite journal|last1=Gou |first1=X.-D. |last2=Feng |first2=Z. |last3=Wei |first3=H.-B. |last4=Lv |first4=Y. |last5=Yang |first5=S.-L. |year=2021 |title=A new Protophyllocladoxylon stem from the Xishanyao Formation (Middle Jurassic) in the Santanghu Basin, Xinjiang, Northwest China |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104474 |doi=10.1016/j.revpalbo.2021.104474 |bibcode=2021RPaPa.29204474G |doi-access=free }}

|

Sp. nov

|

In press

|

Gou & Feng in Gou et al.

|

Middle Jurassic

|

Xishanyao Formation

|

{{Flag|China}}

|

A conifer of uncertain phylogenetic placement, possibly belonging or related to the family Podocarpaceae.

|

=Other conifers=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Cargalostrobus{{Cite journal|last=Gomankov |first=A. V. |year=2021 |title=On conifer fructifications from the Cargala Mines (the Kazanian of the Southern Fore-Urals) |journal=Lethaea Rossica |volume=23 |pages=21–31 |url=http://paleobot.ru/pdf/02_2021_23.pdf }}

|

Gen. et sp. nov

|

Valid

|

Gomankov

|

Permian

|

|

{{Flag|Russia}}
({{Flag|Orenburg Oblast}})

|

A member of Pinales belonging to the family Sashiniaceae. Genus includes new species C. demetrii.

|

Megaporoxylon sinensis{{Cite journal|last1=Wan |first1=M. |last2=Wang |first2=J. |last3=Shi |first3=T. |last4=Wang |first4=K. |last5=Tang |first5=P. |last6=Wang |first6=J. |year=2021 |title=Megaporoxylon sinensis sp. nov., a new coniferous trunk from the Upper Triassic of northern Bogda Mountains, northwestern China |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104536 |doi=10.1016/j.revpalbo.2021.104536 |bibcode=2021RPaPa.29504536W |s2cid=244206721 }}

|

Sp. nov

|

In press

|

Wan et al.

|

Late Triassic (Carnian–Norian)

|

Huangshanjie Formation

|

{{Flag|China}}

|

A coniferous trunk.

|

Taxus huolingolensis{{Cite journal|last1=Dong |first1=C. |last2=Shi |first2=G. |last3=Herrera |first3=F. |last4=Wang |first4=Y. |last5=Wang |first5=Z. |last6=Zhang |first6=B. |last7=Xu |first7=X. |last8=Herendeen |first8=P. S. |last9=Crane |first9=P. R. |year=2021 |title=Leaves of Taxus with cuticle micromorphology from the Early Cretaceous of eastern Inner Mongolia, Northeast China |journal=Review of Palaeobotany and Palynology |volume=298 |pages=Article 104588 |doi=10.1016/j.revpalbo.2021.104588 |s2cid=245558315 |doi-access=free }}

|

Sp. nov

|

In press

|

Dong et al.

|

Early Cretaceous

|

Huolinhe Formation

|

{{Flag|China}}

|

A species of Taxus.

|

Voltzia edithae{{Cite journal|last1=Forte |first1=G. |last2=Kustatscher |first2=E. |last3=Van Konijnenburg-van Cittert |first3=J. H. A. |year=2021 |title=Conifer diversity in the Middle Triassic: new data from the Fossillagerstätte Kühwiesenkopf/Monte Prà della Vacca (Pelsonian, Anisian) in the Dolomites (NE Italy) |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=445–467 |doi=10.1086/714280 |s2cid=233649930 }}

|

Sp. nov

|

Valid

|

Forte, Kustatscher & Van Konijnenburg-van Cittert

|

Middle Triassic (Anisian)

|

|

{{Flag|Italy}}

|

A member of Voltziales.

|

Xenoxylon utahense{{Cite journal|last1=Xie |first1=A. |last2=Gee |first2=C. T. |last3=Bennis |first3=M. B. |last4=Gray |first4=D. |last5=Sprinkel |first5=D. A. |year=2021 |title=A more southerly occurrence of Xenoxylon in North America: X. utahense Xie et Gee sp. nov. from the Upper Jurassic Morrison Formation in Utah, USA, and its paleobiogeographic and paleoclimatic significance |journal=Review of Palaeobotany and Palynology |volume=291 |pages=Article 104451 |doi=10.1016/j.revpalbo.2021.104451 |bibcode=2021RPaPa.29104451X |s2cid=236239787 }}

|

Sp. nov

|

In press

|

Xie & Gee in Xie et al.

|

Late Jurassic

|

Morrison Formation

|

{{Flag|United States}}
({{Flag|Utah}})

|

Fossil wood of a conifer.

|

Zhuotingoxylon{{Cite journal|last1=Wan |first1=M. |last2=Yang |first2=W. |last3=Wang |first3=K. |last4=Liu |first4=L. |last5=Wang |first5=J. |title=Zhuotingoxylon liaoi gen. et sp. nov., a silicified coniferous trunk from the Changhsingian (Permian) of southern Bogda Mountains, northwestern China |year=2021 |journal=Geological Journal |volume=56 |issue=12 |pages=6135–6150 |doi=10.1002/gj.4189 |s2cid=236276022 }}

|

Gen. et sp. nov

|

In press

|

Wan et al.

|

Permian (Changhsingian)

|

Guodikeng Formation

|

{{Flag|China}}

|

A silicified trunk with coniferous affinities. Genus includes new species Z. liaoi.

|

Flowering plants

=Basal angiosperms=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Allonymphaea{{Cite journal|last=Doweld |first=A. B. |year=2021 |title=Fossil Alloceltidoxylon, Allonymphaea, Arecocaryon, Paralnoxylon and Paranyssa and extant Komaroviopsis, Marcanodendron, and Papyrocactus (Magnoliophyta), new replacement generic names |journal=Phytotaxa |volume=524 |issue=2 |pages=92–98 |doi=10.11646/phytotaxa.524.2.3 |s2cid=243482734 }}

|

Nom. nov

|

Valid

|

Doweld

|

Eocene

|

|

{{Flag|Egypt}}

|

A replacement name for Thiebaudia Chandler (1954).

|

=Unplaced non-eudicots=

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Alcainea{{Cite journal|last1=Sender |first1=L. M. |last2=Doyle |first2=J. A. |last3=Upchurch |first3=G. R. |last4=Endress |first4=P. K. |last5=Villanueva-Amadoz |first5=U. |last6=Diez |first6=J. B. |year=2021 |title=Evidence on vegetative and inflorescence morphology of Chloranthaceae (Angiospermae) from the Early Cretaceous (middle–late Albian) of Spain |journal=Journal of Systematic Palaeontology |volume=18 |issue=24 |pages=2015–2042 |doi=10.1080/14772019.2021.1873434 |s2cid=232116303 |url=https://figshare.com/articles/dataset/Evidence_on_vegetative_and_inflorescence_morphology_of_Chloranthaceae_Angiospermae_from_the_Early_Cretaceous_middle_late_Albian_of_Spain/14138245 |url-access=subscription }}

|

Gen. et sp. nov

|

Valid

|

Sender et al.

|

Early Cretaceous (Albian)

|

Escucha Formation

|

{{Flag|Spain}}

|

A member of the family Chloranthaceae. Genus includes new species A. eklundiae.

|

Todziaphyllum

|

Gen. et sp. nov

|

Valid

|

Sender et al.

|

Early Cretaceous (Albian)

|

Escucha Formation

|

{{Flag|Spain}}

|

A member of the family Chloranthaceae. Genus includes new species T. elongatum.

|

=Magnoliids=

==Canellales==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Aristolochia macginitieana{{Cite journal|last1=Freitas |first1=J. |last2=Doweld |first2=A. B. |year=2021 |title=Aristolochia macginitieana (Aristolochiaceae), a replacement name for Aristolochia triangularis MacGinitie non Aristolochia triangularis Chamisso |journal=Phytotaxa |volume=500 |issue=1 |pages=59–60 |doi=10.11646/phytotaxa.500.1.11 |s2cid=236589284 }}

|

Nom. nov

|

Valid

|

Freitas & Doweld

|

Oligocene

|

|

{{Flag|United States}}
({{Flag|California}})

|

An Aristolochia species; a replacement name for Aristolochia triangularis MacGinitie (1937).

|

Cryptocaryoxylon grandoleaceum

|

Sp. nov

|

Valid

|

Akkemik

|

Middle Miocene

|

Kesmekaya Volcanics

|

{{Flag|Turkey}}

|

A member of the family Lauraceae.

|

Laurus elliptica{{Cite journal|last1=Winterscheid |first1=H. |last2=Kvaček |first2=Z. |year=2021 |title=Systematic-taxonomic revision of the flora from the late Oligocene Fossillagerstätte Rott near Bonn (Germany). Part 2: Magnoliidae: Basal angiosperms and magnoliids |journal=Palaeontographica Abteilung B |volume=303 |issue=4–6 |pages=119–155 |doi=10.1127/palb/2021/0077 |bibcode=2021PalAB.303..119W |s2cid=244053928 }}

|

Nom. nov

|

Valid

|

Winterscheid in Winterscheid & Kvaček

|

Oligocene

|

|

{{Flag|Germany}}

|

A species of Laurus; a replacement name for Laurus obovata Weber (1852).

|

Rosarioxylon{{Cite journal|last1=Cevallos-Ferriz |first1=S. R. S. |last2=Catharina |first2=A. S. |last3=Kneller |first3=B. |year=2021 |title=Cretaceous Lauraceae wood from El Rosario, Baja California, Mexico |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104478 |doi=10.1016/j.revpalbo.2021.104478 |bibcode=2021RPaPa.29204478C }}

|

Gen. et sp. nov

|

In press

|

Cevallos-Ferriz, Catharina & Kneller

|

Late Cretaceous (Campanian)

|

Rosario Formation

|

{{Flag|Mexico}}

|

A member of the family Lauraceae. Genus includes new species R. bajacaliforniensis.

|

Winteroxylon oleiferum{{Cite journal|last1=Brea |first1=M. |last2=Iglesias |first2=A. |last3=Wilf |first3=P. |last4=Moya |first4=E. |last5=Gandolfo |first5=M. A. |year=2021 |title=First South American Record of Winteroxylon, Eocene of Laguna del Hunco (Chubut, Patagonia, Argentina): New Link to Australasia and Malesia |journal=International Journal of Plant Sciences |volume=182 |issue=3 |pages=185–197 |doi=10.1086/712427 |issn=1058-5893 |s2cid=232050459 }}

|

Sp. nov

|

Valid

|

Brea et al.

|

Early Eocene

|

Huitrera Formation

|

{{Flag|Argentina}}

|

A member of the family Winteraceae.

|

=Monocots=

==Alismatid monocots==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Bognerospadix{{Cite journal|last1=Stockey |first1=R. A. |last2=Hoffman |first2=G. L. |last3=Rothwell |first3=G. W. |year=2021 |title=Fossil evidence for Paleocene diversification of Araceae: Bognerospadix gen. nov. and Orontiophyllum grandifolium comb. nov. |journal=American Journal of Botany |volume=108 |issue=8 |pages=1417–1440 |doi=10.1002/ajb2.1707 |pmid=34431509 |s2cid=237292226 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Stockey, Hoffman & Rothwell

|

Paleocene

|

|

{{Flag|Canada}}
({{Flag|Alberta}})

|

A member of the family Araceae. Genus includes new species B. speirsiae.

|

==Lilioid monocots==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Mirafloris{{Cite journal|last=Poinar |first=G. O. |title=A Monocot Flower, Mirafloris burmitis gen. et sp. nov. (Monocots: Angiospermae), in Burmese Amber |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=3 |pages=342–348 |doi=10.37819/biosis.002.03.0126 |s2cid=238732295 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Poinar

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A member of the family Liliaceae. Genus includes new species M. burmitis.

|

==Commelinid monocots==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Arecipites invaginatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Arecaceae.

|

Arecocaryon

|

Nom. nov

|

Valid

|

Doweld

|

Eocene

|

Messel pit

|

{{Flag|Germany}}

|

A member of the family Arecaceae; a replacement name for Friedemannia Collinson, Manchester & Wilde (2012).

|

Cyperaceaepollis wesselinghii

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Cyperaceae.

|

Eograminis{{Cite journal|last1=Poinar |first1=G. |last2=Soreng |first2=R. J. |year=2021 |title=A New Genus and Species of Grass, Eograminis balticus (Poaceae: Arundinoideae) in Baltic Amber |journal=International Journal of Plant Sciences |volume=182 |issue=9 |pages=808–816 |doi=10.1086/716781 |s2cid=239036777 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Poinar & Soreng

|

Eocene

|

Baltic amber

|

{{Flag|Russia}}
({{Flag|Kaliningrad Oblast}})

|

Grass belonging to the subfamily Arundinoideae. Genus includes new species E. balticus.

|

Luminidites amazonicus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Arecaceae.

|

Orthogonospermum{{Cite journal|last1=Smith |first1=S. Y. |last2=Kapgate |first2=D. K. |last3=Robinson |first3=S. |last4=Srivastava |first4=R. |last5=Benedict |first5=J. C. |last6=Manchester |first6=S. R. |year=2021 |title=Fossil fruits and seeds of Zingiberales from the Late Cretaceous–early Cenozoic Deccan Intertrappean Beds of India |journal=International Journal of Plant Sciences |volume=182 |issue=2 |pages=91–108 |doi=10.1086/711474 |s2cid=231875495 }}

|

Gen. et sp. nov

|

Valid

|

Smith et al.

|

Late Cretaceous (Maastrichtian)

|

Deccan Intertrappean Beds

|

{{Flag|India}}

|

A member of the family Zingiberaceae. Genus includes new species O. patanense.

|

Sabalites colaniae{{Cite journal|last1=Song |first1=A. |last2=Liu |first2=J. |last3=Liang |first3=S.-Q. |last4=Do |first4=T. V. |last5=Nguyen |first5=H. B. |last6=Deng |first6=W.-Y.-D. |last7=Jia |first7=L.-B. |last8=Del Rio |first8=C. |last9=Srivastava |first9=G. |last10=Feng |first10=Z. |last11=Zhou |first11=Z.-K. |last12=Huang |first12=J. |last13=Su |first13=T. |title=Leaf fossils of Sabalites (Arecaceae) from the Oligocene of northern Vietnam and their paleoclimatic implications |journal=Plant Diversity |year=2022 |volume=44 |issue=4 |pages=406–416 |doi=10.1016/j.pld.2021.08.003 |pmid=35967257 |pmc=9363516 |doi-access=free }}

|

Sp. nov

|

In press

|

Song, Su, Do & Zhou in Song et al.

|

Oligocene

|

Dong Ho Formation

|

{{Flag|Vietnam}}

|

A member of the family Arecaceae belonging to the subfamily Coryphoideae.

|

Trichotomosulcites normalis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Arecaceae.

|

===Commelinid monocot research===

  • A study on the evolutionary history of palms throughout the Cenozoic era, aiming to determine the impact of Cenozoic environmental changes on the diversification and biogeography of palms, is published by Lim et al. (2021).{{Cite journal|last1=Lim |first1=J. Y. |last2=Huang |first2=H. |last3=Farnsworth |first3=A. |last4=Lunt |first4=D. J. |last5=Baker |first5=W. J. |last6=Morley |first6=R. J. |last7=Kissling |first7=W. D. |last8=Hoorn |first8=C. |year=2021 |title=The Cenozoic history of palms: Global diversification, biogeography and the decline of megathermal forests |journal=Global Ecology and Biogeography |volume=31 |issue=3 |pages=425–439 |doi=10.1111/geb.13436 |hdl=1983/fa5f36f6-5dbe-4109-bcaa-f9838a88e337 |s2cid=245284265 |url=https://research-information.bris.ac.uk/ws/files/308458806/Full_text_PDF_final_published_version_.pdf }}
  • Pollens of member of the family Poaceae preserving the same morphological characteristics as that of modern cereal grains are described from a sedimentary core from Lake Acıgöl (Turkey) by Andrieu-Ponel et al. (2021), who interpret this finding as indicative of the presence of proto-cereals in Anatolia since 2.3 million years ago, likely evolving from wild Poaceae as a result of trampling, nitrogen enrichment of soils and browsing by large mammal herds, and evaluate possible benefits from the availability of these proto-cereals for early hominins.{{Cite journal|last1=Andrieu-Ponel |first1=V. |last2=Rochette |first2=P. |last3=Demory |first3=F. |last4=Alçiçek |first4=H. |last5=Boulbes |first5=N. |last6=Bourlès |first6=D. |last7=Helvacı |first7=C. |last8=Lebatard |first8=A.-E. |last9=Mayda |first9=S. |last10=Michaud |first10=H. |last11=Moigne |first11=A.-M. |last12=Nomade |first12=S. |last13=Perrin |first13=M. |last14=Ponel |first14=P. |last15=Rambeau |first15=C. |last16=Vialet |first16=A. |last17=Gambin |first17=B. |last18=Alçiçek |first18=M. C. |title=Continuous presence of proto-cereals in Anatolia since 2.3 Ma, and their possible co-evolution with large herbivores and hominins |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 8914 |doi=10.1038/s41598-021-86423-8 |pmid=33903602 |pmc=8076274 |bibcode=2021NatSR..11.8914A }}

=Basal eudicots=

== Proteales ==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Platanocarpelia{{Cite journal|last1=Maslova |first1=N. P. |last2=Kodrul |first2=T. M. |last3=Kachkina |first3=V. V. |title=Leaves of Ettingshausenia cuneifolia (Bronn) Stiehler (Angiospermae) and Associated Carpels and Stamens from the Turonian of Southern Kazakhstan |year=2021 |journal=Paleontological Journal |volume=55 |issue=10 |pages=1193–1214 |doi=10.1134/S0031030121100063 |s2cid=245540003 }}

|

Gen. et sp. nov

|

Valid

|

Maslova, Kodrul & Kachkina

|

Late Cretaceous (Turonian)

|

|

{{Flag|Kazakhstan}}

|

A member of the family Platanaceae. Genus includes new species P. kyzyljarica.

|

Proteacidites pseudodehaanii

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Proteaceae.

|

=Superasterids=

==Campanulid euasterids==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Pittosporum ettingshausenii{{Cite journal|last=Doweld |first=A. B. |year=2021 |title=New names in Pittosporum, extant and fossil (Pittosporaceae) |journal=Phytotaxa |volume=498 |issue=4 |pages=298–300 |doi=10.11646/phytotaxa.498.4.9 |s2cid=235540500 }}

|

Nom. nov

|

Valid

|

Doweld

|

Miocene

|

|

{{Flag|New Zealand}}

|

A species of Pittosporum; a replacement name for Pittosporum elegans (Ettingshausen) W.R.B. Oliver (1950).

|

Xenopanax

|

Gen. et comb. nov

|

Valid

|

Doweld

|

Eocene

|

|

{{Flag|Russia}}
({{Flag|Kamchatka Krai}})

|

A new genus for "Pittosporum" beringianum Chelebaeva & Akhmetiev (1983).

|

==Lamiid euasterids==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Adina vastanenesis{{Cite journal|last1=Shukla |first1=A. |last2=Mehrotra |first2=R. C. |last3=Verma |first3=P. |last4=Chandra |first4=K. |last5=Singh |first5=A. |year=2021 |title="Out-of-India" dispersal for Adina (tribe Naucleeae; family Rubiaceae): evidence from the early Eocene fossil record from India |journal=Palaeoworld |volume=30 |issue=4 |pages=737–745 |doi=10.1016/j.palwor.2021.01.001 |s2cid=234244901 }}

|

Sp. nov

|

Valid

|

Shukla et al.

|

Early Eocene

|

Cambay Shale Formation

|

{{Flag|India}}

|

A species of Adina.

|

Dicolpopollis? costatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Macoubea.

|

Dolichandra pacei{{Cite journal|last1=Franco |first1=M. J. |last2=Brea |first2=M. |last3=Cerdeño |first3=E. |year=2021 |title=First Bignoniaceae liana from the Miocene of South America and its evolutionary significance |journal=American Journal of Botany |volume=108 |issue=9 |pages=1761–1774 |doi=10.1002/ajb2.1736 |pmid=34591314 |s2cid=238230695 |doi-access=free |hdl=11336/184023 |hdl-access=free }}

|

Sp. nov

|

Valid

|

Franco, Brea & Cerdeño

|

Miocene (Santacrucian)

|

Mariño Formation

|

{{Flag|Argentina}}

|

A species of Dolichandra.

|

Fraxinoxylon beypazariense

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Oleaceae.

|

Fraxinus eoemarginata{{Cite journal|last1=Mathewes |first1=R. |last2=Archibald |first2=S. B. |last3=Lundgren |first3=A. |year=2021 |title=Tips and identification of early Eocene Fraxinus L. samaras from the Quilchena locality, Okanagan Highlands, British Columbia, Canada |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104480 |doi=10.1016/j.revpalbo.2021.104480 |bibcode=2021RPaPa.29304480M }}

|

Sp. nov

|

In press

|

Mathewes, Archibald & Lundgren

|

Early Eocene

|

Quilchena site

|

{{Flag|Canada}}
({{Flag|British Columbia}})

|

A species of Fraxinus.

|

Kapgateophyllum{{Cite journal|last=Deshmukh |first=U. B. |year=2021 |title=Kapgateophyllum gen. nov. (Acanthaceae) |journal=Phytotaxa |volume=500 |issue=2 |pages=147–148 |doi=10.11646/phytotaxa.500.2.8 |s2cid=236585198 }}

|

Nom. nov

|

Valid

|

Deshmukh

|

Late Cretaceous (Maastrichtian) - early Eocene

|

Deccan Intertrappean Beds

|

{{Flag|India}}

|

A member of the family Acanthaceae; a replacement name for Acanthophyllum Ramteke & Kapgate (2014).

|

Ladakhipollenites campbellii

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Vitex.

|

Lymingtonia splendida

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Convolvulaceae.

|

Maryendressantha{{Cite journal|last1=Singh |first1=H. |last2=Judd |first2=W. S. |last3=Samant |first3=B. |last4=Agnihotri |first4=P. |last5=Grimaldi |first5=D. A. |last6=Manchester |first6=S. R. |year=2021 |title=Flowers of Apocynaceae in amber from the early Eocene of India |journal=American Journal of Botany |volume=108 |issue=5 |pages=883–892 |doi=10.1002/ajb2.1651 | issn=0002-9122 |pmid=34018178 |s2cid=235073745 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Singh et al.

|

Early Eocene

|

Cambay amber

|

{{Flag|India}}

|

A member of the family Apocynaceae. Genus includes new species M. succinifera.

|

Perfotricolpites hexacolpatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Merremia.

|

Verrustephanoporites intraverrucosus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Apocynaceae.

|

==Non euasterids==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Anacolosidites reticulatus

|

Sp. nov

|

In press

|

Morley, Huang & Hoorn in Huang et al.

|

Middle and late Eocene

|

Yaw Formation

|

|

Pollen probably derived from plants belonging to the genus Ptychopetalum.

|

Camptotheca manchesterii{{Cite journal|last1=Xie |first1=S.-P. |last2=Zhang |first2=S.-H. |last3=McElwain |first3=J. C. |last4=Zhang |first4=P. |last5=Wang |first5=B. |last6=Zhang |first6=Y. |last7=Yang |first7=Y.-H. |last8=Chen |first8=J.-Y. |year=2021 |title=First occurrence of Camptotheca fruits from late Miocene of southwestern China |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3625–3632 |doi=10.1080/08912963.2021.1881782 |s2cid=234020768 }}

|

Sp. nov

|

In press

|

Xie et al.

|

Late Miocene

|

Bangmai Formation

|

{{Flag|China}}

|

A species of Camptotheca.

|

Diospyros christensenii{{Cite journal|last1=Denk |first1=T. |last2=Bouchal |first2=J. M. |year=2021 |title=Dispersed pollen and calyx remains of Diospyros (Ebenaceae) from the middle Miocene "Plant beds" of Søby, Denmark |journal=GFF |volume=143 |issue=2–3 |pages=292–304 |doi=10.1080/11035897.2021.1907443 |bibcode=2021GFF...143..292D |s2cid=237648462 |url=http://urn.kb.se/resolve?urn=urn:nbn:se:nrm:diva-4502 }}

|

Sp. nov

|

Valid

|

Denk & Bouchal

|

Miocene

|

|

{{Flag|Denmark}}

|

A species of Diospyros.

|

Halesia mosbruggeri{{Cite journal|last=Kvaček |first=Z. |year=2021 |title=Halesia mosbruggeri Kvaček, sp. nov., a new fossil fruit of Halesia L. (Styracaceae) from the Bohemian Miocene (Czech Republic) |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=75–78 |doi=10.1007/s12549-020-00463-y |s2cid=232127283 }}

|

Sp. nov

|

Valid

|

Kvaček

|

Early Miocene

|

Most Basin

|

{{Flag|Czech Republic}}

|

A species of Halesia.

|

Loranthacites tabatingensis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Struthanthus.

|

Mecsekispermum{{Cite journal|last1=Erdei |first1=B. |last2=Hably |first2=L. |year=2021 |title=Fossil Gordonia (s.l.)–like (Theaceae) winged seeds from the early Miocene of the Mecsek Mts, W Hungary |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=59–67 |doi=10.1007/s12549-020-00461-0 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Hably & Erdei

|

Miocene (Burdigalian)

|

Feked Formation

|

{{Flag|Hungary}}

|

Possibly a member of the family Theaceae. Genus includes new species M. gordonioides.

|

Miranthus{{Cite journal|last1=Friis |first1=E. M. |last2=Crane |first2=P. R. |last3=Pedersen |first3=K. R. |title=Early flowers of primuloid Ericales from the Late Cretaceous of Portugal and their ecological and phytogeographic implications |year=2021 |journal=Fossil Imprint |volume=77 |issue=2 |pages=214–230 |doi=10.37520/fi.2021.016 |s2cid=245553031 |url=http://fi.nm.cz/en/clanek/early-flowers-of-primuloid-ericales-from-the-late-cretaceous-of-portugal-and-their-ecological-and-phytogeographic-implications-2/ |doi-access=free }}

|

Gen. et 2 sp. nov

|

Valid

|

Friis, Crane

|

Late Cretaceous (Campanian-Maastrichtian)

|

|

{{Flag|Portugal}}

|

A Primulaceae genus. Genus includes new species M. elegans and M. kvacekii.

|

Nyssa nanningensis{{Cite journal|last1=Xu |first1=S.-L. |last2=Kodrul |first2=T. M. |last3=Maslova |first3=N. P. |last4=Song |first4=H.-Z. |last5=Tobias |first5=A. V. |last6=Wu |first6=X.-K. |last7=Quan |first7=C. |last8=Jin |first8=J.-H. |year=2021 |title=First occurrence of Nyssa endocarps and associated fungi in the Oligocene of South China: palaeogeographical and palaeoecological significance |journal=Papers in Palaeontology |volume=8 |doi=10.1002/spp2.1408 |s2cid=244057027 }}

|

Sp. nov

|

In press

|

Xu & Jin in Xu et al.

|

Late Oligocene

|

Yongning Formation

|

{{Flag|China}}

|

A tupelo.

|

Paranyssa

|

Nom. nov

|

Valid

|

Doweld

|

Paleocene

|

|

{{Flag|United States}}
({{Flag|Montana}})

|

A member of the family Nyssaceae; a replacement name for Browniea Manchester & Hickey (2007).

|

Parsonsidites? minibrenacii

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Amaranthaceae.

|

Ternstroemites klettwitzensis{{Cite journal|last=Striegler |first=U. |title=New leaf species from the upper Miocene flora of the leaf-bearing Wischgrund clay (Lower Lusatia, Brandenburg, Germany) |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=102–110 |doi=10.37520/fi.2021.009 |s2cid=245028668 |url=http://fi.nm.cz/en/clanek/new-leaf-species-from-the-upper-miocene-flora-of-the-leaf-bearing-wischgrund-clay-lower-lusatia-brandenburg-germany-2/ |doi-access=free }}

|

Sp. nov

|

Valid

|

Striegler

|

Miocene (Tortonian)

|

Rauno Formation

|

{{Flag|Germany}}

|

A member of the family Theaceae.

|

=Superrosids=

==Fabids==

===Fabales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Albizia mahuadanrensis{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Spicer |first3=R. A. |last4=Spicer |first4=T. E. V. |last5=Mahato |first5=S. |last6=Bera |first6=S. |last7=Kumar |first7=S. |last8=Khan |first8=M. A. |year=2021 |title=Pliocene Albizia (Fabaceae) from Jharkhand, eastern India: reappraisal of its biogeography during Cenozoic in Southeast Asia |journal=Palaeoworld |volume=31 |pages=153–168 |doi=10.1016/j.palwor.2021.03.004 |s2cid=233521737 |url=http://oro.open.ac.uk/75930/1/1-s2.0-S1871174X21000263-main.pdf }}

|

Sp. nov

|

In press

|

Hazra, Hazra & Khan in Hazra et al.

|

Pliocene

|

Rajdanda Formation

|

{{Flag|India}}

|

A species of Albizia.

|

Albizia palaeoprocera

|

Sp. nov

|

In press

|

Hazra, Hazra & Khan in Hazra et al.

|

Pliocene

|

Rajdanda Formation

|

{{Flag|India}}

|

A species of Albizia.

|

Cercis zhangpuensis{{Cite journal|last1=Wang |first1=Z. |last2=Shi |first2=G. |last3=Sun |first3=B. |last4=Jia |first4=H. |last5=Dong |first5=C. |last6=Yin |first6=S. |last7=Wu |first7=X. |year=2021 |title=A new Cercis (Leguminosae) from the middle Miocene of Fujian, China |journal=Historical Biology: An International Journal of Paleobiology |volume=34 |pages=94–101 |doi=10.1080/08912963.2021.1900170 |s2cid=233645060 }}

|

Sp. nov

|

In press

|

Wang et al.

|

Miocene

|

Fotan Group

|

{{Flag|China}}

|

A species of Cercis.

|

Cladrastis haominiae{{Cite journal|last1=Jia |first1=L.-B. |last2=Huang |first2=J. |last3=Su |first3=T. |last4=Spicer |first4=R. A. |last5=Zhang |first5=S.-T. |last6=Li |first6=S.-F. |last7=Pan |first7=B. |last8=Nam |first8=G.-S. |last9=Huang |first9=Y.-J. |last10=Zhou |first10=Z.-K. |year=2021 |title=Fossil infructescence from southwestern China reveals Paleogene establishment of Cladrastis in Asia |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104456 |doi=10.1016/j.revpalbo.2021.104456 |bibcode=2021RPaPa.29204456J |s2cid=236255120 |url=http://oro.open.ac.uk/76470/1/1-s2.0-S0034666721000804-main.pdf }}

|

Sp. nov

|

In press

|

Jia et al.

|

Paleogene

|

|

{{Flag|China}}

|

A species of Cladrastis.

|

Enterolobiumoxylon{{Cite journal|last1=Pérez-Lara |first1=D. K. |last2=Estrada-Ruiz |first2=E. |last3=Castañeda-Posadas |first3=C. |year=2021 |title=Kingiodendron and Enterolobium Eocene woods from the El Bosque formation, Chiapas, Mexico |journal=Journal of South American Earth Sciences |volume=111 |pages=Article 103477 |doi=10.1016/j.jsames.2021.103477 |bibcode=2021JSAES.11103477P |s2cid=237819942 }}

|

Gen. et sp. nov

|

In press

|

Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas

|

Eocene

|

El Bosque Formation

|

{{Flag|Mexico}}

|

Fossil wood of a member of the family Fabaceae. Genus includes new species E. triserial.

|

Gleditsioxylon fiambalense{{Cite journal|last=Baez |first=J. |year=2021 |title=First fossil record in Tambería formation (Neogene) in Bolsón de Fiambalá, Catamarca province, Argentina: Palaeoenvironmental inferences through Leguminosae woods |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103403 |doi=10.1016/j.jsames.2021.103403 |bibcode=2021JSAES.11003403B }}

|

Sp. nov

|

In press

|

Baez

|

Miocene

|

Tambería Formation

|

{{Flag|Argentina}}

|

A member of Leguminosae.

|

Kingiodendron mexicanus

|

Sp. nov

|

In press

|

Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas

|

Eocene

|

El Bosque Formation

|

{{Flag|Mexico}}

|

Fossil wood of a member of the family Fabaceae.

|

Ladakhipollenites? pseudocolpiconstrictus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Fabaceae.

|

Leguminocarpum lottii{{Cite journal|last1=Li |first1=X. |last2=Manchester |first2=S. R. |last3=Correa-Narvaez |first3=J. E. |last4=Herendeen |first4=P. S. |year=2021 |title=An Extinct Fruit Species of Fabaceae from the Early Eocene of Northwestern Wyoming, USA |journal=International Journal of Plant Sciences |volume=182 |issue=8 |pages=730–746 |doi=10.1086/715634 |s2cid=237505717 }}

|

Sp. nov

|

Valid

|

Li & Manchester in Li et al.

|

Early Eocene

|

Tepee Trail Formation

|

{{Flag|United States}}
({{Flag|Wyoming}})

|

A member of the family Fabaceae.

|

Leguminocarpum olmensis{{Cite journal|last1=Centeno-González |first1=N. K. |last2=Martínez-Cabrera |first2=H. I. |last3=Porras-Múzquiz |first3=H. |last4=Estrada-Ruiz |first4=E. |title=Late Campanian fossil of a legume fruit supports Mexico as a center of Fabaceae radiation |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 41 |doi=10.1038/s42003-020-01533-9 |pmid=33446929 |pmc=7809014 }}

|

Sp. nov

|

Valid

|

Centeno-González et al.

|

Late Cretaceous (Campanian)

|

Olmos Formation

|

{{Flag|Mexico}}

|

A member of the family Fabaceae.

|

Neopapilionia{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Bera |first3=S. |last4=Khan |first4=M. A. |year=2021 |title=First Fossil Legume Flower of Papilionoid Affinity from India |journal=Journal of the Geological Society of India |volume=97 |issue=3 |pages=267–270 |doi=10.1007/s12594-021-1677-3 |issn=0016-7622 |s2cid=232164301 }}

|

Gen. et sp. nov

|

Valid

|

Hazra, Hazra & Khan in Hazra et al.

|

Pliocene

|

Rajdanda Formation

|

{{Flag|India}}

|

A member of the family Fabaceae. Genus includes new species N. indica.

|

Ormosia cyclocarpa{{Cite journal|last1=Li |first1=X.-C. |last2=Manchester |first2=S. R. |last3=Xiao |first3=L. |last4=Wang |first4=Q. |last5=Hu |first5=Y. |last6=Sun |first6=B.-N. |year=2021 |title=Ormosia (Fabaceae: Faboideae) from the Miocene of southeastern China support historical expansion of the tropical genus in East Asia |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3561–3578 |doi=10.1080/08912963.2021.1877700 |s2cid=233806537 }}

|

Sp. nov

|

In press

|

Li et al.

|

Miocene

|

|

{{Flag|China}}

|

A species of Ormosia.

|

Peltophorum asiatica{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Bera |first3=S. |last4=Khan |first4=M. A. |year=2021 |title=First fossil evidence of marginal winged fruits of Peltophorum (Caesalpinioideae: Fabaceae) from India |journal=Brittonia |volume=73 |issue=3 |pages=241–250 |doi=10.1007/s12228-021-09679-4 |s2cid=238855320 }}

|

Sp. nov

|

In press

|

Hazra, Hazra & Khan in Hazra et al.

|

Pliocene

|

Rajdanda Formation

|

{{Flag|India}}

|

A species of Peltophorum.

|

Polyadopollenites minimus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Mimosa.

|

Salpinganthium{{Cite journal|last1=Poinar |first1=G. O. |last2=Chambers |first2=K. L. |year=2021 |title=Salpinganthium hispaniolanum gen. et sp. nov. (Fabaceae: Detarieae), a mid-Tertiary flower in Dominican amber |journal=Journal of the Botanical Research Institute of Texas |volume=15 |issue=2 |pages=559–567 |doi=10.17348/jbrit.v15.i2.1161 |s2cid=245099015 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Poinar & Chambers

|

Burdigalian

|

Dominican amber

|

{{Flag|Dominican Republic}}

|

A member of the family Fabaceae belonging to the tribe Detarieae. Genus includes new species S. hispaniolanum.

|

Striatopollis crassitectatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Macrolobium.

|

Syncolporites foveolatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Swartzia.

|

===Fagales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Eucarpinoxylon kayacikii

|

Sp. nov

|

Valid

|

Akkemik

|

Middle Miocene

|

Kesmekaya Volcanics

|

{{Flag|Turkey}}

|

A member of the family Betulaceae.

|

Fagus dodgei{{cite journal |last1=Wheeler |first1=E. A. |last2=Manchester |first2=S. R. |title=A diverse assemblage of Late Eocene woods from Oregon, western USA |journal=Fossil Imprint |date=2021 |volume=77 |issue=2 |pages=299–329 |doi=10.37520/fi.2021.022|s2cid=246981608 |doi-access=free }}

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Fagus species wood.

|

Lithocarpoxylon ashwillii

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Lithocarpoxylon species wood.

|

Morella absarokensis{{Cite journal|last1=Wheeler |first1=E. A. |last2=Baas |first2=P. |last3=Manchester |first3=S. R. |year=2021 |title=Wood Anatomy of Modern and Fossil Fagales in Relation to Phylogenetic Hypotheses, Familial Classification, and Patterns of Character Evolution |journal=International Journal of Plant Sciences |volume=183 |issue=1 |pages=61–86 |doi=10.1086/717328 |s2cid=244081754 |doi-access=free }}

|

Comb nov

|

valid

|

(Wheeler, Scott, & Barghoorn) Wheeler & Manchester

|

Middle Eocene

|

Lamar River Formation

|

{{Flag|USA}}
{{Flag|Wyoming}}

|

Moved from Myrica absarokensis (1978)

|

Morella scalariformis

|

Comb nov

|

valid

|

(Kruse) Wheeler, Baas, & Manchester

|

Middle Eocene

|

Eden Valley Formation

|

{{Flag|USA}}
{{Flag|Wyoming}}

|

Moved from Myrica scalariformis (1954)

|

Myricamentum{{Cite journal|last1=Wilde |first1=V. |last2=Frankenhäuser |first2=H. |last3=Lenz |first3=O. K. |year=2021 |title=A myricaceous male inflorescence with pollen in situ from the middle Eocene of Europe |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=4 |pages=873–883 |doi=10.1007/s12549-020-00479-4 |s2cid=233417128 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Wilde, Frankenhäuser & Lenz

|

Eocene

|

Eckfelder Maar

|

{{Flag|Germany}}

|

A catkin-like male inflorescence, probably of myricaceous affinity. Genus includes new species M. eckfeldensis.

|

Ostryoxylon gokceadaense

|

Sp. nov

|

Valid

|

Akkemik

|

Middle Miocene

|

Kesmekaya Volcanics

|

{{Flag|Turkey}}

|

A member of the family Betulaceae.

|

Palaeocarpinus borealis{{cite journal |last1=Correa-Narvaez |first1=J. E. |last2=Manchester |first2=S. R. |title=Distribution and Morphological Diversity of Palaeocarpinus (Betulaceae) from the Paleogene of the Northern Hemisphere. |journal=The Botanical Review |date=2021 |volume=88 |issue=2 |pages=161–203 |doi=10.1007/s12229-021-09258-y|s2cid=237795532 }}

|

Comb nov

|

Valid

|

(Heer) Correa & Manchester

|

Paleocene
late Paleocene

|

Menat Formation

|

{{Flag|France}}
{{flag|Puy-de-Dôme}}

|

A Coryloideae species.
First described as Anchietea borealis (1859).
Moved from Atriplex borealis (1912)

|

Palaeocarpinus parva

|

Sp. nov

|

Valid

|

Manchester & Correa

|

Middle Eocene

|

Clarno Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Coryloideae species.

|

Palaeocarpinus pterabaratra

|

Sp. nov

|

Valid

|

Correa & Manchester

|

Paleocene
Middle Paleocene

|

Fort Union Formation
"unnamed lower member"

|

{{Flag|USA}}
{{Flag|Wyoming}}

|

A Coryloideae species.

|

Palaeocarpinus pteravestigia

|

Sp. nov

|

Valid

|

Correa & Manchester

|

Paleocene
Middle Paleocene

|

Fort Union Formation
"unnamed lower member"

|

{{Flag|USA}}
{{Flag|Wyoming}}

|

A Coryloideae species.

|

Palaeocarya indica{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Kumar |first3=S. |last4=Mahato |first4=S. |last5=Bera |first5=M. |last6=Bera |first6=S. |last7=Khan |first7=M. A. |year=2021 |title=First fossil evidence of Palaeocarya (Engelhardioideae: Juglandaceae) from India and its biogeographical implications |journal=Journal of Systematics and Evolution |volume=59 |issue=6 |pages=1307–1320 |doi=10.1111/jse.12736 |s2cid=233897694 |doi-access=free }}

|

Sp. nov

|

Valid

|

Hazra, Hazra & Khan in Hazra et al.

|

Pliocene

|

Rajdanda Formation

|

{{Flag|India}}

|

A member of the family Juglandaceae.

|

Paralnoxylon

|

Nom. nov

|

Valid

|

Doweld

|

Paleocene

|

|

{{Flag|United Kingdom}}

|

A member of the family Betulaceae; a replacement name for Cantia Stopes (1915).

|

Quercoxylon yaltirikii

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Fagaceae.

|

====Fagalean research====

  • A study on wood anatomy in extant and fossil members of Fagales is published by Wheeler, Baas & Manchester who transfer two Eocene species from Myrica to Morella.

===Malpighiales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Euphorbiotheca deccanensis{{Cite journal|last1=Reback |first1=R. G. |last2=Kapgate |first2=D. K. |last3=Wurdack |first3=K. |last4=Manchester |first4=S. R. |year=2021 |title=Fruits of Euphorbiaceae from the Late Cretaceous Deccan Intertrappean Beds of India |journal=International Journal of Plant Sciences |volume=183 |issue=2 |page=000 |doi=10.1086/717691 |s2cid=239507275 }}

|

Sp. nov

|

In press

|

Reback et al.

|

Late Cretaceous (Maastrichtian)

|

Deccan Intertrappean Beds

|

{{Flag|India}}

|

A member of the family Euphorbiaceae.

|

Passiflora appalachiana{{Cite journal|last=Hermsen |first=E. J. |year=2021 |title=Review of the fossil record of Passiflora, with a description of new seeds from the Pliocene Gray Fossil Site, Tennessee, U.S.A. |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=533–550 |doi=10.1086/714282 |s2cid=233701901 }}

|

Sp. nov

|

Valid

|

Hermsen

|

Pliocene

|

Gray Fossil Site

|

{{Flag|United States}}
({{Flag|Tennessee}})

|

A species of Passiflora.

|

Passifloriidites{{Cite journal|last1=D'Apolito |first1=C. |last2=Jaramillo |first2=C. |last3=Harrington |first3=G. |year=2021 |title=Miocene Palynology of the Solimões Formation (Well 1-AS-105-AM), Western Brazilian Amazonia |journal=Smithsonian Contributions to Paleobiology |volume=105 |issue=105 |pages=iv-134 |doi=10.5479/si.16803493 |s2cid=243391850 |url=https://figshare.com/articles/book/Miocene_Palynology_of_the_Solim_es_Formation_Well_1-AS-105-AM_Western_Brazilian_Amazonia/16803493 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Passifloraceae. Genus includes new species P. pseudoperculatus.

|

Populoxylon sebenense

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Salicaceae.

|

Salicoxylon galatianum

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Salicaceae.

|

Verrustephanoporites circularis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Mascagnia.

|

===Oxalidales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Tropidogyne euthystyla{{Cite journal|last1=Poinar |first1=G. O. |last2=Chambers |first2=K. L. |last3=Vega |first3=F. E. |year=2021 |title=Tropidogyne euthystyla sp. nov., a new small-flowered addition to the genus from mid-Cretaceous Myanmar amber |journal=Journal of the Botanical Research Institute of Texas |volume=15 |issue=1 |pages=113–119 |doi=10.17348/jbrit.v15.i1.1053 |s2cid=237700194 |doi-access=free }}

|

Sp. nov

|

Valid

|

Poinar, Chambers & Vega

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A possible member of Cunoniaceae.

|

===Rosales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Alloceltidoxylon

|

Nom. nov

|

Valid

|

Doweld

|

Eocene

|

Clarno Formation

|

{{Flag|United States}}
({{Flag|Oregon}})

|

A flowering plant with possible affinities with urticalean rosids; a replacement name for Scottoxylon Wheeler & Manchester (2002).

|

Celtis popsii

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Celtis species wood.

|

Crataegoxylon sibiricum

|

Sp. nov

|

Valid

|

Dolezych, LePage & Williams

|

Oligocene (Chattian)

|

Korlikov Formation

|

{{Flag|Russia}}
({{Flag|Tomsk Oblast}})

|

|

Morus asiatica{{Cite journal|last1=Patel |first1=R. |last2=Hazra |first2=T. |last3=Rana |first3=R. S. |last4=Hazra |first4=M. |last5=Bera |first5=S. |last6=Khan |first6=M. A. |year=2021 |title=First fossil record of mulberry from Asia |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104459 |doi=10.1016/j.revpalbo.2021.104459 |bibcode=2021RPaPa.29204459P }}

|

Sp. nov

|

In press

|

Patel, Rana & Khan in Patel et al.

|

Early Eocene

|

|

{{Flag|India}}

|

A species of Morus.

|

Prunoidoxylon prunoides

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Rosaceae.

|

Psilatriporites minimus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Celtis.

|

Pyracantha pseudococcinea

|

Sp. nov

|

Valid

|

Striegler

|

Miocene (Tortonian)

|

Rauno Formation

|

{{Flag|Germany}}

|

A species of Pyracantha.

|

Ulmoxylon kasapligilii

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Ulmaceae.

|

Ventilago tibetensis{{Cite journal|last1=Del Rio |first1=C. |last2=Wang |first2=T.-X. |last3=Xu |first3=X.-T. |last4=Sabroux |first4=R. |last5=Spicer |first5=T. E. V. |last6=Liu |first6=J. |last7=Chen |first7=P.-R. |last8=Wu |first8=F.-X. |last9=Zhou |first9=Z.-K. |last10=Su |first10=T. |year=2021 |title=Ventilago (Rhamnaceae) Fruit from the Middle Eocene of the Central Tibet, China |journal=International Journal of Plant Sciences |volume=182 |issue=7 |pages=638–648 |doi=10.1086/715507 |s2cid=236410986 }}

|

Sp. nov

|

Valid

|

Del Rio et al.

|

Middle Eocene

|

|

{{Flag|China}}

|

A species of Ventilago.

|

Zelkovoxylon crystalliferum

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Ulmaceae.

|

==Malvids==

===Malvales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Bombacacidites hooghiemstrae

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Malvaceae.

|

Craigia lincangensis{{Cite journal|last1=Wang |first1=B. |last2=Zhang |first2=S. |last3=Zhang |first3=P. |last4=Yang |first4=Y. |last5=Chen |first5=J. |last6=Zhang |first6=Y. |last7=Xie |first7=S. |year=2021 |title=A new occurrence of Craigia (Malvaceae) from the Miocene of Yunnan and its biogeographic significance |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3402–3412 |doi=10.1080/08912963.2020.1867980 |s2cid=234188949 }}

|

Sp. nov

|

In press

|

Wang & Xie in Wang et al.

|

Late Miocene

|

|

{{Flag|China}}

|

A species of Craigia

|

Dipterocapus fotanensis{{Cite journal|last1=Chen |first1=J. |last2=Han |first2=L. |last3=Hua |first3=Y. |last4=Wu |first4=G. |last5=Sun |first5=B. |year=2021 |title=Fruits and leaves of Dipterocarpus from the Miocene of Zhangpu, Fujian, and its geological significance |journal=Arabian Journal of Geosciences |volume=14 |issue=13 |pages=Article 1270 |doi=10.1007/s12517-021-07445-0 |s2cid=235664238 }}

|

Sp. nov

|

Valid

|

Chen et al.

|

Miocene

|

|

{{Flag|China}}

|

A species of Dipterocarpus

|

Discoidites angulosus{{Cite journal|last1=Huang |first1=H. |last2=Pérez-Pinedo |first2=D. |last3=Morley |first3=R. J. |last4=Dupont-Nivet |first4=G. |last5=Philip |first5=A. |last6=Win |first6=Z. |last7=Aung |first7=D. W. |last8=Licht |first8=A. |last9=Jardine |first9=P. E. |last10=Hoorn |first10=C. |year=2021 |title=At a crossroads: The late Eocene flora of central Myanmar owes its composition to plate collision and tropical climate |journal=Review of Palaeobotany and Palynology |volume=291 |pages=Article 104441 |doi=10.1016/j.revpalbo.2021.104441 |bibcode=2021RPaPa.29104441H |s2cid=235421153 |doi-access=free |hdl=11245.1/6fb3c1f3-e28b-4a90-9d4f-dbe38f07a512 |hdl-access=free }}

|

Sp. nov

|

In press

|

Huang, Morley & Hoorn in Huang et al.

|

Late Eocene

|

Yaw Formation

|

{{Flag|Myanmar}}

|

Pollen probably derived from plants belonging to the genus Brownlowia.

|

Thymelipollis amazonicus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Thymelaeaceae.

|

Tilia asiatica{{Cite journal|last1=Jia |first1=L.B. |last2=Nam |first2=G.S. |last3=Su |first3=T. |last4=Stull |first4=G.W. |last5=Li |first5=S.F. |last6=Huang |first6=Y.J. |last7=Zhou |first7=Z.K. |year=2021 |title=Fossil fruits of Firmiana and Tilia from the middle Miocene of South Korea and the efficacy of the Bering land bridge for the migration of mesothermal plants |journal=Plant Diversity |volume=43 |issue=6 |pages=480–491 |doi=10.1016/j.pld.2020.12.006 |pmid=35024517 |pmc=8720707 |doi-access=free }}

|

Sp. nov

|

In press

|

Jia & Nam in Jia et al.

|

Middle Miocene

|

Pohang Basin

|

{{Flag|South Korea}}

|

A species of Tilia

|

Wataria kvacekii

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Wataria species wood.

|

===Myrtales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Florschuetzia impostora

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen, possibly of a member of the family Lythraceae.

|

Lazarocardenasoxylon{{Cite journal|last1=Estrada-Ruiz |first1=E. |last2=Martínez-Cabrera |first2=H. I. |last3=García-Hernández |first3=I. P. |year=2021 |title=New dicotyledonous woods from the San Carlos Formation (Upper Cretaceous) in Northern Mexico |journal=IAWA Journal |volume=43 |issue=1–2 |pages=66–79 |doi=10.1163/22941932-bja10079 |s2cid=244902117 }}

|

Gen. et sp. nov

|

In press

|

Estrada-Ruiz & Martínez-Cabrera in Estrada-Ruiz, Martínez-Cabrera & García-Hernández

|

Late Cretaceous

|

San Carlos Formation

|

{{Flag|Mexico}}

|

Possibly a member of the family Myrtaceae. Genus includes new species L. aldamense.

|

Lythrum portugalliense{{Cite journal|last1=Vieira |first1=M. |last2=Zetter |first2=R. |last3=Coiro |first3=M. |last4=Grímsson |first4=F. |year=2021 |title=Pliocene Lythrum (loosestrife, Lythraceae) pollen from Portugal and the Neogene establishment of European lineages |journal=Review of Palaeobotany and Palynology |volume=296 |pages=Article 104548 |doi=10.1016/j.revpalbo.2021.104548 | issn=0034-6667 |doi-access=free |hdl=10362/144398 |hdl-access=free }}

|

Sp. nov

|

In press

|

Vieira et al.

|

Pliocene (Piacenzian)

|

|

{{Flag|Portugal}}

|

A species of Lythrum.

|

Syzygium guipingensis{{Cite journal|last1=Li |first1=Y. |last2=Huang |first2=L. |last3=Quan |first3=C. |last4=Jin |first4=J. |last5=Oskolski |first5=A. A. |year=2021 |title=Fossil wood of Syzygium from the Miocene of Guangxi, South China: the earliest fossil evidence of the genus in eastern Asia |journal=IAWA Journal |volume=42 |issue=4 |pages=435–441 |doi=10.1163/22941932-bja10069|issn=0928-1541 |s2cid=240550082 }}

|

Sp. nov

|

In press

|

Li et al.

|

Miocene

|

Erzitang Formation

|

{{Flag|China}}

|

A species of Syzygium.

|

Terminalioxylon mozambicense{{Cite journal|last1=Bamford |first1=M. |last2=Pickford |first2=M. |title=Stratigraphy, chronology and palaeontology of the Tertiary rocks of the Cheringoma Plateau, Mozambique |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=187–213 |doi=10.37520/fi.2021.014 |s2cid=245074611 |url=http://fi.nm.cz/en/clanek/stratigraphy-chronology-and-palaeontology-of-the-tertiary-rocks-of-the-cheringoma-plateau-mozambique-2/ |doi-access=free }}

|

Sp. nov

|

Valid

|

Bamford & Pickford

|

Probably late Eocene

|

|

{{Flag|Mozambique}}

|

A member of the family Combretaceae.

|

Trapa sanyingensis{{Cite journal|last1=Aung |first1=A. T. |last2=Del Rio |first2=C. |last3=Wang |first3=T.-X. |last4=Liu |first4=J. |last5=Spicer |first5=T. E. V. |last6=Su |first6=T. |year=2021 |title=Fossil fruits and pollen grains of Trapa from the Upper Pliocene of the Sanying Formation (Yunnan, China) |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104498 |doi=10.1016/j.revpalbo.2021.104498 |bibcode=2021RPaPa.29304498A }}

|

Sp. nov

|

In press

|

Aung et al.

|

Late Pliocene

|

Sanying Formation

|

{{Flag|China}}

|

A water caltrop.

|

Verrutricolporites pusillus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen, possibly of a member of the family Lythraceae.

|

Xystonia{{Cite journal|last1=Carvalho |first1=M. |last2=Herrera |first2=F. |last3=Gómez |first3=S. |last4=Martínez |first4=C. |last5=Jaramillo |first5=C. |year=2021 |title=Early records of Melastomataceae from the middle-late Paleocene rainforests of South America conflict with Laurasian origins |journal=International Journal of Plant Sciences |volume=182 |issue=5 |pages=401–412 |doi=10.1086/714053 |s2cid=233910850 }}

|

Gen. et sp. nov

|

Valid

|

Carvalho et al.

|

Paleocene

|

Bogotá Formation

|

{{Flag|Colombia}}

|

A member of the family Melastomataceae. Genus includes new species X. simonae.

|

===Sapindales===

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Aceroxylon aceroides

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Sapindaceae.

|

Ampelorhiza{{Cite journal|last1=Jud |first1=N. A. |last2=Allen |first2=S. E. |last3=Nelson |first3=C. W. |last4=Bastos |first4=C. L. |last5=Chery |first5=J. G. |title=Climbing since the early Miocene: The fossil record of Paullinieae (Sapindaceae) |year=2021 |journal=PLOS ONE |volume=16 |issue=4 |pages=e0248369 |doi=10.1371/journal.pone.0248369 |pmid=33826635 |pmc=8026063 |bibcode=2021PLoSO..1648369J |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Jud et al.

|

Early Miocene

|

Cucaracha Formation

|

{{Flag|Panama}}

|

A member of the family Sapindaceae belonging to the subfamily Sapindoideae and the tribe Paullinieae. Genus includes new species A. heteroxylon.

|

Anacardium gassonii{{Cite journal|last1=Rodríguez-Reyes |first1=O. |last2=Estrada-Ruiz |first2=E. |last3=Monje Dussán |first3=C. |last4=de Andrade Brito |first4=L. |last5=Terrazas |first5=T. |title=A new Oligocene-Miocene tree from Panama and historical Anacardium migration patterns |year=2021 |journal=PLOS ONE |volume=16 |issue=6 |pages=e0250721 |doi=10.1371/journal.pone.0250721 |pmid=34077439 |pmc=8171895 |bibcode=2021PLoSO..1650721R |doi-access=free }}

|

Sp. nov

|

Valid

|

Rodríguez-Reyes, Estrada-Ruiz & Terrazas in Rodríguez-Reyes et al.

|

Oligocene-Miocene

|

|

{{Flag|Panama}}

|

A species of Anacardium.

|

Atalantioxylon thanobolensis{{Cite journal|last1=Soomro |first1=N. |last2=Mangi |first2=J. |last3=Panhwer |first3=M. |last4=Jatoi |first4=G. |last5=Khuhro |first5=S. |last6=Khokhar |first6=Q. |last7=Khan |first7=S. |last8=Mengal |first8=A. |last9=Shaikh |first9=N. |year=2021 |title=Anatomical characteristics of fossil wood collected from the Manchar Formation (Miocene), Thano Bula Khan, Sindh, Pakistan |journal=Italian Botanist |volume=11 |pages=1–8 |doi=10.3897/italianbotanist.11.60344 |s2cid=233274837 |doi-access=free }}

|

Sp. nov

|

Valid

|

Soomro et al.

|

Miocene

|

Manchar Formation

|

{{Flag|Pakistan}}

|

Fossil wood of a member of the family Rutaceae.

|

Melia santangensis{{Cite journal|last1=Liu |first1=W.-Q. |last2=Xu |first2=S.-L. |last3=Fu |first3=Q.-Y. |last4=Quan |first4=C. L. |last5=Jin |first5=J.-H. |title=Late Oligocene Melia (Meliaceae) from the Nanning Basin of South China and it's biogeographical implication |year=2021 |journal=Journal of Palaeogeography |volume=10 |issue=1 |pages=Article number 16 |doi=10.1186/s42501-021-00097-x |bibcode=2021JPalG..10...16L |s2cid=235690851 |doi-access=free }}

|

Sp. nov

|

Valid

|

Liu, Xu & Jin in Liu et al.

|

Late Oligocene

|

Yongning Formation

|

{{Flag|China}}

|

A species of Melia.

|

Pistacia terrazasae

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Pistacia species wood.

|

Proteacidites poriscabratus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Sapindaceae.

|

Sorindeioxylon

|

Gen. et sp. nov

|

Valid

|

Bamford & Pickford

|

Probably late Eocene

|

Mazamba Formation

|

{{Flag|Mozambique}}

|

A member of the family Anacardiaceae. Genus includes new species S. gorongosense.

|

Syncolporites tenuicolpatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the genus Serjania.

|

Tetradium nanningense{{Cite journal|last1=Huang |first1=L.-L. |last2=Jin |first2=J.-H. |last3=Quan |first3=C. |last4=Oskolski |first4=A. A. |title=Earliest fossil record of the genus Tetradium (Rutaceae) in Asia: implications for its evolution and palaeoecology |year=2021 |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=2065–2074 |doi=10.1002/spp2.1394 |s2cid=237688959 }}

|

Sp. nov

|

Valid

|

Huang et al.

|

Late Oligocene

|

Yongning Formation

|

{{Flag|China}}

|

A species of Tetradium.

|

Verrutricolporites simplex

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Pollen of a member of the family Simaroubaceae.

|

==Non eurosid superrosids==

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Hamamelidoxylon crystalliferum

|

Sp nov

|

Valid

|

Wheeler & Manchester

|

Late Eocene

|

John Day Formation

|

{{Flag|USA}}
{{Flag|Oregon}}

|

A Hamamelidoxylon species wood.

|

Liquidambar guipingensis{{Cite journal|last1=Huang |first1=L.-L. |last2=Jin |first2=J.-H. |last3=Quan |first3=C. |last4=Oskolski |first4=A. A. |year=2021 |title=New occurrences of Altingiaceae fossil woods from the Miocene and upper Pleistocene of South China with phytogeographic implications |journal=Journal of Palaeogeography |volume=10 |issue=4 |pages=482–493 |doi=10.1016/j.jop.2021.11.001 |bibcode=2021JPalG..10..482H |s2cid=252959116 |doi-access=free }}

|

Sp. nov

|

In press

|

Huang et al.

|

Miocene

|

Erzitang Formation

|

{{Flag|China}}

|

A species of Liquidambar.

|

Liquidambaroxylon efeae{{Cite journal|last=Akkemik |first=Ü. |year=2021 |title=A re-examination of the angiosperm wood record from the early and middle Miocene of Turkey, and new species descriptions |journal=Acta Palaeobotanica |volume=61 |issue=1 |pages=42–94 |doi=10.35535/acpa-2021-0004 |s2cid=237741731 |doi-access=free }}

|

Sp. nov

|

Valid

|

Akkemik

|

Early Miocene

|

Hançili Formation

|

{{Flag|Turkey}}

|

A member of the family Altingiaceae.

|

Obirafructus{{Cite journal|last1=Kajita |first1=Y. |last2=Suzuki |first2=M. H. |last3=Nishida |first3=H. |year=2021 |title=A Multicarpellary Apocarpous Gynoecium from the Late Cretaceous (Coniacian–Santonian) of the Upper Yezo Group of Obira, Hokkaido, Japan: Obirafructus kokubunii gen. & sp. nov. |journal=Acta Phytotaxonomica et Geobotanica |volume=72 |issue=1 |pages=1–21 |doi=10.18942/apg.202009 }}

|

Gen. et sp. nov

|

Valid

|

Kajita & Nishida in Kajita, Suzuki & Nishida

|

Late Cretaceous (Coniacian–Santonian)

|

Haborogawa Formation

|

{{Flag|Japan}}

|

A member of Saxifragales of uncertain phylogenetic placement. Genus includes new species O. kokubunii.

|

Paleoaltingia{{Cite journal|last1=Lai |first1=Y. |last2=Gandolfo |first2=M. A. |last3=Crepet |first3=W. L. |last4=Nixon |first4=K. C. |year=2021 |title=Paleoaltingia gen. nov., a new genus of Altingiaceae from the Late Cretaceous of New Jersey |journal=American Journal of Botany |volume=108 |issue=3 |pages=461–471 |doi=10.1002/ajb2.1618 |pmid=33660257 |s2cid=232115011 |doi-access=free }}

|

Gen. nov

|

Valid

|

Lai et al.

|

Late Cretaceous (Turonian)

|

|

{{Flag|United States}}
({{Flag|New Jersey}})

|

A member of Altingiaceae. Genus includes P. ovum-dinosauri and P. polyodonta.

|

= Other angiosperms =

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Araliaephyllum vittenburgii{{Cite journal|last1=Golovneva |first1=L. |last2=Bugdaeva |first2=E. |last3=Volynets |first3=E. |last4=Sun |first4=Y. |last5=Zolina |first5=A. |title=Angiosperm diversification in the Early Cretaceous of Primorye, Far East of Russia |year=2021 |journal=Fossil Imprint |volume=77 |issue=2 |pages=231–255 |doi=10.37520/fi.2021.017 |s2cid=245545364 |url=http://fi.nm.cz/clanek/angiosperm-diversification-in-the-early-cretaceous-of-primorye-far-east-of-russia/ |doi-access=free }}

|

Sp. nov

|

Valid

|

Golovneva & Volynets in Golovneva et al.

|

Early Cretaceous (Albian)

|

Galenki Formation

|

{{Flag|Russia}}
({{Flag|Primorsky Krai}})

|

A flowering plant of uncertain phylogenetic placement.

|

Baderadea{{Cite journal|last1=Pessoa |first1=E. M. |last2=Ribeiro |first2=A. C. |last3=Jud |first3=N. A. |year=2021 |title=A eudicot leaf from the Lower Cretaceous (Aptian, Araripe Basin) Crato Konservat-Lagerstätte |journal=American Journal of Botany |volume=108 |issue=10 |pages=2055–2065 |doi=10.1002/ajb2.1751 | issn=0002-9122 |pmid=34647319 |s2cid=238858698 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Pessoa, Ribeiro & Jud

|

Early Cretaceous (Aptian)

|

Crato Formation

|

{{Flag|Brazil}}

|

A herbaceous eudicot similar to some members of Ranunculales. Genus includes new species B. pinnatissecta.

|

Byttneripollis rugulatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Dilcherifructus{{Cite journal|last=Wang |first=X. |title=The Currently Earliest Angiosperm Fruit from the Jurassic of North America |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=4 |pages=416–422 |doi=10.37819/biosis.001.04.0160 |s2cid=245724946 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Wang

|

Middle Jurassic

|

Simón Formation

|

{{Flag|Mexico}}

|

Possibly a fruit of an early flowering plant. Genus includes new species D. mexicana.

|

Farabeipollis deccanensis{{Cite journal|last1=Sonkusare |first1=H. |last2=Samant |first2=B. |last3=Mohabey |first3=D. M. |year=2021 |title=Palynoassemblage from intertrappean sediments of Satpura Group, Betul district, Madhya Pradesh: implications in understanding age and palaeoclimate |journal=Journal of the Palaeontological Society of India |volume=66 |issue=1 |pages=35–54 |url=https://www.researchgate.net/publication/353006021 }}

|

Sp. nov

|

Valid

|

Sonkusare, Samant & Mohabey

|

Late Cretaceous (Maastrichtian)

|

Deccan Intertrappean Beds

|

{{Flag|India}}

|

Pollen of a flowering plant of uncertain affinity.

|

Florigerminis{{cite book |last1=Cui |first1=D.-F. |last2=Hou |first2=Y. |last3=Yin |first3=P. |last4=Wang |first4=X. |year=2021 |chapter=A Jurassic flower bud from the Jurassic of China |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |journal=Geological Society, London, Special Publications |publisher=The Geological Society of London |doi=10.1144/SP521-2021-122 |s2cid=244737990 }}

|

Gen. et sp. nov

|

In press

|

Cui et al.

|

Middle-Late Jurassic

|

Jiulongshan Formation

|

{{Flag|China}}

|

A possible flower bud.
The type species is F. jurassica.
First announced online 2021, Final article published 2022.

|

File:Florigerminis F2.A-B.jpg]]

Gansufructus{{cite journal |last1=Du |first1=B. |last2=Zhang |first2=M. |last3=Sun |first3=B. |last4=Li |first4=A. |last5=Zhang |first5=J. |last6=Yan |first6=D. |last7=Xie |first7=S. |last8=Wu |first8=J. |year=2021 |title=An Exceptionally Well-Preserved Herbaceous Eudicot from the Early Cretaceous (late Aptian-early Albian) of Northwest China |journal=National Science Review |volume=8 |issue=12 |pages=nwab084 |doi=10.1093/nsr/nwab084 |pmid=34987839 |pmc=8692937 }}

|

Gen. et sp. nov

|

In press

|

Du in Du et al.

|

Early Cretaceous (late Aptian-early Albian)

|

Zhonggou Formation

|

{{Flag|China}}

|

A eudicot of uncertain phylogenetic placement. Genus includes new species G. saligna.

|

Inaperturopollenites tectatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Ladakhipollenites? corvattatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Ladakhipollenites? endoporatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Ladakhipollenites nanus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Ladakhipollenites? sphaericus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Margocolporites bilinearis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Margocolporites incertus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Multiporopollenites intermedius

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Nigericolpites{{Cite journal|last=Hernández |first=J. A. R. |year=2021 |title=Nigericolpites: a replacement name for the illegitimate Maastrichtian magnoliopsid pollen genus Clavatricolpites Hoeken-Klink. (Angiospermae: Magnoliopsida) |journal=Grana |volume=60 |issue=5 |pages=370–371 |doi=10.1080/00173134.2020.1827025|issn=0017-3134 |s2cid=234286062 }}

|

Nom. nov

|

Valid

|

Hernández

|

Late Cretaceous (Maastrichtian)

|

|

{{Flag|Nigeria}}

|

Pollen of a flowering plant; a replacement name for Clavatricolpites Hoeken-Klinkenberg (1964).

|

Psilaperiporites circinatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Psilaperiporites depressus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Psilastephanocolporites ectoporatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Psilastephanocolporites pseudomarinamensis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Ranunculacidites pontoreticulatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retibrevitricolpites pseudoretibolus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retiperiporites retiporatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retistephanocolpites liberalis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retistephanocolporites loxocolpatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitrescolpites benjaminensis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitrescolpites brevicolpatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitrescolpites grossus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitrescolpites kriptoporus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitrescolpites marginatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitriporites discretus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Retitriporites sifonis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites apertus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites? colpiverrucosus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites crassinexinicus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites crassitectatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites grossomurus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites guttatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites lolongatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites protoguttatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites pseudocrassopolaris

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites pseudopilatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites pseudoscabratus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites quantulus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Rhoipites vilis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Tetracolporopollenites nanus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Tetracolporopollenites xatanawensis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil pollen.

|

Other plants

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Adelocladoxis{{Cite journal|last1=Durieux |first1=T. |last2=Lopez |first2=M. A. |last3=Bronson |first3=A. W. |last4=Tomescu |first4=A. M. F. |year=2021 |title=A new phylogeny of the cladoxylopsid plexus: contribution of an early cladoxylopsid from the Lower Devonian (Emsian) of Quebec |journal=American Journal of Botany |volume=108 |issue=10 |pages=2066–2095 |doi=10.1002/ajb2.1752|issn=0002-9122 |pmid=34664712 |s2cid=239028044 |doi-access=free }}

|

Gen. et sp. nov

|

Valid

|

Durieux et al.

|

Devonian (Emsian)

|

Battery Point Formation

|

{{Flag|Canada}}
({{Flag|Quebec}})

|

A member of Cladoxylopsida. Genus includes new species A. praecox.

|

Baragwanathia brevifolioides{{Cite journal|last1=Kraft |first1=P. |last2=Kvaček |first2=Z. |title=Baragwanathia brevifolioides, a nomen novum for B. brevifolia P.Kraft et Kvaček, 2017 |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=53–54 |doi=10.37520/fi.2021.006 |s2cid=245034157 |url=http://fi.nm.cz/en/clanek/baragwanathia-brevifolioides-a-nomen-novum-for-b-brevifolia-p-kraft-et-kvacek-2017-2/ |doi-access=free }}

|

Nom. nov

|

Valid

|

Kraft & Kvaček

|

Silurian (Přídolí)

|

Požáry Formation

|

{{Flag|Czech Republic}}

|

A member of the family Drepanophycaceae; a replacement name for Baragwanathia brevifolia Kraft & Kvaček (2017).

|

Camarozonosporites fossulatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Lycopodiaceae.

|

Cingulatisporites cristatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the genus Selaginella.

|

Cingulatisporites matisiensis

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the genus Phaeoceros.

|

Closterium mosbruggeri{{Cite journal|last1=Ivanov |first1=D. |last2=Belkinova |first2=D. |year=2021 |title=Closterium mosbruggeri sp. nov.: a new fossil species from the middle Miocene of Northwest Bulgaria |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=69–74 |doi=10.1007/s12549-020-00476-7 |s2cid=231859320 }}

|

Sp. nov

|

Valid

|

Ivanov & Belkinova

|

Miocene (Serravallian)

|

|

{{Flag|Bulgaria}}

|

A green alga, a species of Closterium.

|

Colpodexylon mergae{{Cite journal|last1=Harris |first1=C. |last2=Gess |first2=R. W. |last3=Prestianni |first3=C. |last4=Bamford |first4=M. K. |year=2021 |title=A Late Devonian refugium for Colpodexylon (Lycopsida) at high latitude |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104481 |doi=10.1016/j.revpalbo.2021.104481 |bibcode=2021RPaPa.29304481H |url=https://orbi.uliege.be/bitstream/2268/295475/1/Harris%20et%20al.%202021%20Colpodexylon.pdf }}

|

Sp. nov

|

In press

|

Harris et al.

|

Devonian (Famennian)

|

Witpoort Formation

|

{{Flag|South Africa}}

|

A member of Lycopsida.

|

Colpodexylon pullumpedes

|

Sp. nov

|

In press

|

Harris et al.

|

Devonian (Famennian)

|

Witpoort Formation

|

{{Flag|South Africa}}

|

A member of Lycopsida.

|

Cordaabaxicutis martii{{Cite journal|last1=Šimůnek |first1=Z. |last2=Lojka |first2=R. |year=2021 |title=Carboniferous cuticles from the Lubná coal seam (Kladno Formation, Kladno-Rakovník Basin, Czech Republic) |journal=Palaeontographica Abteilung B |volume=303 |issue=4–6 |pages=77–117 |doi=10.1127/palb/2021/0076 |bibcode=2021PalAB.303...77S |s2cid=244561451 }}

|

Sp. nov

|

Valid

|

Šimůnek & Lojka

|

Carboniferous (Pennsylvanian)

|

Kladno Formation

|

{{Flag|Czech Republic}}

|

Cordaitalean cuticles.

|

Cordaabaxicutis papillosus

|

Sp. nov

|

Valid

|

Šimůnek & Lojka

|

Carboniferous (Pennsylvanian)

|

Kladno Formation

|

{{Flag|Czech Republic}}

|

Cordaitalean cuticles.

|

Cordaadaxicutis raristomatus

|

Sp. nov

|

Valid

|

Šimůnek & Lojka

|

Carboniferous (Pennsylvanian)

|

Kladno Formation

|

{{Flag|Czech Republic}}

|

Cordaitalean cuticles.

|

Cynodontium luthii{{Cite journal|last1=Bippus |first1=A. C. |last2=Rothwell |first2=G. W. |last3=Stockey |first3=R. A. |year=2021 |title=Cynodontium luthii sp. nov.: a permineralized moss gametophyte from the Late Cretaceous of the North Slope of Alaska |journal=American Journal of Botany |volume=108 |issue=3 |pages=495–504 |doi=10.1002/ajb2.1617 |pmid=33650114 |s2cid=232091615 |doi-access=free }}

|

Sp. nov

|

Valid

|

Bippus, Rothwell & Stockey

|

Late Cretaceous

|

|

{{Flag|United States}}
({{Flag|Alaska}})

|

A moss belonging to the family Rhabdoweisiaceae, a species of Cynodontium.

|

Dayvaultia{{Cite journal|last1=Manchester |first1=S. R. |last2=Zhang |first2=X. |last3=Hotton |first3=C. L. |last4=Wing |first4=S. L. |last5=Crane |first5=P. R. |year=2021 |title=Distinctive quadrangular seed-bearing structures of gnetalean affinity from the Late Jurassic Morrison Formation of Utah, USA |journal=Journal of Systematic Palaeontology |volume=19 |issue=10 |pages=743–760 |doi=10.1080/14772019.2021.1968522 |s2cid=239021014 |url=https://www.tandfonline.com/doi/abs/10.1080/14772019.2021.1968522 |url-access=subscription }}

|

Gen. et sp. nov

|

Valid

|

Manchester et al.

|

Late Jurassic

|

Morrison Formation

|

{{Flag|United States}}
({{Flag|Utah}})

|

A seed-bearing structure of gnetalean affinity. Genus includes new species D. tetragona.

|

Distefanopolia{{Cite journal|last1=Barattolo |first1=F. |last2=Romano |first2=R. |last3=Conrad |first3=M. |title=Evidence of external gametophores in puzzling Late Triassic–Early Jurassic dasycladalean green algae |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=4 |pages=901–919 |doi=10.4202/app.00883.2021 |doi-access=free }}

|

Gen. et comb. nov

|

Valid

|

Barattolo, Romano & Conrad

|

Late Triassic and possibly Early Jurassic

|

|

{{Flag|Austria}}

{{Flag|Czech Republic}}

{{Flag|Germany}}

{{Flag|Greece}}

{{Flag|Italy}}

{{Flag|Oman}}

{{Flag|Slovakia}}

|

A green alga belonging to the group Dasycladales and the family Bornetelleae. Genus includes "Heteroporella" micropora Di Stefano & Senowbari-Daryan (1985), "Heteroporella" macropora Di Stefano, 1981 ex Di Stefano & Senowbari-Daryan (1985), "Chinianella" zanklii Ott (1967), "Chinianella" crosii Ott (1968) and "Heteroporella" carpatica Bystrický (1967).

|

Distichophytum mogilatii{{Cite journal|last=Naugolnykh |first=S. V. |year=2021 |title=A new species of Distichophytum Mägdefrau, 1938 from the Lower Devonian of Siberia (the Torgashino locality, Krasnojarsk krai, Russia) |journal=Wulfenia |volume=28 |pages=141–150 |url=https://www.researchgate.net/publication/356613631 }}

|

Sp. nov

|

Valid

|

Naugolnykh

|

Devonian

|

|

{{Flag|Russia}}
({{Flag|Krasnoyarsk Krai}})

|

A member of the family Zosterophyllaceae.

|

Dragastanella{{Cite journal|last1=Barattolo |first1=F. |last2=Bucur |first2=I. I. |last3=Marian |first3=A. V. |year=2021 |title=Deciphering voids in Dasycladales, the case of Dragastanella transylvanica, a new Lower Cretaceous triploporellacean genus and species from Romania |journal=Journal of Paleontology |volume=95 |issue=5 |pages=889–905 |doi=10.1017/jpa.2021.40|bibcode=2021JPal...95..889B | issn=0022-3360 |s2cid=236346831 |doi-access=free }}

|

Gen. et sp. et comb. nov

|

Valid

|

Barattolo, Bucur & Marian

|

Early Cretaceous

|

|

{{Flag|Italy}}

{{Flag|Romania}}

{{Flag|Spain}}

|

A green alga belonging to the group Dasycladales. Genus includes new species D. transylvanica, as well as "Zittelina" hispanica Masse, Arias & Vilas (1993), "Zittelina" massei Bucur, Granier & Săsăran (2010) and "Triploporella" matesina Barattolo (1980).

|

Elandia{{Cite journal|last1=Gess |first1=R. W. |last2=Prestianni |first2=C. |year=2021 |title=An early Devonian flora from the Baviaanskloof Formation (Table Mountain Group) of South Africa |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 11859 |doi=10.1038/s41598-021-90180-z |pmid=34088916 |pmc=8178408 |bibcode=2021NatSR..1111859G }}

|

Gen. et sp. nov

|

Valid

|

Gess & Prestianni

|

Devonian (Lochkovian?)

|

Baviaanskloof Formation

|

{{Flag|South Africa}}

|

An early polysporangiophyte. Genus includes new species E. itshoba.

|

Flabellopteris{{Cite journal|last1=Gess |first1=R. W. |last2=Prestianni |first2=C. |year=2021 |title=Flabellopteris lococannensis gen. et sp. nov.: A new fern-like plant from the Famennian of South Africa |journal=Review of Palaeobotany and Palynology |volume=297 |pages=Article 104585 |doi=10.1016/j.revpalbo.2021.104585 |s2cid=245209478 |url=https://orbi.uliege.be/bitstream/2268/302493/1/1-s2.0-S0034666721002098-main.pdf }}

|

Gen. et sp. nov

|

In press

|

Gess & Prestianni

|

Devonian (Famennian)

|

Witpoort Formation

|

{{Flag|South Africa}}

|

A fern-like plant of uncertain affinities. Genus includes new species F. lococannensis.

|

Foraminisporis connexus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the genus Anthoceros.

|

Frullania kachinensis{{Cite journal|last1=Li |first1=Y. |last2=Wang |first2=Y.-D. |last3=Feldberg |first3=K. |last4=Wang |first4=Q. |last5=Yang |first5=X.-J. |year=2021 |title=A new leafy liverwort of Frullania (Frullaniaceae, Porellales) from the mid-Cretaceous Kachin amber, Myanmar |journal=Geological Journal |volume=56 |issue=10 |pages=5046–5057 |doi=10.1002/gj.4222 |s2cid=238795680 }}

|

Sp. nov

|

Valid

|

Li et al.

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A liverwort, a species of Frullania.

|

Frullania palaeoafricana{{Cite journal|last1=Bouju |first1=V. |last2=Feldberg |first2=K. |last3=Kaasalainen |first3=U. |last4=Schäfer-Verwimp |first4=A. |last5=Hedenäs |first5=L. |last6=Buck |first6=W. R. |last7=Wang |first7=B. |last8=Perrichot |first8=V. |last9=Schmidt |first9=A. R. |title=Miocene Ethiopian amber: a new source of fossil cryptogams |journal=Journal of Systematics and Evolution |year=2022 |volume=60 |issue=4 |pages=932–954 |doi=10.1111/jse.12796 |s2cid=236586190 |url=https://hal-insu.archives-ouvertes.fr/insu-03231697/file/BOUJU-2021.pdf }}

|

Sp. nov

|

In press

|

Bouju et al.

|

Miocene

|

Ethiopian amber

|

{{Flag|Ethiopia}}

|

A liverwort, a species of Frullania.

|

Frullania shewanensis

|

Sp. nov

|

In press

|

Bouju et al.

|

Miocene

|

Ethiopian amber

|

{{Flag|Ethiopia}}

|

A liverwort, a species of Frullania.

|

Gilboaphyton fuyunensis{{Cite journal|last1=Liu |first1=B.-C. |last2=Bai |first2=J. |last3=Wang |first3=Y. |last4=Yang |first4=N. |last5=Xu |first5=H.-H. |year=2021 |title=On the discovery of Gilboaphyton (Lycopsida) from the Upper Devonian of East Junggar, Xinjiang, and its global distribution |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104473 |doi=10.1016/j.revpalbo.2021.104473 |bibcode=2021RPaPa.29204473L }}

|

Sp. nov

|

In press

|

Liu et al.

|

Late Devonian

|

Kaxiweng Formation

|

{{Flag|China}}

|

A member of Protolepidodendrales.

|

Guazia{{Cite journal|last1=Wang |first1=D.-M. |last2=Liu |first2=L. |last3=Zhou |first3=Y. |last4=Qin |first4=M. |last5=Meng |first5=M.-C. |last6=Guo |first6=Y. |last7=Xue |first7=J.-Z. |year=2021 |title=Guazia, the earliest ovule without cupule but with unique integumentary lobes |journal=National Science Review |volume=9 |issue=4 |pages=nwab196 |doi=10.1093/nsr/nwab196 |pmid=35386924 |pmc=8982201 }}

|

Gen. et sp. nov

|

In press

|

Wang et al.

|

Late Devonian

|

Wutong Formation

|

{{Flag|China}}

|

A seed plant of uncertain phylogenetic placement. Genus includes new species G. dongzhiensis.

|

Ixostrobus daohugouensis{{Cite journal|last1=Na |first1=Y. |last2=Sun |first2=C. |last3=Wang |first3=H. |last4=Huang |first4=T. |last5=Bevitt |first5=J. |last6=Li |first6=Y. |last7=Li |first7=T. |last8=Zhao |first8=Y. |last9=Li |first9=N. |year=2021 |title=Application of neutron tomography in studying new material of Ixostrobus Raciborski from the Middle Jurassic of Inner Mongolia, China |journal=Geological Journal |volume=56 |issue=9 |pages=4618–4626 |doi=10.1002/gj.4196 |s2cid=237869542 }}

|

Sp. nov

|

In press

|

Na & Sun in Na et al.

|

Middle Jurassic

|

|

{{Flag|China}}

|

A member of Czekanowskiales.

|

Kenrickia{{Cite journal|last1=Toledo |first1=S. |last2=Bippus |first2=A. C. |last3=Atkinson |first3=B. A. |last4=Bronson |first4=A. W. |last5=Tomescu |first5=A. M. F. |year=2021 |title=Taxon sampling and alternative hypotheses of relationships in the euphyllophyte plexus that gave rise to seed plants: insights from an Early Devonian radiatopsid |journal=New Phytologist |volume=232 |issue=2 |pages=914–927 |doi=10.1111/nph.17511 |pmid=34031894 |s2cid=235199240 |doi-access=free }}

|

Gen. et sp. nov

|

In press

|

Toledo et al.

|

Devonian (Emsian)

|

Battery Point Formation

|

{{Flag|Canada}}

|

An early euphyllophyte belonging to the group Radiatopses. Genus includes new species K. bivena.

|

Krommia

|

Gen. et sp. nov

|

Valid

|

Gess & Prestianni

|

Devonian (Lochkovian?)

|

Baviaanskloof Formation

|

{{Flag|Brazil}}

{{Flag|South Africa}}

|

An early polysporangiophyte. Genus includes new species K. parvapila.

|

Lejeunea abyssinicoides

|

Sp. nov

|

In press

|

Bouju et al.

|

Miocene

|

Ethiopian amber

|

{{Flag|Ethiopia}}

|

A liverwort, a species of Lejeunea.

|

Lycaugea{{Cite journal|last1=Meyer-Berthaud |first1=B. |last2=Decombeix |first2=A.-L. |last3=Blanchard |first3=R. |year=2021 |title=Lycaugea edieae gen. et sp. nov., a Late Devonian lycopsid from New South Wales, Australia |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=418–429 |doi=10.1086/714350 |s2cid=233666757 |url=https://hal.inrae.fr/hal-03231982/file/2021%20Meyer%20Berthaud%20et%20al%20Lycaugea%20.pdf }}

|

Gen. et sp. nov

|

Valid

|

Meyer-Berthaud, Decombeix & Blanchard

|

Devonian (Famennian)

|

|

{{Flag|Australia}}

|

A lycopsid. Genus includes new species L. edieae.

|

Lycopodiumsporites amazonicus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Spores of a member of the family Lycopodiaceae.

|

Melvillipteris sonidia{{Cite journal|last1=Bai |first1=L. |last2=Huang |first2=P. |last3=Yang |first3=N. |last4=Ju |first4=W. |last5=Liu |first5=J. |last6=Basinger |first6=J. F. |last7=Xu |first7=H. |last8=Xue |first8=J. |year=2021 |title=A new Late Devonian flora from Sonid Zuoqi, Inner Mongolia, northeastern China |journal=Journal of Paleontology |volume=96 |issue=2 |pages=462–484 |doi=10.1017/jpa.2021.93 |s2cid=239867911 |url=https://zenodo.org/record/5301469 }}

|

Sp. nov

|

In press

|

Bai et al.

|

Devonian (probably Famennian)

|

|

{{Flag|China}}

|

A member of Rhacophytales.

|

Mesochara adobensis{{Cite journal|last1=De Sosa Tomas |first1=A. |last2=Martín-Closas |first2=C. |last3=Vallati |first3=P. |last4=Krause |first4=M. |title=Early Cretaceous Mesochara-rich assemblages from central Patagonia, Argentina, predate the origin of homogenous Charoidean floras by about 30 million years |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105017 |doi=10.1016/j.cretres.2021.105017|issn=0195-6671 |s2cid=239707025 |hdl=2445/181533 |hdl-access=free }}

|

Sp. nov

|

In press

|

De Sosa Tomas et al.

|

Early Cretaceous

|

Los Adobes Formation

|

{{Flag|Argentina}}

|

A member of Charophyta.

|

Mesochara dobrogeica{{Cite journal|last1=Sanjuan |first1=J. |last2=Vicente |first2=A. |last3=Pérez-Cano |first3=J. |last4=Stoica |first4=M. |last5=Martín-Closas |first5=C. |year=2021 |title=Early Cretaceous charophytes from south Dobrogea (Romania). Biostratigraphy and palaeobiogeography |journal=Cretaceous Research |volume=122 |pages=Article 104762 |doi=10.1016/j.cretres.2021.104762 |bibcode=2021CrRes.12204762S |hdl=2445/181959 |s2cid=234160551 |doi-access=free |hdl-access=free }}

|

Sp. nov

|

In press

|

Sanjuan et al.

|

Early Cretaceous (Berriasian)

|

|

{{Flag|Romania}}

|

A member of Charophyta.

|

Mixoxylon{{Cite journal|last1=Chernomorets |first1=O. |last2=Sakala |first2=J. |year=2021 |title=Mixoxylon australe gen. et sp. nov., a unique homoxylous wood with non-angiosperm affinity from the Lower Cretaceous of Antarctica (Albian, James Ross Island) |journal=Antarctic Science |volume=33 |issue=5 |pages=493–501 |doi=10.1017/S0954102021000389 |bibcode=2021AntSc..33..493C |s2cid=243834316 }}

|

Gen. et sp. nov

|

In press

|

Chernomorets & Sakala

|

Early Cretaceous (Albian)

|

Whisky Bay Formation

|

Antarctica

|

A homoxylous wood of uncertain systematic affinities. Genus includes new species M. australe.

|

Mtshaelo

|

Gen. et sp. nov

|

Valid

|

Gess & Prestianni

|

Devonian (Lochkovian?)

|

Baviaanskloof Formation

|

{{Flag|South Africa}}

|

An early polysporangiophyte. Genus includes new species M. kougaensis.

|

Omniastrobus{{Cite journal|last1=Bonacorsi |first1=N. K. |last2=Gensel |first2=P. G. |last3=Hueber |first3=F. M. |last4=Leslie |first4=A. B. |year=2021 |title=Omniastrobus gen. nov., an Emsian plant with implications for the evolution of heterospory in the Early Devonian |journal=International Journal of Plant Sciences |volume=182 |issue=3 |pages=198–209 |doi=10.1086/712356 |s2cid=232050463 }}

|

Gen. et sp. nov

|

Valid

|

Bonacorsi et al.

|

Devonian (Emsian)

|

Campbellton Formation

|

{{Flag|Canada}}

|

A lycophyte. Genus includes new species O. dawsonii.

|

Palaeonitella trifurcata{{Cite journal|last1=Martín-Closas |first1=C. |last2=Segura-Altés |first2=R. |last3=Pérez-Cano |first3=J. |last4=Bover-Arnal |first4=T. |last5=Sanjuan |first5=J. |year=2021 |title=Palaeonitella trifurcata n. sp., a cortoid-building charophyte from the Lower Cretaceous of Catalonia |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104523 |doi=10.1016/j.revpalbo.2021.104523 |bibcode=2021RPaPa.29504523M |doi-access=free |hdl=2445/181500 |hdl-access=free }}

|

Sp. nov

|

Valid

|

Martín-Closas et al.

|

Early Cretaceous (Barremian–Aptian)

|

|

{{Flag|Spain}}

|

A member of Charophyta belonging to the family Characeae.

|

Paratingia wuhaia{{Cite journal|last1=Wang |first1=J. |last2=Hilton |first2=J. |last3=Pfefferkorn |first3=H. W. |last4=Wang |first4=S. |last5=Zhang |first5=Y. |last6=Bek |first6=J. |last7=Pšenička |first7=J. |last8=Seyfullah |first8=L. J. |last9=Dilcher |first9=D. |year=2021 |title=Ancient noeggerathialean reveals the seed plant sister group diversified alongside the primary seed plant radiation |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=11 |pages=e2013442118 |doi=10.1073/pnas.2013442118 | issn=0027-8424 |pmid=33836571 |pmc=7980368 |bibcode=2021PNAS..11813442W |doi-access=free }}

|

Sp. nov

|

Valid

|

Wang et al.

|

Permian (Asselian)

|

Taiyuan Formation

|

{{Flag|China}}

|

A progymnosperm belonging to the group Noeggerathiales and the family Tingiostachyaceae.

|

Permotheca? musaformis{{Cite journal |last1=Foraponova |first1=T. S. |last2=Karasev |first2=E. V. |title=Systematic problems of pollen organs of the genus Permotheca Zalessky from the Permian of Subangara |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=691–706 |doi=10.1134/S0031030121060058 |s2cid=245010226 |url=https://www.elibrary.ru/item.asp?id=46621802 |url-access=subscription }}

|

Sp. nov

|

Valid

|

Foraponova & Karasev

|

Permian

|

|

{{Flag|Russia}}

|

A pteridosperm.

|

Radula heinrichsii{{Cite journal|last1=Feldberg |first1=K. |last2=Schäfer-Verwimp |first2=A. |last3=Renner |first3=M. A. M. |last4=von Konrat |first4=M. |last5=Bechteler |first5=J. |last6=Müller |first6=P. |last7=Wang |first7=Y.-D. |last8=Schneider |first8=H. |last9=Schmidt |first9=A. R. |title=Liverworts from Cretaceous amber |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104987 |doi=10.1016/j.cretres.2021.104987 |bibcode=2021CrRes.12804987F |s2cid=238782100 }}

|

Sp. nov

|

In press

|

Feldberg et al.

|

Cretaceous

|

Burmese amber

|

{{Flag|Myanmar}}

|

A liverwort, a species of Radula.

|

Rehamnia{{Cite journal|last1=Oukassou |first1=M. |last2=Naugolnykh |first2=S. V. |year=2021 |title=Rehamnia (Lycopodiophyta), a new enigmatic Late Devonian plant from Morocco |journal=Journal of African Earth Sciences |volume=182 |pages=Article 104274 |doi=10.1016/j.jafrearsci.2021.104274 |bibcode=2021JAfES.18204274O }}

|

Gen. et sp. nov

|

In press

|

Oukassou & Naugolnykh

|

Late Devonian

|

|

{{Flag|Morocco}}

|

A member of Lycopodiophyta of uncertain phylogenetic placement. Genus includes new species R. michardis.

|

Ricciopsis baojishanensis{{Cite journal|last1=Han |first1=L. |last2=Yang |first2=T. |last3=Wang |first3=H.-J. |last4=Cai |first4=J.-H. |last5=Liang |first5=W.-Y. |last6=Bao |first6=L. |last7=Chen |first7=H.-Y. |last8=Zhang |first8=L. |last9=Li |first9=W.-J. |last10=Yan |first10=D.-F. |year=2021 |title=Liverwort fossils from the Late Triassic of Baiyin, Gansu, and their geological significance |journal=Palaeoworld |volume=in press |doi=10.1016/j.palwor.2021.09.009 |s2cid=242525785 }}

|

Sp. nov

|

In press

|

Han & Yan in Han et al.

|

Late Triassic

|

Nanying'er Formation

|

{{Flag|China}}

|

A liverwort.

|

Salopella laidae{{cite journal |last1=McSweeney |first1=F. R. |last2=Shimeta |first2=J. |last3=Buckeridge |first3=J. S. |year=2021 |title=Early land plants from the Lower Devonian of central Victoria, Australia, including a new species of Salopella |journal=Memoirs of Museum Victoria |volume=80 |pages=193–205 |doi=10.24199/j.mmv.2021.80.11 |s2cid=242026265 |doi-access=free }}

|

Sp. nov

|

Valid

|

McSweeney, Shimeta & Buckeridge

|

Devonian (Pragian)

|

Yea Formation

|

{{Flag|Australia}}

|

An early land plant of uncertain affinities.

|

Skyttegaardia{{Cite journal|last1=Friis |first1=E. M. |last2=Crane |first2=P. R. |last3=Pedersen |first3=K. R. |year=2021 |title=Microsporangiophores from the Early Cretaceous (Berriasian) of Bornholm, Denmark, with comments on a pre-angiosperm xerophytic flora |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104487 |doi=10.1016/j.revpalbo.2021.104487 |bibcode=2021RPaPa.29304487F }}

|

Gen. et sp. nov

|

In press

|

Friis, Crane & Pedersen

|

Early Cretaceous (Berriasian)

|

|

{{Flag|Denmark}}

|

A plant of uncertain phylogenetic placement, possibly close to cycads. Genus includes new species S. galtieri.

|

Thysananthus aethiopicus

|

Sp. nov

|

In press

|

Bouju et al.

|

Miocene

|

Ethiopian amber

|

{{Flag|Ethiopia}}

|

A liverwort belonging to the family Lejeuneaceae.

|

Velascoa{{Cite journal|last1=Flores Barragan |first1=M. A. |last2=Velasco de León |first2=M. P. |last3=Ortega Chavez |first3=E. |year=2021 |title=New genus for megaphyllous leaves from the Upper Paleozoic of Mexico Velascoa pueblensis gen. nov |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103408 |doi=10.1016/j.jsames.2021.103408 |bibcode=2021JSAES.11003408F }}

|

Gen. et sp. nov

|

Junior homonym

|

Flores Barragan, Velasco de León & Ortega Chavez

|

Permian

|

Matzitzi Formation

|

{{Flag|Mexico}}

|

Fossil leaves of a plant of uncertain phylogenetic placement, with a morphology similar to Ginkgophyta. Genus includes new species V. pueblensis. The generic name is preoccupied by Velascoa Calderón & Rzedowski (1997).

|

Vitinellopsis{{Cite journal|last1=Vachard |first1=D. |last2=Bucur |first2=I. I. |last3=Munnecke |first3=A. |year=2021 |title=Vitinellopsis nov. gen., a new calcareous alga (Chlorophyta, Bryopsidales) from the Silurian of Gotland (Sweden), and the tribe Vitinelleae nov. nom |journal=Geobios |volume=70 |pages=75–85 |doi=10.1016/j.geobios.2021.10.001 |s2cid=244766381 }}

|

Gen. et sp. nov

|

In press

|

Vachard, Bucur & Munnecke

|

Silurian

|

|

{{Flag|Sweden}}

|

A green alga belonging to the group Bryopsidales. Genus includes new species V. gotlandica.

|

Zosterophyllum confertum{{Cite journal|last1=Gossmann |first1=R. |last2=Poschmann |first2=M. J. |last3=Giesen |first3=P. |last4=Schultka |first4=S. |title=A stratigraphically significant new zosterophyllopsid from the Rhenish Lower Devonian (W Germany) |journal=Palaeobiodiversity and Palaeoenvironments |year=2022 |volume=102 |issue=3 |pages=503–519 |doi=10.1007/s12549-021-00509-9 |s2cid=238479823 |url=https://link.springer.com/article/10.1007/s12549-021-00509-9 |url-access=subscription }}

|

Sp. nov

|

In press

|

Gossmann et al.

|

Early Devonian

|

|

{{Flag|Germany}}

|

A zosterophyll.

|

Palynology

class="wikitable sortable" align="center" width="100%"
Name

!Novelty

!Status

!Authors

!Age

!Type locality

!Location

!Notes

!Images

Camarozonosporites trilobatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

A spore of lycopodialean affinity.

|

Echinatisporis infantulus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Erlansonisporites duwaensis{{Cite book|last1=Li |first1=W.B. |last2=Batten |first2=D. J. |last3=Li |first3=J.G. |last4=Peng |first4=J.G. |year=2021 |title=Mesozoic Megaspores and Palynomorphs from Tarim Basin, Northwest China |series=Palaeontologia Sinica |volume=202 |pages=1–250 |isbn=978-7030696526 }}

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Erlansonisporites exquisitus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Erlansonisporites perbellus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Erlansonisporites textilis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Ginkgomonocolpites{{Cite journal|last=Hernández |first=J. A. R. |year=2021 |title=A replacement name for the poorly known illegitimate Paleogene genus Psilamonocolpites Y.K.Mathur (Ginkgoales) |journal=Phytotaxa |volume=500 |issue=1 |pages=57–58 |doi=10.11646/phytotaxa.500.1.10 |s2cid=236595043 }}

|

Nom. nov

|

Valid

|

Hernández

|

Paleogene

|

|

{{Flag|India}}

|

A gymnosperm pollen; a replacement name for Psilamonocolpites Mathur (1966).

|

Hamulatisporis bareanus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Henrisporites longibaculiformis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Horstisporites comitus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Horstisporites denticulatus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Horstisporites subtilis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Horstisporites tarimensis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Hughesisporites reticulatus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Hughesisporites unicus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Ischyosporites dubius

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

A spore of dicksoniaceous affinity.

|

Ischyosporites granulatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Laevigatosporites indigestus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Luntaispora

|

Gen. et sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

Genus includes new species L. laevigata.

|

Microfoveolatosporis simplex

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Minerisporites tarimensis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Minerisporites triangularis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Narkisporites conicus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Narkisporites densibaculatus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Narkisporites densiconicus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Narkisporites tarimensis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Neoraistrickia dubia

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Noniasporites triassicus{{cite book |last1=Ghosh |first1=A. K. |last2=Chatterjee |first2=R. |last3=Pramanik |first3=S. |last4=Kar |first4=R. |year=2021 |chapter=Radiation of Flora in the Early Triassic Succeeding the End Permian Crisis: Evidences from the Gondwana Supergroup of Peninsular India |editor1=S. Banerjee |editor2=S. Sarkar |title=Mesozoic Stratigraphy of India |series=Society of Earth Scientists Series |pages=87–113 |publisher=Springer |doi=10.1007/978-3-030-71370-6_3 |isbn=978-3-030-71369-0 |s2cid=243201506 }}

|

Sp. nov

|

Valid

|

Ghosh et al.

|

Early Triassic

|

Panchet Formation

|

{{Flag|India}}

|

A megaspore.

|

Otynisporites tarimensis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Polypodiisporites discretus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Psilatriletes delicatus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Punctatosporites latrubessei

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Rotverrusporites amazonicus

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

Stellibacutriletes

|

Gen. et 4 sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

Genus includes new species S. capillaris, S. gracilis, S. rarus and S. solidus.

|

Striatriletes inconspicuus

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Tarimispora

|

Gen. et 2 sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

Genus includes new species T. auriculata and T. perfecta.

|

Tricristatispora trilobata

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Tricristatispora yingmailensis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Trileites plicatilis

|

Sp. nov

|

Valid

|

Li et al.

|

Mesozoic

|

|

{{Flag|China}}

|

|

Verrucatotriletes pseudovirueloides

|

Sp. nov

|

Valid

|

D'Apolito, Jaramillo & Harrington

|

Miocene

|

Solimões Formation

|

{{Flag|Brazil}}

|

Fossil spores.

|

=Palynological research=

  • Strother & Foster (2021) describe an assemblage of fossil spores from the Ordovician (Tremadocian) of Australia, representing a morphology that was intermediate morphology between confirmed land plant spores and earlier forms of uncertain phylogenetic placement, and evaluate the implications of these fossils for the knowledge of the evolution of land plants from their algal ancestors.{{Cite journal|last1=Strother |first1=P. K. |last2=Foster |first2=C. |year=2021 |title=A fossil record of land plant origins from charophyte algae |journal=Science |volume=373 |issue=6556 |pages=792–796 |doi=10.1126/science.abj2927 |pmid=34385396 |bibcode=2021Sci...373..792S |s2cid=236991210 }}
  • A study on the fossil pollen record from New Zealand, dating from 100 million years ago to the present, is published by Prebble et al. (2021), who report evidence indicating that Cretaceous diversification was closely followed by an increase in flowering plants frequency, but their maximum frequency did not occur until the Eocene.{{cite journal |last1=Prebble |first1=J. G. |last2=Kennedy |first2=E. M. |last3=Reichgelt |first3=T. |last4=Clowes |first4=C. |last5=Womack |first5=T. |last6=Mildenhall |first6=D. C. |last7=Raine |first7=J. I. |last8=Crouch |first8=E. M. |year=2021 |title=A 100 million year composite pollen record from New Zealand shows maximum angiosperm abundance delayed until Eocene |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=566 |pages=Article 110207 |doi=10.1016/j.palaeo.2020.110207 |bibcode=2021PPP...56610207P |s2cid=233656482 }}
  • A study on changes of abundance in spores and pollen record from the Danish Basin, and on their implications for the knowledge of the impact of the Triassic–Jurassic extinction event on land plants, is published by Lindström (2021).{{cite journal |last=Lindström |first=S. |year=2021 |title=Two-phased Mass Rarity and Extinction in Land Plants During the End-Triassic Climate Crisis |journal=Frontiers in Earth Science |volume=9 |pages=Article 780343 |doi=10.3389/feart.2021.780343 |bibcode=2021FrEaS...9.1079L |doi-access=free }}
  • A study on the vegetation history in the southwestern Balkans, as indicated by pollen from the sedimentary record in the Lake Ohrid extending to 1.36 million years ago, is published by Donders et al. (2021).{{Cite journal|last1=Donders |first1=T. |last2=Panagiotopoulos |first2=K. |last3=Koutsodendris |first3=A. |last4=Bertini |first4=A. |last5=Mercuri |first5=A. M. |last6=Masi |first6=A. |last7=Combourieu-Nebout |first7=N. |last8=Joannin |first8=S. |last9=Kouli |first9=K. |last10=Kousis |first10=I. |last11=Peyron |first11=O. |last12=Torri |first12=P. |last13=Florenzano |first13=A. |last14=Francke |first14=A. |last15=Wagner |first15=B. |last16=Sadori |first16=L. |year=2021 |title=1.36 million years of Mediterranean forest refugium dynamics in response to glacial–interglacial cycle strength |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=34 |pages=e2026111118 |doi=10.1073/pnas.2026111118 |pmid=34400496 |pmc=8403972 |bibcode=2021PNAS..11826111D |doi-access=free }}

Research

  • A study on changes of the morphological complexity of reproductive structures of land plants throughout their evolutionary history, based on data from fossil and extant land plants, is published by Leslie, Simpson & Mander (2021).{{Cite journal|last1=Leslie |first1=A. B. |last2=Simpson |first2=C. |last3=Mander |first3=L. |year=2021 |title=Reproductive innovations and pulsed rise in plant complexity |journal=Science |volume=373 |issue=6561 |pages=1368–1372 |doi=10.1126/science.abi6984 |pmid=34529461 |bibcode=2021Sci...373.1368L |s2cid=237547326 |url=http://oro.open.ac.uk/79109/1/Leslie%20et%20al%20accepted%20manuscript.pdf }}
  • Revision of Silurian (Wenlock to Přídolí) assemblages of polysporangiophytes with dispersed spores and cryptospores, aiming to determine the relationship between Silurian plant evolution and climate changes linked with perturbations of the global carbon cycle, is published by Pšenička et al. (2021).{{Cite journal|last1=Pšenička |first1=J. |last2=Bek |first2=J. |last3=Frýda |first3=J. |last4=Žárský |first4=V. |last5=Uhlířová |first5=M. |last6=Štorch |first6=P. |year=2021 |title=Dynamics of Silurian Plants as Response to Climate Changes |journal=Life |volume=11 |issue=9 |pages=Article 906 |doi=10.3390/life11090906 |pmid=34575055 |pmc=8470493 |bibcode=2021Life...11..906P |doi-access=free }}
  • Reconstruction of the structure and development of the rooting system of Asteroxylon mackiei is presented by Hetherington et al. (2021).{{Cite journal|last1=Hetherington |first1=A. J. |last2=Bridson |first2=S. L. |last3=Jones |first3=A. L. |last4=Hass |first4=H. |last5=Kerp |first5=H. |last6=Dolan |first6=L. |year=2021 |title=An evidence-based 3D reconstruction of Asteroxylon mackiei, the most complex plant preserved from the Rhynie chert |journal=eLife |volume=10 |pages=e69447 |doi=10.7554/eLife.69447 |pmid=34425940 |pmc=8384418 |doi-access=free }}
  • A study on factors influencing the extent of arboreal vegetation during the Late Paleozoic icehouse is published by Matthaeus et al. (2021), who interpret their findings as indicating that Pangaea could have supported widespread arboreal plant growth and forest cover based on leaf water constraints, but the forest extent was restricted because of impact of freezing on plants, and estimate that contracting forest cover increased net global surface runoff by up to 6.1%.{{Cite journal|last1=Matthaeus |first1=W. J. |last2=Macarewich |first2=S. I. |last3=Richey |first3=J. D. |last4=Wilson |first4=J. P. |last5=McElwain |first5=J. C. |last6=Montañez |first6=I. P. |last7=DiMichele |first7=W. A. |last8=Hren |first8=M. T. |last9=Poulsen |first9=C. J. |last10=White |first10=J. D. |year=2021 |title=Freeze tolerance influenced forest cover and hydrology during the Pennsylvanian |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=42 |pages=e2025227118 |doi=10.1073/pnas.2025227118 |pmid=34635589 |pmc=8594576 |bibcode=2021PNAS..11825227M |doi-access=free }}
  • Description of the reproductive organs of the lycopsids from the Upper Devonian Wutong Formation (China), and a study on the ability of the sporophyll units for wind dispersal, is published by Zhou et al. (2021), who name new form species Lepidophylloides longshanensis and Lepidophylloides changxingensis.{{Cite journal|last1=Zhou |first1=Y. |last2=Wang |first2=D.-M. |last3=Liu |first3=L. |last4=Huang |first4=P. |year=2021 |title=The morphometric of lycopsid sporophylls and the evaluation of their dispersal potential: an example from the Upper Devonian of Zhejiang Province, China |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 198 |doi=10.1186/s12862-021-01933-3 |pmid=34732141 |pmc=8565055 |doi-access=free }}
  • An exceptionally well preserved Brasilodendron-like lycopsid forest containing over 150 upright stumps is described from an early Permian postglacial landscape of western Gondwana (Paraná Basin, Brazil) by Mottin et al. (2021).{{Cite journal|last1=Mottin |first1=T. E. |last2=Iannuzzi |first2=R. |last3=Vesely |first3=F. F. |last4=Montañez |first4=I. P. |last5=Griffis |first5=N. |last6=Canata |first6=R. E. |last7=Barão |first7=L. M. |last8=da Silveira |first8=D. M. |last9=Garcia |first9=A. M. |year=2021 |title=A glimpse of a Gondwanan postglacial fossil forest |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=588 |pages=Article 110814 |doi=10.1016/j.palaeo.2021.110814 |s2cid=245582665 }}
  • A study on the anatomy of Stigmaria asiatica is published by Chen et al. (2021).{{Cite journal|last1=Chen |first1=B.-Y. |last2=Wan |first2=M.-L. |last3=Zhou |first3=W.-M. |last4=Wang |first4=S.-J. |last5=Wang |first5=J. |year=2021 |title=Anatomy of Stigmaria asiatica Jongmans et Gothan from the Asselian (lowermost Permian) of Wuda Coalfield, Inner Mongolia, North China |journal=Palaeoworld |volume=31 |issue=2 |pages=311–323 |doi=10.1016/j.palwor.2021.05.001| issn=1871-174X |s2cid=236366665 }}
  • Stump casts of Sigillaria, preserving traces of internal anatomy, are described from the earliest Permian Wuda Tuff (China) by D'Antonio et al. (2021).{{Cite journal|last1=D'Antonio |first1=M. P. |last2=Boyce |first2=C. K. |last3=Zhou |first3=W.-M. |last4=Pfefferkorn |first4=H. W. |last5=Wang |first5=J. |title=Primary tissues dominated ground-level trunk diameter in Sigillaria: evidence from the Wuda Tuff, Inner Mongolia |year=2021 |journal=Journal of the Geological Society |volume=179 |issue=2 |doi=10.1144/jgs2021-021 |s2cid=238701484 }}
  • A study aiming to determine probable causes of the world-wide proliferation of members of Isoetales, particularly Pleuromeia, during and in the aftermath of the Permian–Triassic extinction event, and evaluating the implications of this proliferation for the knowledge of environmental stresses during and in the aftermath of this extinction event, is published by Looy, van Konijnenburg-van Cittert & Duijnstee (2021).{{Cite journal|last1=Looy |first1=C. V. |last2=van Konijnenburg-van Cittert |first2=J. H. A. |last3=Duijnstee |first3=I. A. P. |year=2021 |title=Proliferation of Isoëtalean Lycophytes During the Permo-Triassic Biotic Crises: A Proxy for the State of the Terrestrial Biosphere |journal=Frontiers in Earth Science |volume=9 |pages=Article 615370 |doi=10.3389/feart.2021.615370 |bibcode=2021FrEaS...9...55L |doi-access=free }}
  • New fossil material of Saportaea salisburioides, providing new information on leaf morphology and growth of this plant, is described from the Permian Umm Irna Formation (Jordan) by Kerp et al. (2021), who interpret their findings as indicating that Saportaea grandifolia and Baiera virginiana were synonyms of S. salisburioides, and possibly indicating that the fructification belonging to the genus Nystroemia is a part of Saportaea.{{Cite journal|last1=Kerp |first1=H. |last2=Blomenkemper |first2=P. |last3=Hamad |first3=A. A. |last4=Bomfleur |first4=B. |year=2021 |title=Saportaea Fontaine et White 1880 – An enigmatic, long-ranging, widely distributed but rare type of late Palaeozoic and early Mesozoic foliage |journal=Review of Palaeobotany and Palynology |volume=296 |pages=Article 104542 |doi=10.1016/j.revpalbo.2021.104542 |s2cid=244595344 }}
  • Description of Geinitzia reichenbachii from its gross morphology to the cellular scale, and a study on the likely ecology of this conifer, is published by Moreau et al. (2021).{{Cite journal|last1=Moreau |first1=J.-D. |last2=Philippe |first2=M. |last3=Néraudeau |first3=D. |last4=Dépré |first4=E. |last5=Le Couls |first5=M. |last6=Fernandez |first6=V. |last7=Beurel |first7=S. |year=2021 |title=Paleohistology of the Cretaceous resin-producing conifer Geinitzia reichenbachii using X-ray synchrotron microtomography |journal=American Journal of Botany |volume=108 |issue=9 |pages=1745–1760 |doi=10.1002/ajb2.1722 |pmid=34495546 |s2cid=237441544 |doi-access=free }}
  • A study on the evolutionary history of the family Cycadaceae, based on genomic data and fossil record, is published by Liu et al. (2021).{{Cite journal|last1=Liu |first1=J. |last2=Lindstrom |first2=A. J. |last3=Marler |first3=T. E. |last4=Gong |first4=X. |year=2021 |title=Not that young: combining plastid phylogenomic, plate tectonic and fossil evidence indicates a Palaeogene diversification of Cycadaceae |journal=Annals of Botany |volume=129 |issue=2 |pages=217–230 |doi=10.1093/aob/mcab118 |pmid=34520529 |pmc=8796677 |doi-access=free }}
  • Well-preserved recurved cupules of seed plants are described from the Lower Cretaceous of China by Shi et al. (2021), who interpret the structure of these cupules as consistent with the recurved form and development of the second integument in the bitegmic anatropous ovules of flowering plants, and evaluate the implications of these fossils for the knowledge of the origin of the flowering plants.{{Cite journal|last1=Shi |first1=G. |last2=Herrera |first2=F. |last3=Herendeen |first3=P. S. |last4=Clark |first4=E. G. |last5=Crane |first5=P. R. |year=2021 |title=Mesozoic cupules and the origin of the angiosperm second integument |journal=Nature |volume=594 |issue=7862 |pages=223–226 |doi=10.1038/s41586-021-03598-w |pmid=34040260 |bibcode=2021Natur.594..223S |s2cid=235217720 }}
  • Taxonomically diverse flora from the Seafood Salad locality, found ~65 m below the Cretaceous-Paleogene boundary in the Hell Creek Formation (Montana, United States), is described by Wilson, Wilson Mantilla & Strӧmberg (2021), who study the affinities of plants of this locality and compare them with other Late Cretaceous floras of the Western Interior.{{Cite journal|last1=Wilson |first1=P. K. |last2=Wilson Mantilla |first2=G. P. |last3=Strӧmberg |first3=C. A. E. |year=2021 |title=Seafood Salad: A diverse latest Cretaceous flora from eastern Montana |journal=Cretaceous Research |volume=121 |pages=Article 104734 |doi=10.1016/j.cretres.2020.104734 |bibcode=2021CrRes.12104734W |s2cid=233543523 }}
  • A study on the timing of the origin of the flowering plants, based on data from fossil record and from the diversity of extant members of this group, is published by Silvestro et al. (2021), who interpret their findings as indicating that several flowering plant families originated in the Jurassic.{{Cite journal|last1=Silvestro |first1=D. |last2=Bacon |first2=C. D. |last3=Dong |first3=W. |last4=Zhang |first4=Q. |last5=Donoghue |first5=P. C. J. |last6=Antonelli |first6=A. |last7=Xing |first7=Y. |year=2021 |title=Fossil data support a pre-Cretaceous origin of flowering plants |journal=Nature Ecology & Evolution |volume=5 |issue=4 |pages=449–457 |doi=10.1038/s41559-020-01387-8 |pmid=33510432 |hdl=1983/11314812-19f6-4cbc-84a2-28791cb16d53 |s2cid=231763334 |url=https://research-information.bris.ac.uk/ws/files/264149147/Silvestro_et_al_AF.pdf }}
  • A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).{{cite journal |last1=Xiao |first1=L. |last2=Labandeira |first2=C. |last3=Dilcher |first3=D. |last4=Ren |first4=D. |year=2021 |title=Florivory of Early Cretaceous flowers by functionally diverse insects: implications for early angiosperm pollination |journal=Proceedings of the Royal Society B: Biological Sciences |volume=288 |issue=1953 |pages=Article ID 20210320 |doi=10.1098/rspb.2021.0320 |pmid=34132112 |pmc=8207559 }}
  • New fossil material of Callianthus dilae is described from the Lower Cretaceous Yixian Formation (China) by Wang et al. (2021), who reconstruct the whole plant of Callianthus, interpreting it as an aquatic flowering plant.{{Cite journal|last1=Wang |first1=X. |last2=Shih |first2=C. |last3=Liu |first3=Z.-J. |last4=Lin |first4=L. |last5=Singh |first5=K. J. |year=2021 |title=Reconstructing the Callianthus plant – an early aquatic angiosperm from the Lower Cretaceous of China |journal=Cretaceous Research |volume=128 |pages=Article 104983 |doi=10.1016/j.cretres.2021.104983|bibcode=2021CrRes.12804983W |issn=0195-6671 |doi-access=free }}
  • A study on the anatomy of the epidermal features of the floating leaves of Quereuxia angulata from the Upper Cretaceous Yong'ancun Formation (China) is published by Liang et al. (2021).{{Cite journal|last1=Liang |first1=F. |last2=Tian |first2=N. |last3=Sun |first3=W. |last4=Wu |first4=Q. |last5=Liu |first5=B. |last6=Wang |first6=H. |year=2021 |title=Epidermal features of the floating leaves of Quereuxia angulata (Newberry) Krištofovič, an aquatic angiosperm from the Upper Cretaceous of Northeast China |journal=Cretaceous Research |volume=125 |pages=Article 104835 |doi=10.1016/j.cretres.2021.104835 |bibcode=2021CrRes.12504835L }}
  • A study on plant extinction and ecological change in tropical forests resulting from the Cretaceous–Paleogene extinction event, based on data from fossil pollen and leaves from Colombia, is published by Carvalho et al. (2021), who report evidence indicative of a long interval of low plant diversity in the Neotropics after the end-Cretaceous extinction, and the emergence of forests with a structure resembling modern Neotropical rainforests, with a closed canopy and multistratal structure dominated by flowering plants, during the Paleocene.{{Cite journal|last1=Carvalho |first1=M. R |last2=Jaramillo |first2=C. |last3=de la Parra |first3=F. |last4=Caballero-Rodríguez |first4=D. |last5=Herrera |first5=F. |last6=Wing |first6=S. |last7=Turner |first7=B. L. |last8=D'Apolito |first8=C. |last9=Romero-Báez |first9=M. |last10=Narváez |first10=P. |last11=Martínez |first11=C. |last12=Gutierrez |first12=M. |last13=Labandeira |first13=C. |last14=Bayona |first14=G. |last15=Rueda |first15=M. |last16=Paez-Reyes |first16=M. |last17=Cárdenas |first17=D. |last18=Duque |first18=Á. |last19=Crowley |first19=J. L. |last20=Santos |first20=C. |last21=Silvestro |first21=D. |year=2021 |title=Extinction at the end-Cretaceous and the origin of modern Neotropical rainforests |journal=Science |volume=372 |issue=6537 |pages=63–68 |doi=10.1126/science.abf1969 |pmid=33795451 |bibcode=2021Sci...372...63C |s2cid=232484243 }}
  • A study on the impact of the mid-Eocene greenhouse warming event on floras from southernmost South America is published by Fernández et al. (2021).{{Cite journal|last1=Fernández |first1=D. A. |last2=Palazzesi |first2=L. |last3=González Estebenet |first3=M. S. |last4=Tellería |first4=M. C. |last5=Barreda |first5=V. D. |title=Impact of mid Eocene greenhouse warming on America's southernmost floras |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 176 |doi=10.1038/s42003-021-01701-5 |pmid=33564110 |pmc=7873257 }}
  • Evidence from middle Eocene-middle Miocene tuffaceous deposits of central and northern Patagonia, indicating that soils, vegetation, insects and mammal herbivores began to record diverse traits related to the presence of grasslands with mosaic vegetation since middle Eocene, is presented by Bellosi et al. (2021).{{Cite journal|last1=Bellosi |first1=E. |last2=Genise |first2=J. F. |last3=Zucol |first3=A. |last4=Bond |first4=M. |last5=Kramarz |first5=A. |last6=Sánchez |first6=M. V. |last7=Krause |first7=J. M. |title=Diverse evidence for grasslands since the Eocene in Patagonia |year=2021 |journal=Journal of South American Earth Sciences |volume=108 |pages=Article 103357 |doi=10.1016/j.jsames.2021.103357 |bibcode=2021JSAES.10803357B |s2cid=235567426 }}
  • A study on Middle Miocene microfloral assemblages from ten localities in the Madrid Basin (Spain), providing evidence of prevalence of open habitats with grass-dominated, savannah-like vegetation under a warm and semi-arid climatic regime in the Iberian Peninsula in the Middle Miocene, is published by Casas-Gallego et al. (2021).{{Cite journal|last1=Casas-Gallego |first1=M. |last2=Postigo-Mijarra |first2=J. M. |last3=Rivas-Carballo |first3=M. R. |last4=Valle-Hernández |first4=M. F. |last5=Morín-de Pablos |first5=J. |last6=Barrón |first6=E. |title=Early evidence of continental aridity and open-habitat grasslands in Europe as revealed by the Middle Miocene microflora of the Madrid Basin |year=2021 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=581 |pages=Article 110603 |doi=10.1016/j.palaeo.2021.110603 |bibcode=2021PPP...58110603C |s2cid=238847370 }}
  • Crump et al. (2021) present a record of vegetation from the Last Interglacial based on ancient DNA from lake sediment from the Baffin Island (Canada), and report evidence of major ecosystem changes in the Arctic in response to warmth, including a ~400 km northward range shift of dwarf birch relative to today.{{Cite journal|last1=Crump |first1=S. E. |last2=Fréchette |first2=B. |last3=Power |first3=M. |last4=Cutler |first4=S. |last5=de Wet |first5=G. |last6=Raynolds |first6=M. K. |last7=Raberg |first7=J. H. |last8=Briner |first8=J. P. |last9=Thomas |first9=E. K. |last10=Sepúlveda |first10=J. |last11=Shapiro |first11=B. |last12=Bunce |first12=M. |last13=Miller |first13=G. H. |year=2021 |title=Ancient plant DNA reveals High Arctic greening during the Last Interglacial |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=13 |pages=e2019069118 |doi=10.1073/pnas.2019069118 |pmid=33723011 |pmc=8020792 |bibcode=2021PNAS..11819069C |doi-access=free }}

Deaths

  • Alan Graham (1934–2021), passed away on 8 July 2021. Graham earned his PhD in 1962 under the guidance of Chester A. Arnold, and was noted for a career studying the Cenozoic paleobotany of the Caribbean and Central America. {{cite journal |last1=Jaramillo |first1=C. |last2=Jarzen |first2=D. M. |title=Alan Keith Graham (1934–2021) |journal=Palynology |date=2022 |volume=46 |issue=1 |pages=1–4 |doi=10.1080/01916122.2021.1971121 |bibcode=2022Paly...4671121J |s2cid=238700678 |doi-access=free }}

References

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Category:2021 in paleontology

Category:Paleobotany