2021 in paleomalacology
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This list of fossil molluscs described in 2021 is a list of new taxa of fossil molluscs that were described during the year 2021, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2021.
Ammonites
=New taxa=
=Research=
- A modern review of the palaeobiology of heteromorph ammonoids is published by Hoffmann et al. (2021), including details of their anatomy, buoyancy, locomotion, predators, diet, palaeoecology, and extinction.{{Cite journal|last1=Hoffmann |first1=R. |last2=Slattery |first2=J. S. |last3=Kruta |first3=I. |last4=Linzmeier |first4=B. J. |last5=Lemanis |first5=R. E. |last6=Mironenko |first6=A. |last7=Goolaerts |first7=S. |last8=De Baets |first8=K. |last9=Peterman |first9=D. J. |last10=Klug |first10=C. |year=2021 |title=Recent advances in heteromorph ammonoid palaeobiology |journal=Biological Reviews |volume=96 |issue=2 |pages=576–610 |doi=10.1111/brv.12669 |doi-access=free |pmid=33438316 }}
- A study on the impact of the Permian–Triassic extinction event on the morphological diversity of ammonites is published by Dai, Korn & Song (2021).{{Cite journal |last1=Dai |first1=X. |last2=Korn |first2=D. |last3=Song |first3=H. |title=Morphological selectivity of the Permian-Triassic ammonoid mass extinction |year=2021 |journal=Geology |volume=49 |issue=9 |pages=1112–1116 |doi=10.1130/G48788.1 |bibcode=2021Geo....49.1112D |s2cid=236274865 |doi-access=free }}
- A study on the morphology and ontogeny of whorl profiles of the late Anisian ceratitid ammonites is published by Bischof et al. (2021).{{Cite journal|last1=Bischof |first1=E. A. |last2=Schlüter |first2=N. |last3=Korn |first3=D. |last4=Lehmann |first4=J. |year=2021 |title=Ontogeny of highly variable ceratitid ammonoids from the Anisian (Middle Triassic) |journal=PeerJ |volume=9 |pages=e10931 |doi=10.7717/peerj.10931 |pmid=33717689 |pmc=7934684 |doi-access=free }}
- An exceptionally preserved specimen of Sigaloceras enodatum, providing information on the anatomy of ammonite muscles, is described from the Middle Jurassic (Callovian) Kellaways Sand Member of Colne Gravel (Gloucestershire, United Kingdom) by Cherns et al. (2021), who evaluate the implications of this specimen for the knowledge of the swimming mechanism of ammonites.{{Cite journal|last1=Cherns |first1=L. |last2=Spencer |first2=A. R. T. |last3=Rahman |first3=I. A. |last4=Garwood |first4=R. J. |last5=Reedman |first5=C. |last6=Burca |first6=G. |last7=Turner |first7=M. J. |last8=Hollingworth |first8=N. T. J. |last9=Hilton |first9=J. |year=2021 |title=Correlative tomography of an exceptionally preserved Jurassic ammonite implies hyponome-propelled swimming |journal=Geology |volume=50 |issue=4 |pages=397–401 |doi=10.1130/G49551.1 |hdl=10141/622963 |s2cid=245051452 |url=http://pure-oai.bham.ac.uk/ws/files/155784759/Cherns_et_al_2021_Author_Accepted_Manuscript.pdf }}
- Soft parts of a perisphinctid belonging to the genus Subplanites, separated from the conch either taphonomically or during a failed predation, are described from the Tithonian conservation deposits of Wintershof (southern Germany) by Klug et al. (2021).{{Cite journal|last1=Klug |first1=C. |last2=Schweigert |first2=G. |last3=Tischlinger |first3=H. |last4=Pochmann |first4=H. |year=2021 |title=Failed prey or peculiar necrolysis? Isolated ammonite soft body from the Late Jurassic of Eichstätt (Germany) with complete digestive tract and male reproductive organs |journal=Swiss Journal of Palaeontology |volume=140 |issue=1 |pages=Article 3 |doi=10.1186/s13358-020-00215-7 |pmid=33505352 |pmc=7813712 |doi-access=free }}
- The first physical compression experiments on model ammonite septa are performed by Johnson et al. (2021), who attempt to determine whether the increasingly complex fractal margins of ammonite septa might have evolved to increase resistance to shell-crushing predators.{{Cite journal|last1=Johnson |first1=E. H. |last2=DiMarco |first2=B. M. |last3=Peterman |first3=D. J. |last4=Carter |first4=A. M. |last5=Allmon |first5=W. D. |year=2021 |title=Did shell-crushing predators drive the evolution of ammonoid septal shape? |journal=Paleobiology |volume=47 |issue=4 |pages=666–679 |doi=10.1017/pab.2021.13 |bibcode=2021Pbio...47..666J |s2cid=234854806 }}
- A study aiming to determine the function of complex septa in the ammonite shell, providing evidence of a relationship between septal complexity and capacity for liquid retention within chambers of the shell, likely improving buoyancy regulation, is published by Peterman et al. (2021).{{Cite journal|last1=Peterman |first1=D. J. |last2=Ritterbush |first2=K. A. |last3=Ciampaglio |first3=C. N. |last4=Johnson |first4=E. H. |last5=Inoue |first5=S. |last6=Mikami |first6=T. |last7=Linn |first7=T. J. |title=Buoyancy control in ammonoid cephalopods refined by complex internal shell architecture |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 8055 |doi=10.1038/s41598-021-87379-5 |pmid=33850189 |pmc=8044186 |bibcode=2021NatSR..11.8055P }}
- Two specimens of Peltoceras athleta having both female and male features are described from Callovian limestone beds in Méron near Montreuil-Bellay (France) by Frau & Boursicot (2021), who interpret these specimens as cases of intersexuality in ammonites.{{Cite journal|last1=Frau |first1=C. |last2=Boursicot |first2=P.-Y. |year=2021 |title=Another lesson from beautiful monsters: the case of 'sex reversals' in the Ammonoidea and their significance |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 133 |doi=10.1186/s12862-021-01857-y |pmid=34174830 |pmc=8236137 |doi-access=free }}
- Soft tissue remains preserved within the conch of a specimen of Proleymeriella schrammeni from the Albian of Germany are described by Hoffmann et al. (2021).{{Cite journal|last1=Hoffmann |first1=R. |last2=Morón-Alfonso |first2=D. |last3=Klug |first3=C. |last4=Tanabe |first4=K. |year=2021 |title=Ammonoid soft tissue remains revealed by computed tomography |journal=Swiss Journal of Palaeontology |volume=140 |issue=1 |pages=Article 11 |doi=10.1186/s13358-021-00226-y |doi-access=free |hdl=11336/135617 |hdl-access=free }}
- A study on the arrangement in space and probable nature of hook-like structures found with specimens of Rhaeboceras halli is published by Smith et al. (2021), who interpret these structures as representing part of the brachial crown armature, and propose a hypothetical reconstruction of an ammonite brachial crown.{{Cite journal|last1=Smith |first1=C. P. A. |last2=Landman |first2=N. H. |last3=Bardin |first3=J. |last4=Kruta |first4=I. |title=New evidence from exceptionally "well-preserved" specimens sheds light on the structure of the ammonite brachial crown |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 11862 |doi=10.1038/s41598-021-89998-4 |doi-access=free |pmid=34088905 |pmc=8178333 |bibcode=2021NatSR..1111862S }}
- A study on the systematics and phylogeny of the genus Aegocrioceras is published by Weinkauf, Hoffmann & Wiedenroth (2021).{{Cite journal|last1=Weinkauf |first1=M. F. G. |last2=Hoffmann |first2=R. |last3=Wiedenroth |first3=K. |year=2021 |title=Evolutionary–phylogenetic pathway of the Cretaceous ammonite genus Aegocrioceras and its relationship to Juddiceras spp. and Crioceratites spp. |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=2113–2139 |doi=10.1002/spp2.1397 |s2cid=238723092 |doi-access=free }}
- A study on the hydrostatic properties of Baculites compressus, and on their implications for the knowledge of the swimming capabilities and ecology of this ammonite and orthoconic cephalopods in general, is published by Peterman & Ritterbush (2021).{{Cite journal|last1=Peterman |first1=D. J. |last2=Ritterbush |first2=K. A. |year=2021 |title=Vertical escape tactics and movement potential of orthoconic cephalopods |journal=PeerJ |volume=9 |pages=e11797 |doi=10.7717/peerj.11797 |doi-access=free |pmid=34316410 |pmc=8288114 }}
- Revision of the fossil material of Parapuzosia seppenradensis, including historic specimens from Europe and the United States and new material from England and Mexico, and a study on the ontogeny, distribution and evolution of this ammonite, is published by Ifrim et al. (2021).{{Cite journal|last1=Ifrim |first1=C. |last2=Stinnesbeck |first2=W. |last3=González González |first3=A. H. |last4=Schorndorf |first4=N. |last5=Gale |first5=A. S. |year=2021 |title=Ontogeny, evolution and palaeogeographic distribution of the world's largest ammonite Parapuzosia (P.) seppenradensis (Landois, 1895) |journal=PLOS ONE |volume=16 |issue=11 |pages=e0258510 |doi=10.1371/journal.pone.0258510 |pmid=34758037 |pmc=8580234 |bibcode=2021PLoSO..1658510I |doi-access=free }}
- A study on the habitat depth preferences of Hoploscaphites constrictus from the European Boreal Chalk Sea, as indicated by stable isotope thermometry of aptychi and by data from predation marks on the scaphitid moulds, is published by Machalski et al. (2021).{{Cite journal|last1=Machalski |first1=M. |last2=Owocki |first2=K. |last3=Dubicka |first3=Z. |last4=Malchyk |first4=O. |last5=Wierny |first5=W. |title=Stable isotopes and predation marks shed new light on ammonoid habitat depth preferences |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 22730 |doi=10.1038/s41598-021-02236-9 |pmid=34815505 |pmc=8611083 |bibcode=2021NatSR..1122730M |doi-access=free }}
- A study on the relationship between sutural complexity and body size in Cretaceous ammonites is published by Pérez-Claros (2021).{{Cite journal |last=Pérez-Claros |first=J. A. |title=Sutural complexity and body size in Cretaceous ammonoids: macroevolutionary decoupling of correlated features |year=2021 |journal=Cretaceous Research |volume=132 |pages=Article 105120 |doi=10.1016/j.cretres.2021.105120 |s2cid=245422498 |doi-access=free |hdl=10630/24004 |hdl-access=free }}
Other cephalopods
=New taxa=
=Research=
- New fossil material with shell features that characterize early cephalopods is described from the Cambrian (Terreneuvian) Bonavista Formation (Canada) by Hildenbrand et al. (2021), who interpret this finding as possibly representing the earliest cephalopods known to date, potentially pushing the origin of cephalopods back in time by about 30 million years, to the time before the first occurrence of euarthropods.{{Cite journal|last1=Hildenbrand |first1=A. |last2=Austermann |first2=G. |last3=Fuchs |first3=D. |last4=Bengtson |first4=P. |last5=Stinnesbeck |first5=W. |title=A potential cephalopod from the early Cambrian of eastern Newfoundland, Canada |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 388 |doi=10.1038/s42003-021-01885-w |pmid=33758350 |pmc=7987959 |s2cid=232338352 |doi-access=free }}
- Two specimens of Syrionautilus libanoticus preserving soft parts, including a specimen showing an imprint of the conch, are described from the Cenomanian of Hadjoula (Lebanon) by Klug et al. (2021).{{Cite journal|last1=Klug |first1=C. |last2=Pohle |first2=A. |last3=Roth |first3=R. |last4=Hoffmann |first4=R. |last5=Wani |first5=R. |last6=Tajika |first6=A. |year=2021 |title=Preservation of nautilid soft parts inside and outside the conch interpreted as central nervous system, eyes, and renal concrements from the Lebanese Cenomanian |journal=Swiss Journal of Palaeontology |volume=140 |issue=1 |pages=Article 15 |doi=10.1186/s13358-021-00229-9 |pmid=34721283 |pmc=8549922 |doi-access=free }}
- A study on the diversity patterns and spatial structure of belemnite assemblages in Western Tethys Ocean during the Early Jurassic is published by Neige, Weis & Fara (2021).{{Cite journal|last1=Neige |first1=P. |last2=Weis |first2=R. |last3=Fara |first3=E. |year=2021 |title=Ups and downs of belemnite diversity in the Early Jurassic of Western Tethys |journal=Palaeontology |volume=64 |issue=2 |pages=263–283 |doi=10.1111/pala.12522 |bibcode=2021Palgy..64..263N |doi-access=free }}
- A study on morphological changes of belemnites across the Pliensbachian-Toarcian boundary event, based on data from the Peniche GSSP section (Portugal), is published by Nätscher et al. (2021).{{Cite journal|last1=Nätscher |first1=P. S. |last2=Dera |first2=G. |last3=Reddin |first3=C. J. |last4=Rita |first4=P. |last5=De Baets |first5=K. |title=Morphological response accompanying size reduction of belemnites during an Early Jurassic hyperthermal event modulated by life history |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 14480 |doi=10.1038/s41598-021-93850-0 |doi-access=free |pmid=34262074 |pmc=8280180 |bibcode=2021NatSR..1114480N }}
- A study on chemistry, organization and genesis of circular structures (superficially resembling chromatophores) preserved in coleoid cephalopod specimens from the Jurassic of Germany and the Cretaceous of Lebanon is published by Klug et al. (2021).{{Cite journal|last1=Klug |first1=C. |last2=Di Silvestro |first2=G. |last3=Hoffmann |first3=R. |last4=Schweigert |first4=G. |last5=Fuchs |first5=D. |last6=Clements |first6=T. |last7=Gueriau |first7=P. |year=2021 |title=Taphonomic patterns mimic biologic structures: diagenetic Liesegang rings in Mesozoic coleoids and coprolites |journal=PeerJ |volume=9 |pages=e10703 |doi=10.7717/peerj.10703 |pmid=33520466 |pmc=7811783 |doi-access=free }}
- A well-preserved soft-body imprint of a fossil squid is described from the lower Oligocene of the Krasnodar region (Russia) by Mironenko et al. (2021), who interpret this finding as the first unquestionable representative of Teuthida in the fossil record known to date, and report preservation of fish remains in the stomach contents of this squid, indicative of its piscivorous diet.{{Cite journal|last1=Mironenko |first1=A. A. |last2=Boiko |first2=M. S. |last3=Bannikov |first3=A. F. |last4=Arkhipkin |first4=A. I. |last5=Bizikov |first5=V. A. |last6=Košťák |first6=M. |year=2021 |title=First discovery of the soft-body imprint of an Oligocene fossil squid indicates its piscivorous diet |journal=Lethaia |volume=54 |issue=5 |pages=793–805|doi=10.1111/let.12440 |s2cid=238646997 }}
- A specimen of Jeletzkyteuthis coriacea holding a specimen of Parabelopeltis flexuosa in its arms, with the apex very close to its jaws, is described from the Toarcian Posidonia Shale (Germany) by Klug et al. (2021), who interpret this finding as a case of frozen predatory behaviour of two different octobrachian species.{{Cite journal |last1=Klug |first1=C. |last2=Schweigert |first2=G. |last3=Fuchs |first3=D. |last4=De Baets |first4=K. |year=2021 |title=Distraction sinking and fossilized coleoid predatory behaviour from the German Early Jurassic |journal=Swiss Journal of Palaeontology |volume=140 |issue=1 |pages=Article 7 |doi=10.1186/s13358-021-00218-y |s2cid=232237598 |pmid=33815267 |pmc=7965854 |doi-access=free }}
- Redescription of the gladius of the holotype specimen of Necroteuthis hungarica is published by Košťák et al. (2021), who reinterpret this species as a member of the family Vampyroteuthidae linking Mesozoic loligosepiids with extant Vampyroteuthis.{{Cite journal|last1=Košťák |first1=M. |last2=Schlögl |first2=J. |last3=Fuchs |first3=D. |last4=Holcová |first4=K. |last5=Hudáčková |first5=N. |last6=Culka |first6=A. |last7=Fözy |first7=I. |last8=Tomašových |first8=A. |last9=Milovský |first9=R. |last10=Šurka |first10=J. |last11=Mazuch |first11=M. |title=Fossil evidence for vampire squid inhabiting oxygen-depleted ocean zones since at least the Oligocene |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 216 |doi=10.1038/s42003-021-01714-0 |pmid=33603225 |pmc=7893013 }}
- Klompmaker & Landman (2021) describe the oldest drill holes produced by octopodoids in lucinid bivalve specimens from the Campanian of South Dakota, indicating that the drilling habit evolved early in the evolutionary history of Octopodoidea.{{Cite journal|last1=Klompmaker |first1=A. A. |last2=Landman |first2=N. H. |title=Octopodoidea as predators near the end of the Mesozoic Marine Revolution |journal=Biological Journal of the Linnean Society |year=2021 |volume=132 |issue=4 |pages=894–899 |doi=10.1093/biolinnean/blab001 |doi-access=free }}
Gastropods
=New taxa=
| class="wikitable sortable" align="center" width="100%" |
| Name
! Novelty ! Status ! Authors ! Age ! Type locality ! Country ! Notes ! Images |
|---|
| Acirsa ambrosii{{Cite journal|last1=Brunetti |first1=M. M. |last2=Cresti |first2=M. |title=Tre nuove specie di Gasteropodi per il Pliocene di Orciano Pisano |year=2021 |journal=Bollettino Malacologico |volume=57 |issue=1 |pages=106–113 }}
| Sp. nov | Valid | Brunetti & Cresti | | | {{Flag|Italy}} | A species of Acirsa. | |
| Acirsa (Notacirsa) texana
| Sp. nov | Valid | Garvie | | | {{Flag|United States}} | A species of Acirsa. | |
| Acteon pseudotypica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Acteon. | |
| Acutitomaria{{Cite journal |last1=Karapunar |first1=B. |last2=Nützel |first2=A. |title=Slit-band gastropods (Pleurotomariida) from the Upper Triassic St. Cassian Formation and their diversity dynamics in the Triassic |year=2021 |journal=Zootaxa |volume=5042 |issue=1 |pages=1–165 |doi=10.11646/zootaxa.5042.1.1 |pmid=34811018 |s2cid=244213844 }}
| Gen. et sp. nov | Valid | Karapunar & Nützel | | | {{Flag|Italy}} | A member of Pleurotomariida. Genus includes new species A. kustatscherae. | |
| Alatostrombus{{Cite journal|last=Raven |first=J. G. M. |title=Notes on molluscs from NW Borneo 8. Stromboidea (Gastropoda, Caenogastropoda). Addendum, including the description of two new genera and new extinct species |year=2021 |journal=Vita Malacologica |volume=20 |pages=23–50 }}
| Gen. et sp. et comb. nov | Valid | Raven | Miocene | | {{Flag|Indonesia}} {{Flag|Malaysia}} | A member of the family Strombidae. Genus includes new species A. acutalatus, as well as "Strombus" sedanensis Martin (1899). | |
| Alvania absuturalis{{cite book |last1=Chirli |first1=C. |last2=Forli |first2=M. |year=2021 |title=The Family Rissoidae Gray, 1847 from Miocene to Present-day in the Mediterranean Basin |pages=1–466 |publisher=Edizioni Danaus |isbn=978-8-89-760331-3 }}
| Sp. nov | Valid | Chirli & Forli | Pliocene | | {{Flag|Italy}} | A species of Alvania. | |
| Alvania ameliae
| Sp. nov | Valid | Chirli & Forli | Pleistocene | | {{Flag|Italy}} | A species of Alvania. | |
| Alvania benestarensis
| Sp. nov | Valid | Chirli & Forli | Miocene | | {{Flag|Italy}} | A species of Alvania. | |
| Alvania cerreti{{cite journal|last1=Gardella |first1=F. |last2=Bertaccini |first2=E. |last3=Bertamini |first3=R. |last4=Bongiardino |first4=C. |last5=Petracci |first5=P. |last6=Tabanelli |first6=C. |year=2021 |title=The malacofauna of the "spungone": Alvania cerreti n. sp. |journal=Quaderno di Studi e Notizie di Storia Naturale della Romagna |volume=53 |pages=19–25 |url=https://www.researchgate.net/publication/352462652 }}
| Sp. nov | Valid | Gardella et al. | Pliocene | | {{Flag|Italy}} | A species of Alvania. | |
| Alvania pluricosticillata
| Sp. nov | Disputed | Chirli & Forli | Plio-Pleistocene | | {{Flag|Italy}} | Originally described as a species of Alvania. Argued to be a junior synonym of the extant taxon Crisilla gagliniae by Amati & Oliverio (2024).{{cite journal|last1=Amati |first1=B. |last2=Oliverio |first2=M. |year=2024 |title=Alvania pluricosticillata Chirli & Forli, 2021 junior synonym of Crisilla gagliniae (Amati, 1985) |journal=Bollettino Malacologico |volume=60 |issue=1 |pages=1–4 |doi=10.53559/BollMalacol.2023.17 }} | |
| Amaea (Confusiscala) paleocaenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Ammonicera danieli{{Cite journal|last1=Chernyshev |first1=A. V. |last2=Goedert |first2=J. L. |title=New species of latest Eocene/earliest Oligocene microgastropods (Heterobranchia: Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA |year=2021 |journal=Zootaxa |volume=4981 |issue=3 |pages=469–480 |doi=10.11646/zootaxa.4981.3.3 |pmid=34186710 |s2cid=235686756 }}
| Sp. nov | Valid | Chernyshev & Goedert | Late Eocene/early Oligocene | | {{Flag|United States}} | A species of Ammonicera. | |
| Ammonicera rolani
| Sp. nov | Valid | Chernyshev & Goedert | Late Eocene/early Oligocene | Gries Ranch Formation | {{Flag|United States}} | A species of Ammonicera. | |
| Ampezzogyra angulata{{Cite journal|last1=Hausmann |first1=I. M. |last2=Nützel |first2=A. |last3=Roden |first3=V. J. |last4=Reich |first4=M. |title=Palaeoecology of tropical marine invertebrate assemblages from the Late Triassic of Misurina, Dolomites, Italy |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=1 |pages=143–192 |doi=10.4202/app.00659.2019 |doi-access=free }}
| Sp. nov | Valid | Nützel & Hausmann in Hausmann et al. | Late Triassic (Carnian) | San Cassiano Formation | {{Flag|Italy}} | A member of the family Stuoraxidae. | |
| Amphidromus ubaldii{{Cite journal|last=Dharma |first=B. |title=New Species and Subspecies of Recent and Fossil Amphidromus Albers, 1850 from Java and Sumatra (Gastropoda: Camaenidae) |year=2021 |journal=The Festivus |volume=53 |issue=4 |pages=298–306 |doi=10.54173/F534298 |s2cid=240261318 |doi-access=free }}
| Sp. nov | Valid | Dharma | Late Pliocene | | {{Flag|Indonesia}} | A species of Amphidromus. | |
| Amplitomaria
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" spuria Münster (1841). | |
| Anacithara akisi{{Cite journal|last=Kolokotronis |first=D. |title=New records of tropical gastropods from the Pliocene of Nicosia with the description of Anacithara akisi sp. nov. (Gastropoda: Horaiclavidae) |year=2021 |journal=Gloria Maris |volume=60 |issue=4 |pages=178–183 |url=https://www.researchgate.net/publication/357528076 }}
| Sp. nov | Valid | Kolokotronis | Pliocene | | {{Flag|Cyprus}} | A species of Anacithara. | |
| Anceps siminescui
| Nom. nov | Valid | Harzhauser | Middle-late Miocene | | {{Flag|Moldova}} | A Trochidae member; | |
| Angulatella
| Gen. et sp. nov | Valid | Nützel & Hausmann in Hausmann et al. | Late Triassic (Carnian) | San Cassiano Formation | {{Flag|Italy}} | A member of the family Prostyliferidae. The type species is A. bizzarinii. | |
| Antithala{{Cite journal|last1=Harzhauser |first1=M. |last2=Landau |first2=B. |title=An overlooked diversity—the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea |year=2021 |journal=Zootaxa |volume=4982 |issue=1 |page=170 |doi=10.11646/zootaxa.4982.1.1 |issn=1175-5326 |pmid=34186703 |s2cid=235687490 }}
| Gen. et comb. et 2 sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Bosnia and Herzegovina}} {{Flag|Hungary}} {{Flag|Poland}} {{Flag|Romania}} {{Flag|Ukraine}} | A member of the family Costellariidae. The type species is "Mitra" sturi Hoernes & Auinger (1880); genus also includes "Mitra" neugeboreni Hoernes & Auinger (1880), as well as new species A. claviformis and A. filipescui. | |
| Apiocypraea cerretensis{{Cite journal|last1=Fehse |first1=D. |last2=Bertaccini |first2=E. |last3=Gardella |first3=F. |last4=Tabanelli |first4=C. |title=Revision of the Mediterranean Pliocene Apiocypraea F.A. Schilder, 1925 (Gastropoda: Eocypraeidae) with special consideration of shells of the "Spungone" area, Romagna, Northern Italy |year=2021 |journal=Archiv für Molluskenkunde: International Journal of Malacology |volume=150 |issue=2 |pages=147–171 |doi=10.1127/arch.moll/150/147-171 |s2cid=245406059 }}
| Sp. nov | Valid | Fehse et al. | | | {{Flag|Italy}} | A member of the family Eocypraeidae. | |
| Aplus abessensis{{Cite journal|last=Lozouet |first=P. |title=Buccinoidea (Mollusca, Gastropoda, Neogastropoda) de l'Oligocène supérieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France) |year=2021 |journal=Cossmanniana |volume=22 |pages=3–129 }}
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Aplus. | |
| Aplus grandiculus
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Aplus. | |
| Aplus littoralis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Aplus. | |
| Aplus pseudoaquitanensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Aplus. | |
| Aquilofusus? antiqua
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Ascheria canni{{Cite journal|last1=Kaim |first1=A. |last2=Little |first2=C. T. S. |last3=Kennedy |first3=W. J. |last4=Mears |first4=E. M. |last5=Anderson |first5=L. M. |title=Late Cretaceous hydrothermal vent communities from the Troodos ophiolite, Cyprus: systematics and evolutionary significance |year=2021 |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=1927–1947 |doi=10.1002/spp2.1370 |s2cid=237895585 |url=https://eprints.whiterose.ac.uk/175499/1/Kaim%20et%20al%202021%20Late%20Cretceous%20hydrothermal%20vent%20communities%20Troodos%20Ophioloite%20Cyprus%20Papers%20in%20Palaeontology.pdf }}
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A member of the family Hokkaidoconchidae. | |
| Asiaharpa{{Cite journal|last=Raven |first=J. G. M. |title=Notes on molluscs from nw Borneo 9. Harpidae (Neogastropoda), with the description of Asiaharpa sarawakiana gen. nov., spec. nov. |year=2021 |journal=Vita Malacologica |volume=20 |pages=51–60 }}
| Gen. et sp. nov | Valid | Raven | Miocene | | {{Flag|Malaysia}} | A member of the family Harpidae. Genus includes new species A. sarawakiana. | |
| Attenuata? danica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Possibly a species of Attenuata. | |
| Auingeria
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Czech Republic}} {{Flag|Germany}} {{Flag|Hungary}} | A member of the family Columbellidae. The type species is "Columbella" bellardii Hörnes (1852); genus also includes "Columbella (Anachis)" austriaca Hoernes & Auinger (1880) and "Pyrene (Anachis) bellardii" grussbachensis Csepreghy-Meznerics (1969; raised to the rank of a separate species Auingeria grussbachensis). | |
| Axonella{{Cite journal |last1=Pacaud |first1=J.-M. |last2=Lhomme |first2=J.-F. |last3=Renaud |first3=G. |title=Description d'un nouveau genre et d'une espèce nouvelle d'Hemisinidae (Mollusca, Gastropoda) du Cuisien (Yprésien inférieur) du bassin de Paris |year=2021 |journal=Folia Conchyliologica |volume=62 |pages=9–14 |url=https://www.researchgate.net/publication/355717416 }}
| Gen. et comb. et sp. nov | Valid | Pacaud, Lhomme & Renaud | | | {{Flag|France}} {{Flag|Spain}} | A member of the family Hemisinidae. The type species is "Cerithium" palense Rouault (1850); genus also includes "Potamides" granosus Doncieux (1908), as well as new species A. falsiacum. | |
| Babylonia chattica{{Cite journal|last=Lozouet |first=P. |title=Turbinelloidea, Mitroidea, Olivoidea, Babyloniidae et Harpidae (Gastropoda, Neogastropoda) de l'Oligocène supérieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France) |year=2021 |journal=Cossmanniana |volume=23 |pages=3–69 }}
| Sp. nov | Valid | Lozouet | | | {{Flag|France}} | A species of Babylonia. | |
| Babylonia edigittenbergeri{{Cite journal|last=Raven |first=J. G. M. |title=Notes on molluscs from NW Borneo 7. Babyloniidae (Gastropoda, Neogastropoda) |year=2021 |journal=Vita Malacologica |volume=20 |pages=14–22 }}
| Sp. nov | Valid | Raven | Miocene | Miri Formation | {{Flag|Malaysia}} | A species of Babylonia. | |
| Babylonia sarawakiana
| Sp. nov | Valid | Raven | Miocene | Miri Formation | {{Flag|Malaysia}} | A species of Babylonia. | |
| Bandellina compacta
| Sp. nov | Valid | Nützel & Hausmann in Hausmann et al. | Late Triassic (Carnian) | San Cassiano Formation | {{Flag|Italy}} | A member of the family Cornirostridae. | |
| Barycypraea abbasi{{Cite journal|last1=Dovesi |first1=M. |last2=Parsons |first2=J. |title=Two new species of Barycypraea (Gastropoda: Cypraeidae) from the Miocene of Java, Indonesia |year=2021 |journal=The Festivus |volume=53 |issue=2 |pages=116–127 |doi=10.54173/F532116 |s2cid=240518287 |doi-access=free }}
| Sp. nov | Valid | Dovesi & Parsons | Miocene (Serravallian) | | {{Flag|Indonesia}} | A species of Barycypraea. | |
| Barycypraea alessandrodovesi
| Sp. nov | Valid | Dovesi & Parsons | Miocene (Serravallian) | Nyalindung Formation | {{Flag|Indonesia}} | A species of Barycypraea. | |
| Bela pediensis{{Cite journal|last=Kolokotronis |first=D. |title=Bela pediensis sp. nov.: A new species of Bela (Gastropoda: Mangeliidae) from the Pliocene of Nicosia |year=2021 |journal=Gloria Maris |volume=60 |issue=3 |pages=128–130 |url=https://www.academia.edu/51810425 }}
| Sp. nov | Valid | Kolokotronis | Pliocene | | {{Flag|Cyprus}} | A species of Bela. | |
| Bellacolumbella
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Czech Republic}} {{Flag|Hungary}} {{Flag|Romania}} | A member of the family Columbellidae. The type species is "Columbella (Nitidella)" karreri Hoernes & Auinger (1880); genus also includes "Columbella (Nitidella)" embryonalis Boettger (1902) and "Columbella (Nitidella)" katharinae Hoernes & Auinger (1880). | |
| Bellardithala
| Nom. et 4 sp. nov | Valid | Harzhauser & Landau | Miocene and Pliocene | Baden Formation Dej Formation | {{Flag|Austria}} {{Flag|Czech Republic}} {{Flag|Italy}} {{Flag|Poland}} {{Flag|Romania}} {{Flag|Ukraine}} {{Flag|Hungary}}? | A member of the family Costellariidae; a replacement name for Micromitra Bellardi (1888). The type species is "Voluta" obsoleta Brocchi (1814); genus also includes new species B. kovaci, B. baluki, B. fedosovi and B. dacica. | |
| Bieleraxis
| Gen. et 2 sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species B. discoidea and B. altis. | |
| Bithynia disregularis{{Cite journal|last1=Li |first1=Z. |last2=Li |first2=Y. |last3=Zhang |first3=Y. |last4=Xie |first4=K. |last5=Li |first5=Z. |title=New Discovery of Oligocene Gastropod Opercula from the Yehucheng Formation in Northwestern China |year=2021 |journal=Acta Geologica Sinica (English Edition) |volume=96 |pages=61–70 |doi=10.1111/1755-6724.14727 |s2cid=236413377 }}
| Sp. nov | In press | Li et al. | Oligocene | | {{Flag|China}} | A species of Bithynia. | |
| Bithynia obliquus
| Sp. nov | In press | Li et al. | Oligocene | Yehucheng Formation | {{Flag|China}} | A species of Bithynia. | |
| Bithynia parainclinimarga
| Sp. nov | In press | Li et al. | Oligocene | Yehucheng Formation | {{Flag|China}} | A species of Bithynia. | |
| Bithynia paramonolithic
| Sp. nov | In press | Li et al. | Oligocene | Yehucheng Formation | {{Flag|China}} | A species of Bithynia. | |
| Bittium prekoeneni
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Bittium. | |
| Bulimulus frenguellii{{Cite journal|last1=Cabrera |first1=F. |last2=Breure |first2=A. S. H. |last3=Norbis |first3=W. |last4=Martínez |first4=S. |title=Description of a new Late Cretaceous species of Bulimulus Leach, 1814 (Orthalicoidea: Bulimulidae) using geometric morphometric analysis |year=2021 |journal=Journal of Molluscan Studies |volume=87 |issue=2 |pages=eyab006 |doi=10.1093/mollus/eyab006 }}
| Sp. nov | Valid | Cabrera et al. | | | {{Flag|Uruguay}} | A species of Bulimulus. | |
| Burminella{{Cite journal|last1=Balashov |first1=I. A. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |title=A mid-Cretaceous land snail Burminella artiukhini gen. et sp. nov. from Burmese amber: a "missing link" between Pupinidae and other Cyclophoroidea? (Caenogastropoda) |journal=Cretaceous Research |year=2021 |volume=127 |pages=Article 104941 |doi=10.1016/j.cretres.2021.104941 |bibcode=2021CrRes.12704941B | issn=0195-6671 |doi-access=free }}
| Gen. et sp. nov | In press | Balashov, Perkovsky & Vasilenko | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | A land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea. Genus includes new species B. artiukhini. | |
| Bursa transeuntis{{Cite journal|last1=Forli |first1=M. |last2=Smriglio |first2=C. |title=A new species of Tonnoidea Suter, 1813 (Gastropoda Bursidae) for the Italian Pliocene: Bursa transeuntis n. sp. |year=2021 |journal=Biodiversity Journal |volume=12 |issue=2 |pages=483–492 |doi=10.31396/Biodiv.Jour.2021.12.2.483.492 |s2cid=237810888 |doi-access=free }}
| Sp. nov | Valid | Forli & Smriglio | Miocene and Pliocene | | {{Flag|Hungary}} {{Flag|Italy}} {{Flag|Poland}} | A species of Bursa. | |
| Caecum texanum
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Caecum. | |
| Califrapana{{Cite journal|last1=Powell |first1=C. L. |last2=Houart |first2=R. |title=Califrapana: a new genus of California and Bája California late Oligocene to early Miocene muricids previously attributed to the genus Rapana (Mollusca: Gastropoda: Muricidae) |year=2021 |journal=PaleoBios |volume=38 |pages=ucmp_paleobios_52825 |doi=10.5070/P9381052825 |s2cid=236578864 |url=https://escholarship.org/uc/item/3wr5675z |doi-access=free }}
| Gen. et comb. nov | Valid | Powell & Houart | Late Oligocene to early Miocene | | {{Flag|Mexico}} {{Flag|United States}} | A member of the family Muricidae. The type species is "Purpura" vaquerosensis Arnold (1907). | |
| Calyptraphorus velusus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Camponaxis bandeli{{Cite journal|last1=Pieroni |first1=V. |last2=Monari |first2=S. |last3=Todd |first3=J. A. |title=A new caenogastropod from the upper Rhaetian of Lombardy: Palaeobiogeographical history and implications for the Early Jurassic gastropod recovery |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=1 |pages=193–206 |doi=10.4202/app.00792.2020 |doi-access=free }}
| Sp. nov | Valid | Pieroni, Monari & Todd | Late Triassic (Carnian) | | {{Flag|Italy}} | A Tofanellidae member | |
| Campylacrum texanum
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Canariella (Majorata) gerti{{Cite journal|last=Groh |first=K. |title=Description of a new Canariella P. Hesse, 1918 (Stylommatophora: Canariellidae) from the Quaternary of Furteventura, Canary Islands |year=2021 |journal=Basteria. Journal of the Netherlands Malacological Society |volume=85 |issue=2 |pages=126–129 }}
| Sp. nov | Valid | Groh | Quaternary | | {{Flag|Canary Islands}} | A species of Canariella. | |
| Cancellotomaria
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" subcancellata d'Orbigny (1850). | |
| Cancilla nanostriatula{{Cite journal|last1=Harzhauser |first1=M. |last2=Landau |first2=B. |title=The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea |year=2021 |journal=Zootaxa |volume=4983 |issue=1 |page=172 |doi=10.11646/zootaxa.4983.1.1 |pmid=34186702 |s2cid=235687636 }}
| Sp. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Austria}} {{Flag|Bulgaria}} {{Flag|Hungary}} | A species of Cancilla. | |
| Cancilla wagreichi
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} | A species of Cancilla. | |
| Capulus gelus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Capulus. | |
| Cassis kalmani{{Cite journal|last1=Vicián |first1=Z. |last2=Kovács |first2=Z. |title=Middle Eocene Tonnoidea (Caenogastropoda) from the Hungarian Paleogene Basin |year=2021 |journal=Fragmenta Palaeontologica Hungarica |volume=37 |pages=13–47 |doi=10.17111/FragmPalHung.2021.37.13 |s2cid=252349779 |url=https://www.researchgate.net/publication/361606471 }}
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A species of Cassis. | |
| Cassis rasae{{Cite journal|last1=Petuch|first1=E. J. |last2=Berschauer |first2=D. P. |title=Two new Cassis (Gastropoda: Cassidae) and a new Malea (Gastropoda: Tonnidae) from the Pliocene and Pleistocene Beds of southern Florida |year=2021 |journal=The Festivus |volume=53 |issue=1 |pages=3–10 |doi=10.54173/F531003 |s2cid=240504008 }}
| Sp. nov | Valid | Petuch & Berschauer | | | {{Flag|United States}} | A species of Cassis. | |
| Cassis viliusi
| Sp. nov | Valid | Petuch & Berschauer | Pleistocene | | {{Flag|United States}} | A species of Cassis. | |
| Cenorhytis
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species C. groesbecki. | |
| Ceratopea? moridunensis{{Cite journal|last1=Ebbestad |first1=J. O. R. |last2=Cope |first2=J. C. W. |title=A low diversity Sinuites gastropod community from the Floian, Early Ordovician, of South Wales |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=2 |pages=319–335 |doi=10.4202/app.00854.2020 |s2cid=236651641 |doi-access=free }}
| Sp. nov | Valid | Ebbestad & Cope | | | {{Flag|United Kingdom}} | A member of the family Raphistomatidae. | |
| Cerithiella acuta
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Cerithiella. | |
| Cerithiscala parvibrazosa
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Cerithiscala tehuacana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Chemnitzia silvai
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Chemnitzia. | |
| Chevallieria texanopsis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Chicomurex parvus{{Cite journal|last1=Merle |first1=D. |last2=Landau |first2=B. M. |last3=Breitenberger |first3=A. E. |title=New Muricidae (Mollusca, Gastropoda) from the Miocene of Java (Indonesia) |year=2021 |journal=Basteria. Journal of the Netherlands Malacological Society |volume=85 |issue=1 |pages=21–33 |url=https://www.researchgate.net/publication/351159966 }}
| Sp. nov | Valid | Merle, Landau & Breitenberger | | | {{Flag|Indonesia}} | A species of Chicomurex. | |
| Chicoreus javanus
| Sp. nov | Valid | Merle, Landau & Breitenberger | Miocene | | {{Flag|Indonesia}} | A species of Chicoreus. | |
| Chicoreus martini
| Sp. nov | Valid | Merle, Landau & Breitenberger | Miocene | | {{Flag|Indonesia}} | A species of Chicoreus. | |
| Chicoreus solidus
| Sp. nov | Valid | Merle, Landau & Breitenberger | Miocene | | {{Flag|Indonesia}} | A species of Chicoreus. | |
| Clava (Semivertagus) scriba
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Clavatula hirmetzli{{Cite journal|last1=Kovács |first1=Z. |last2=Vicián |first2=Z. |title=Conoidea (Neogastropoda) assemblage from the Lower Badenian (Middle Miocene) deposits of Letkés (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae) |year=2021 |journal=Földtani Közlöny |volume=151 |issue=2 |pages=137–158 |doi=10.23928/foldt.kozl.2021.151.2.137 |s2cid=237929980 |doi-access=free }}
| Sp. nov | Valid | Kovács & Vicián | Middle Miocene | Lajta Limestone Formation | {{Flag|Hungary}} | A species of Clavatula. | |
| Clavatula santhai
| Sp. nov | Valid | Kovács & Vicián | Middle Miocene | Lajta Limestone Formation | {{Flag|Hungary}} | A species of Clavatula. | |
| Clavatula szekelyhidiae
| Sp. nov | Valid | Kovács & Vicián | Middle Miocene | Lajta Limestone Formation | {{Flag|Hungary}} | A species of Clavatula. | |
| Cochlespira (Tahusyrinx) laevis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Cochlespira. | |
| Conilithes herodus{{Cite journal|last1=Psarras |first1=C. |last2=Koskeridou |first2=E. |last3=Merle |first3=D. |title=Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 1: genera Conilithes Swainson, 1840 and Conus (Kalloconus) da Motta, 1991 |year=2021 |journal=Geodiversitas |volume=43 |issue=24 |pages=1309–1339 |doi=10.5252/geodiversitas2021v43a24 |s2cid=244932519 |url=https://sciencepress.mnhn.fr/en/periodiques/geodiversitas/43/24 |doi-access=free }}
| Sp. nov | Valid | Psarras, Koskeridou & Merle | Miocene (Tortonian) | | {{Flag|Greece}} | A species of Conilithes. | |
| Conradconfusus nodulinus{{Cite journal|last=Garvie |first=C. L. |title=Eight new species of Eocene Gastropoda from Texas based on the Megahan collection |year=2021 |journal=Proceedings of the Biological Society of Washington |volume=134 |issue=1 |pages=196–208 |doi=10.2988/0006-324X-134.1.196 |s2cid=240502013 }}
| Sp. nov | Valid | Garvie | | | {{Flag|United States}} | | |
| Conradconfusus texanus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Conus (Kalloconus) asterousiaensis
| Sp. nov | Valid | Psarras, Koskeridou & Merle | Miocene (Tortonian) | | {{Flag|Greece}} {{Flag|Italy}} | A species of Conus. | |
| Conus (Kalloconus) helladicus
| Sp. nov | Valid | Psarras, Koskeridou & Merle | Miocene (Tortonian) | | {{Flag|Greece}} | A species of Conus. | |
| Cordieria biouesensis{{Cite journal |last=Pacaud |first=J.-M. |title=Remarques taxonomiques et nomenclaturales sur les mollusques gastéropodes du Paléogène de France et description d'espèces nouvelles. Partie 3. Conoidea |year=2021 |journal=Xenophora Taxonomy |volume=31 |pages=41–48 |url=https://www.academia.edu/44938873 |access-date=2021-01-23 |archive-date=2021-02-21 |archive-url=https://web.archive.org/web/20210221083216/https://www.academia.edu/44938873/Remarques_taxonomiques_et_nomenclaturales_sur_les_mollusques_gast%C3%A9ropodes_du_Pal%C3%A9og%C3%A8ne_de_France_et_description_d_esp%C3%A8ces_nouvelles_Partie_3_Conoidea |url-status=live }}
| Sp. nov | Valid | Pacaud | | | {{Flag|France}} | A species of Cordieria. | |
| Coronia argilla
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Costoanachis venzoi{{Cite journal|last1=Harzhauser |first1=M. |last2=Landau |first2=B. M. |title=The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group |year=2021 |journal=Zootaxa |volume=5025 |issue=1 |pages=1–75 |doi=10.11646/zootaxa.5025.1.1 |pmid=34810943 |s2cid=238735080 }}
| Nom. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Italy}} | A species of Costoanachis; a replacement name for Anachis (Costoanachis) subcorrugata Venzo & Pelosio (1963). | |
| Crassaseila
| Gen. et 2 sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species C. ripleyana and C. paleocenica. | |
| Crassispira plioibericostricta{{Cite journal|last1=Brunetti |first1=M. M. |last2=Forli |first2=M. |title=Some notes on the Genus Crassispira Swainson, 1840 (Gastropoda: Pseudomelatomidae) with the description of two new species from the Italian and Spanish Pliocene |year=2021 |journal=Bollettino Malacologico |volume=57 |issue=2 |pages=182–191 |url=https://www.researchgate.net/publication/357910548 }}
| Sp. nov | Valid | Brunetti & Forli | Pliocene (Zanclean) | | {{Flag|Spain}} | A species of Crassispira. | |
| Crassispira pseudoglabra
| Sp. nov | Valid | Brunetti & Forli | Pliocene (Piacenzian) | | {{Flag|Italy}} | A species of Crassispira. | |
| Crenaturricula minuta
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Creonella hebetertia
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Cretatortulosa cretakachinensis
| Sp. nov | In press | Yu, Salvador & Jarzembowski | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | | |
| Cretatortulosa gignens{{Cite journal|last1=Jochum |first1=A. |last2=Yu |first2=T. |last3=Neubauer |first3=T. A. |title=Mother snail labors for posterity in bed of mid-Cretaceous amber |year=2021 |journal=Gondwana Research |volume=97 |pages=68–72 |doi=10.1016/j.gr.2021.05.006 |bibcode=2021GondR..97...68J |doi-access=free }}{{Cite journal|last1=Jiang |first1=R. |last2=Shi |first2=C. |last3=Zhang |first3=H. |last4=Cai |first4=H. |last5=Long |first5=X. |last6=Wang |first6=S. |last7=Engel |first7=M. S. |title=Comment on "Mother snail labors for posterity in bed of mid-Cretaceous amber" by A. Jochum, T. Yu and T.A. Neubauer, Gondwana Research, Volume 97, Pages 68–72 |journal=Gondwana Research |year=2022 |volume=101 |pages=21–23 |doi=10.1016/j.gr.2021.07.015 |bibcode=2022GondR.101...21J | issn=1342-937X }}{{Cite journal|last1=Jochum |first1=A. |last2=Yu |first2=T. |last3=Neubauer |first3=T. A. |title=Reply to Jiang et al.'s comments on the article "Mother snail labors for posterity in bed of mid-Cretaceous amber" by Jochum, Yu and Neubauer (2021) Gondwana Research 97, 68–72 |year=2021 |journal=Gondwana Research |volume=101 |pages=24–25 |doi=10.1016/j.gr.2021.07.016|issn=1342-937X |doi-access=free }}
| Sp. nov | In press | Jochum, Yu & Neubauer | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | A member of the family Pupinidae. | |
| Cryptochorda paleocaenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Cyathopoma pingyiensis{{Cite journal|last1=Yu |first1=T. |last2=Wang |first2=H. |last3=Fang |first3=Y. |last4=Salvador |first4=R. B. |last5=Li |first5=S. |last6=Jarzembowski |first6=E. A. |last7=Yu |first7=H. |last8=Zhang |first8=H. |title=Non-marine gastropods from the Cretaceous–Paleogene transition in the Pingyi Basin, eastern China |year=2021 |journal=Proceedings of the Geologists' Association |volume=133 |pages=40–46 |doi=10.1016/j.pgeola.2021.10.003 |s2cid=244516164 }}
| Sp. nov | In press | Yu et al. | Cretaceous–Paleogene transition | Pingyi Basin | {{Flag|China}} | A species of Cyathopoma. | |
| Cylichna (Cylichnopsis) elliptica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Cylichna. | |
| Cymatiella dulaii
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A species of Cymatiella. | |
| Cymatiella tzankovi
| Nom. nov | Valid | Vicián & Kovács | Paleocene | | {{Flag|Bulgaria}} | A species of Cymatiella; a replacement name for Eutritonium (Sassia) rutoti Tzankov (1940). | |
| Cymatium danianum
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Cymatium. | |
| Cyphoma guraboensis{{Cite journal |last1=Groves |first1=L. T. |last2=Landau |first2=B. M. |title=Neogene paleontology in the northern Dominican Republic. 25. The Superfamily Cypraeoidea (Families Cypraeidae, Ovulidae, Triviidae, and Eratoidae) (Mollusca: Gastropoda) |year=2021 |journal=Bulletins of American Paleontology |volume=401 |issue=401 |pages=1–110 |doi=10.32857/bap.2021.401.01 |doi-access=free }}
| Sp. nov | Valid | Groves & Landau | Early Pliocene | | {{Flag|Dominican Republic}} | A species of Cyphoma. | |
| Cypraeorbis alisonkayae
| Sp. nov | Valid | Groves & Landau | Miocene | | {{Flag|Dominican Republic}} | A species of Cypraeorbis. | |
| Cyprioconcha
| Gen. et sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A member of the family Hokkaidoconchidae. The type species is C. robertsoni. | |
| Defensina
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Bosnia and Herzegovina}} {{Flag|Bulgaria}} {{Flag|Romania}} | A member of the family Columbellidae. The type species is "Columbella (Mitrella)" bucciniformis Hoernes & Auinger (1880). | |
| Dermomurex wanneri
| Nom. nov | Valid | Merle, Landau & Breitenberger | Miocene | | {{Flag|Indonesia}} | A species of Dermomurex; a replacement name for Murex acuticostatus Wanner & Hahn (1935). | |
| Desbruyeresia kambiaensis
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A species of Desbruyeresia. | |
| Desbruyeresia kinousaensis
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A species of Desbruyeresia. | |
| Desbruyeresia memiensis
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A species of Desbruyeresia. | |
| Discotectonica paracutissima
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Discotectonica. | |
| Doliella parnitens
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Doliella. | |
| Domiporta amoena
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Austria}} {{Flag|Bulgaria}} {{Flag|Romania}} | A species of Domiporta. | |
| Domiporta pulchra
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} | A species of Domiporta. | |
| Domiporta turpis
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} | A species of Domiporta. | |
| Ederazyga
| Gen. et sp. et comb. nov | Valid | Pieroni, Monari & Todd | Late Triassic (Carnian to Rhaetian) | | {{Flag|Iran}} {{Flag|Italy}} {{Flag|Slovenia}} | A possible Zygopleuridae member. | |
| Ellobium mizutanii{{Cite journal|last1=Kawase |first1=M. |last2=Ichihara |first2=T. |title=A new species of Ellobium (Gastropoda: Panpulmonata: Ellobiidae) from the Miocene Mizunami Group in Gifu Prefecture, central Japan |year=2021 |journal=Bulletin of the Mizunami Fossil Museum |volume=48 |pages=1–7 |doi=10.50897/bmfm.48.0_1 }}
| Sp. nov | Valid | Kawase & Ichihara | Early Miocene | | {{Flag|Japan}} | A species of Ellobium. | |
| Entemnotrochus kathiawarensis{{Cite journal|last1=Bose |first1=K. |last2=Das |first2=S. S. |last3=Mondal |first3=S. |title=An updated generic classification of Cenozoic pleurotomariid gastropods, with new records from the Oligocene and early Miocene of India |year=2021 |journal=Journal of Paleontology |volume=95 |issue=4 |pages=763–776 |doi=10.1017/jpa.2021.4 |bibcode=2021JPal...95..763B |issn=0022-3360 |s2cid=233801673 |url=https://zenodo.org/record/4391425 }}
| Sp. nov | Valid | Bose, Das & Mondal | Miocene (Burdigalian) | | {{Flag|India}} | A species of Entemnotrochus. | |
| Eoacteon paracutissima
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Eoancilla lapicidina
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Eocithara (?) laurencei
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Eopleurotoma atacica
| Nom. nov | Valid | Pacaud | | | {{Flag|France}} | A species of Eopleurotoma; a replacement name for Pleurotoma (Eopleurotoma) romani Doncieux (1908). | |
| Eotrivia pacaudi{{Cite journal|last=Fehse |first=D. |title=Zur Identität von Eotrivia faracii (de Gregorio, 1880) und Beschreibung einer neuen Art in der Gattung Eotrivia F. A. Schilder, 1924 (Mollusca, Gastropoda) |year=2021 |journal=Conchylia |volume=52 |issue=1–2 |pages=79–84 }}
| Sp. nov | Valid | Fehse | Eocene | | {{Flag|Italy}} | | |
| Eoturris fallsensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Eoworthenia frydai
| Nom. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida; a replacement name for Worthenia rarissima Barrande. | |
| Episcomitra antibellardii
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} | A species of Episcomitra. | |
| Episcomitra leopoldiana
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Czech Republic}} {{Flag|Poland}} | A species of Episcomitra. | |
| Episcomitra missile
| Sp. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Austria}} {{Flag|Hungary}} | A species of Episcomitra. | |
| Episcomitra neubaueri
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} | A species of Episcomitra. | |
| Episcomitra pontonxensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Episcomitra. | |
| Episcomitra pseudoincognita
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} {{Flag|Ukraine}} | A species of Episcomitra. | |
| Episcomitra vermeuleni{{Cite journal|last=Pacaud |first=J.-M. |title=Description d'une espèce nouvelle de Mitridae (Mollusca, Gastropoda) du Priabonien (Éocène superieur) de Dnipro (Oblast de Dnipropetrovsk, Ukraine) |year=2021 |journal=Belokrysia |volume=3 |pages=1–11 |url=https://www.researchgate.net/publication/355978782 }}
| Sp. nov | Valid | Pacaud | Eocene (Priabonian) | | {{Flag|Ukraine}} | A species of Episcomitra. | |
| Eulimella ariejansseni
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Eulimella. | |
| Euthema myanmarica{{Cite journal|last=Balashov |first=I. |title=The first records of mollusks from mid-Cretaceous Hkamti amber (Myanmar), with the description of a land snail, Euthema myanmarica n. sp. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) |year=2021 |journal=Journal of Paleontology |volume=95 |issue=5 |pages=994–1003 |doi=10.1017/jpa.2021.26 |bibcode=2021JPal...95..994B |s2cid=233690461 }}
| Sp. nov | Valid | Balashov | Cretaceous | Hkamti amber | {{Flag|Myanmar}} | A species of Euthema. | |
| Euthria mellianensis
| Sp. nov | Valid | Lozouet | Miocene | | {{Flag|France}} | A species of Euthria. | |
| Euthria tarusatensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Euthria. | |
| Euthriofusus argillensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | | |
| Falsifusus problematicus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Fedosovia
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Czech Republic}} {{Flag|Italy}} {{Flag|Romania}} | A member of the family Costellariidae. The type species is "Mitra (Callithea)" fuchsi Hoernes & Auinger (1880). | |
| Fraudiziba
| Gen. et nom. et sp. nov | Valid | Harzhauser & Landau | Miocene and Pliocene | Karaman Basin | {{Flag|Austria}} {{Flag|Bosnia and Herzegovina}} {{Flag|Czech Republic}} {{Flag|Hungary}} {{Flag|Italy}} {{Flag|Poland}} {{Flag|Romania}} {{Flag|Slovakia}} {{Flag|Turkey}} {{Flag|Ukraine}} | A member of the family Mitridae. The type species is F. paratethyca (a replacement name for Mitra austriaca Csepreghy-Meznerics, 1950); genus also includes new species F. ottomanica. | |
| Fulgurofusus spinosus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Fulgurofusus. | |
| Fusulculus texanus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Fusulculus. | |
| Galba prima{{Cite journal|last1=Yu |first1=T. |last2=Neubauer |first2=T. A. |last3=Jochum |first3=A. |title=First freshwater gastropod preserved in amber suggests long-distance dispersal during the Cretaceous Period |year=2021 |journal=Geological Magazine |volume=158 |issue=7 |pages=1327–1334 |doi=10.1017/S0016756821000285 |bibcode=2021GeoM..158.1327Y |s2cid=235529278 }}
| Sp. nov | Valid | Yu, Neubauer & Jochum | Cretaceous | Burmese amber | {{Flag|Myanmar}} | A species of Galba. | |
| Gemmula bearrizensis
| Nom. nov | Valid | Pacaud | Eocene (Priabonian) | | {{Flag|France}} | A species of Gemmula; a replacement name for Drillia pellati Boussac (1911). | |
| Gemmula osca
| Sp. nov | Valid | Pacaud | Eocene (Bartonian) | | {{Flag|Spain}} | A species of Gemmula. | |
| Gentilifulgur
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species G. praemortoni. | |
| Gibbula lovellreevei
| Nom. nov | Valid | Harzhauser | Pliocene | | {{Flag|United Kingdom}} | A Trochidae member; | |
| Gibbula pizzolatoi{{Cite journal|last=Spadini |first=V. |title=The genus Gibbula Risso, 1826 (Trochidae: Cantharidinae) from the Pliocene of Siena |year=2021 |journal=Bollettino Malacologico |volume=57 |issue=1 |pages=82–91 }}
| Sp. nov | Valid | Spadini | Pliocene | | {{Flag|Italy}} | A species of Gibbula. | |
| Gibbula steiningeri
| Nom. nov | Valid | Harzhauser | Early Miocene | | {{Flag|Austria}} | A Trochidae member; | |
| Gibbula tavanii
| Nom. nov | Valid | Harzhauser | Miocene | | {{Flag|Libya}} | A Trochidae member; | |
| Gibbuliculus{{Cite journal|last=Harzhauser |first=M. |title=The Cainozoic to present-day record of Circum-Mediterranean, NE Atlantic and North Sea Cantharidinae and Trochinae (Trochoidea, Gastropoda)—a synopsis |year=2021 |journal=Zootaxa |volume=4902 |issue=1 |pages=zootaxa.4902.1.1 |doi=10.11646/zootaxa.4902.1.1 |pmid=33757113 |s2cid=232339137 }}
| Gen. et comb. et nom. nov | Valid | Harzhauser | Oligocene to Pleistocene | | {{Flag|Europe}} | A Trochidae member; | |
| "Gibbulinella" sandbergeri
| Nom. nov | Valid | Harzhauser & Neubauer | Eocene | | {{Flag|Italy}} | A member of the family Streptaxidae; a replacement name for Pupa simplex Sandberger (1870). | |
| Haedropleura dellabellai
| Sp. nov | Valid | Brunetti & Cresti | Pliocene | | {{Flag|Italy}} | A species of Haedropleura. | |
| Hammatospira cancellata{{Cite journal |last=Mazaev |first=A. V. |title=Lower Carboniferous Gastropods of Dombar Hills (Viséan–Serpukhovian Boundary Beds, Western Kazakhstan) |year=2021 |journal=Paleontological Journal |volume=55 |issue=8 |pages=876–905 |doi=10.1134/S0031030121080049 |s2cid=245330703 }}
| Sp. nov | Valid | Mazaev | Early Carboniferous | | {{Flag|Kazakhstan}} | | |
| Hespererato canae
| Sp. nov | Valid | Groves & Landau | Late Miocene and early Pliocene | | {{Flag|Dominican Republic}} | A species of Hespererato. | |
| Hespererato praeclusa
| Sp. nov | Valid | Groves & Landau | Early–middle Miocene | | {{Flag|Dominican Republic}} | A species of Hespererato. | |
| Hexaplex nascium
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Hexaplex. | |
| Hirsuticyclus canaliculatus{{cite book |last=Yu |first=T. |year=2021 |chapter=New data on gastropod genus Hirsuticyclus (Caenogastropoda, Cyclophoridae) from mid-Cretaceous Kachin amber |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |journal=Geological Society, London, Special Publications |volume=521 |pages=153–160 |publisher=The Geological Society of London |doi=10.1144/SP521-2021-127 |s2cid=244091697 }}
| Sp. nov | In press | Yu | Cretaceous | Burmese amber | {{Flag|Myanmar}} | A member of the family Cyclophoridae. | |
| Hochheimia{{Cite journal |last1=Harzhauser |first1=M. |last2=Neubauer |first2=T. A. |title=A review of the land snail faunas of the European Cenozoic – composition, diversity and turnovers |year=2021 |journal=Earth-Science Reviews |volume=217 |pages=Article 103610 |doi=10.1016/j.earscirev.2021.103610 |bibcode=2021ESRv..21703610H |s2cid=233835790 }}
| Nom. nov | Valid | Harzhauser & Neubauer | Oligocene | | {{Flag|Germany}} | A member of the family Hygromiidae; a replacement name for Palaeotrichia Nordsieck (2014). | |
| Hokkaidoconcha morisseaui
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A member of the family Hokkaidoconchidae. | |
| Holospira? fallsensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Possibly a species of Holospira. | |
| ?Hydrocena praecursor{{Cite journal|last1=Yu |first1=T. |last2=Neubauer |first2=T. A. |title=The oldest fossil Hydrocenidae found in mid-Cretaceous Burmese amber (Gastropoda: Cycloneritida) |year=2021 |journal=Cretaceous Research |volume=122 |pages=Article 104765 |doi=10.1016/j.cretres.2021.104765 |bibcode=2021CrRes.12204765T |s2cid=233585283 }}
| Sp. nov | In press | Yu & Neubauer | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | A member of the family Hydrocenidae. | |
| Jponsia lacetania{{Cite journal |last1=Pacaud |first1=J.-M. |last2=Goret |first2=B. |title=Description d'une espèce nouvelle de Jponsia Pacaud & Harzhauser, 2012 (Mollusca, Gastropoda, Pachychilidae) de l'Ilerdien (Yprésien inférieur) du bassin de Tremp (Espagne) |year=2021 |journal=Folia Conchyliologica |volume=60 |pages=9–18 |url=https://www.researchgate.net/publication/349367822 |access-date=2021-02-20 |archive-date=2021-02-21 |archive-url=https://web.archive.org/web/20210221083217/https://www.researchgate.net/publication/349367822_Description_d%27une_espece_nouvelle_de_Jponsia_Pacaud_Harzhauser_2012_Mollusca_Gastropoda_Pachychilidae_de_l%27Ilerdien_Ypresien_inferieur_du_bassin_de_Tremp_Espagne_Folia_Conchyliologica_60_9-18_text-fig |url-status=live }}
| Sp. nov | Valid | Pacaud & Goret | | | {{Flag|Spain}} | A member of the family Pachychilidae. | |
| Kapalmerella gelus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Keepingia elongata
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | | |
| Laevispira
| Gen. et comb. et sp. nov | Valid | Raven | Miocene and Pliocene | | {{Flag|Brunei}} {{Flag|India}} {{Flag|Indonesia}} | A member of the family Strombidae. Genus includes "Dolomena" bruneiensis Harzhauser, Raven & Landau (2018), "Strombus" glaber Martin (1879), "Strombus" karikalensis Cossmann (1903), "Strombus" triangulatus Martin (1879) and "Strombus" varinginensis Martin (1899), as well as new species L. seriaensis. | |
| Laevitrivia{{Cite journal|last1=Fehse |first1=D. |last2=Vicián |first2=Z. |title=Contributions to the knowledge of the Triviidae. XLII: The Genus Laevitrivia n. gen |year=2021 |journal=Conchylia |volume=52 |issue=1–2 |pages=55–78 }}
| Gen. et comb. et 5 sp. nov | Valid | Fehse & Vicián | Miocene and Pliocene | | {{Flag|France}} {{Flag|Italy}} | A member of the family Triviidae. The type species is "Cypraea" pisolina Lamarck, (1810); genus also includes new species L. annae, L. bertaccinii, L. pliopisolina, L. pseudocalva and L. ventricostata. | |
| Latirus? asperpellis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Possibly a species of Latirus. | |
| Laubella subsulcata
| Sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. | |
| Lineacingulum
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" texturata Münster (1841). | |
| Lineaetomaria
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" decorata Münster (1841). | |
| Liostomia wilvanderstoelae{{Cite journal|last1=Landau |first1=B. M. |last2=Micali |first2=P. |title=The Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. Part 13: Murchisonelloidea and Pyramidelloidea |year=2021 |journal=Cainozoic Research |volume=21 |issue=2 |pages=201–393 |url=https://www.researchgate.net/publication/357117740 }}
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Liostomia. | |
| Littorina elongata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Littorina. | |
| Littorina (Prosthenodon) incertus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Littorina. | |
| Lupira danianum
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Lutema gracilis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Lyria elongata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Lyria. | |
| Lyria tehuacana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Lyria. | |
| Malea hyaducki
| Sp. nov | Valid | Petuch & Berschauer | | | {{Flag|United States}} | A species of Malea. | |
| Martaia
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Austria}} {{Flag|Bulgaria}} {{Flag|Czech Republic}} {{Flag|Hungary}} {{Flag|Romania}} {{Flag|Ukraine}} | A member of the family Columbellidae. The type species is "Columbella (Anachis)" dujardini Hoernes & Auinger (1880); genus also includes "Columbella (Anachis)" zitteli Hoernes & Auinger (1880). | |
| Mathilda (Echinimathilda) duogenta
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Mathilda. | |
| Melongena pontonxensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Melongena. | |
| Metula bulbiformis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Metula. | |
| Metula nana
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Metula. | |
| Metula pseudomajor
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Metula. | |
| Metula rivulisensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Metula. | |
| Mimospira llangynogensis
| Sp. nov | Valid | Ebbestad & Cope | Ordovician (Floian) | | {{Flag|United Kingdom}} | A member of the family Clisospiridae. | |
| Mitrella dacica
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Hungary}} {{Flag|Romania}} {{Flag|Slovakia}} | A species of Mitrella. | |
| Mitrella demaintenonae
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation Dej Formation | {{Flag|Austria}} {{Flag|Romania}} | A species of Mitrella. | |
| Mitrella elongatissima
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Bosnia and Herzegovina}} | A species of Mitrella. | |
| Mitrella fortis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Mitrella. | |
| Mitrella tarbelliana
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Mitrella. | |
| Mitrella viennensis
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Bulgaria}} {{Flag|Czech Republic}} {{Flag|Kazakhstan}} {{Flag|Poland}} {{Flag|Romania}} | A species of Mitrella. | |
| Moniquia astibiai
| Sp. nov | Valid | Pacaud & Goret | Eocene (Priabonian) | | {{Flag|Spain}} | A member of the family Pachychilidae. | |
| Monoplex s.l. szakonyii
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A member of the family Cymatiidae. | |
| Mulderia mulderi
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A member of the family Pyramidellidae. | |
| Multitrochia
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species M. tehuacana. | |
| Nannamoria alquezae{{Cite journal|last=Hawke |first=A. L. |title=New Species of Nannamoria from the Pliocene and Miocene of Australia (Caenogastropoda: Volutidae) |year=2021 |journal=The Festivus |volume=53 |issue=4 |pages=245–260 |doi=10.54173/F534245 |s2cid=240263447 |doi-access=free }}
| Sp. nov | Valid | Hawke | Miocene | | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria cadella
| Sp. nov | Valid | Hawke | Miocene | Cadel Formation | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria costatum
| Sp. nov | Valid | Hawke | Pliocene | Cameron Inlet Formation | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria flindersi
| Sp. nov | Valid | Hawke | Pliocene | Cameron Inlet Formation | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria gnotuki
| Sp. nov | Valid | Hawke | | | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria hiscocki
| Sp. nov | Valid | Hawke | Pliocene | Cameron Inlet Formation | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria malonei
| Sp. nov | Valid | Hawke | | | {{Flag|Australia}} | A species of Nannamoria. | |
| Nannamoria persimilis
| Sp. nov | Valid | Hawke | Miocene | | {{Flag|Australia}} | A species of Nannamoria. | |
| Naria praehelvola{{Cite journal|last1=Fehse |first1=D. |last2=Vicián |first2=Z. |title=New Miocene Cypraeids (Mollusca: Gastropoda: Cypraeidae) from Indonesia |year=2021 |journal=Conchylia |volume=51 |issue=3–4 |pages=35–53 }}
| Sp. nov | Valid | Fehse & Vicián | Miocene | | {{Flag|Indonesia}} | A species of Naria. | |
| Natica (Carinacca) kincaidensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Natica. | |
| Nebularia soliphila
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} | A species of Nebularia. | |
| Nemaspira{{Cite journal |last=Mazaev |first=A. V. |title=Revision of the gastropod family Gosseletinidae Wenz: genera Gosseletina Fischer, 1885, Globodoma Mazaev, 2006, and Nemaspira gen. nov. |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=600–607 |doi=10.1134/S0031030121060071 |bibcode=2021PalJ...55..600M |s2cid=245010385 |url=https://www.elibrary.ru/item.asp?id=46621790 |access-date=2021-10-15 |archive-date=2021-10-28 |archive-url=https://web.archive.org/web/20211028171205/https://www.elibrary.ru/item.asp%3Fid%3D46621790 |url-status=dead |url-access=subscription }}
| Gen. et comb. et 2 sp. nov | Valid | Mazaev | Early Carboniferous to Late Triassic | | {{Flag|Belgium}} {{Flag|Russia}} {{Flag|Thailand}} {{Flag|United Kingdom}} {{Flag|United States}} {{Flag|China}}? | A member of the family Gosseletinidae. Genus includes several species formerly assigned to the genera Gosseletina and Globodoma, as well as new species N. shamaevi and N. skuini. | |
| Nitidiclavus senesi{{Cite journal|last=Biskupič |first=K. |title=A new species of Nitidiclavus (Neogastropoda: Drilliidae) from the Miocene Paratethys and an overview of the paleoecology and distribution of related species in the Cainozoic of Europe |year=2021 |journal=Basteria. Journal of the Netherlands Malacological Society |volume=85 |issue=2 |pages=163–176 }}
| Sp. nov | Valid | Biskupič | Miocene | | {{Flag|Slovakia}} | A member of the family Drilliidae. | |
| Niveria callosa{{Cite journal|last=Fehse |first=D. |title=Contributions to the knowledge of the Triviidae. XLI: The Mystery of Oligocene Triviidae |year=2021 |journal=Conchylia |volume=52 |issue=1–2 |pages=43–54 }}
| Sp. nov | Valid | Fehse | Late Oligocene | | {{Flag|France}} | A species of Niveria. | |
| Niveria novosperara
| Sp. nov | Valid | Fehse | Late Oligocene | | {{Flag|France}} | A species of Niveria. | |
| Nodocingulum
| Gen. et comb. et 2 sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. The type species is "Pleurotomaria" coronata Münster (1841); genus also includes new species N. ernstkittli and possibly also N? turris. | |
| Nodolargena{{Cite journal |last=Garvie |first=C. L. |title=The macrofauna of the Tehuacana Limestone Member (Danian, Kincaid Formation) of Central Texas, with the Description of a few new taxa from the Pisgah Member |year=2021 |journal=Bulletins of American Paleontology |volume=399–400 |issue=399–400 |pages=1–292 |doi=10.32857/bap.2021.399.01 |doi-broken-date=1 November 2024 }}
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A member of the new family Nodolargenidae. Genus includes new species N. frosti. | |
| Nodovoluta
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species N. coronata. | |
| Odostomia malagensis
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Odostomia. | |
| Oligonalia
| Gen. et comb. nov | Valid | Lozouet | Oligocene | | {{Flag|France}} | A new genus for "Siphonalia" riparia Lozouet (1999). | |
| Olinderia
| Gen. et comb. et 3 sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A member of the family Charitodoronidae. The type species is "Charitodoron" tauzini Lozouet (1991); genus also includes new species O. lestrillensis, O. poustagnacensis and O. verdunensis. | |
| Olivella tehuacanensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Olivella. | |
| Ondina pinguis
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Ondina. | |
| Onustus undosus
| Sp. nov | Valid | Raven | Miocene | | {{Flag|Brunei}} | A species of Onustus. | |
| Opalia tehuacana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Opalia. | |
| Orbitestella kieli
| Sp. nov | Valid | Chernyshev & Goedert | Late Eocene/early Oligocene | Gries Ranch Formation | {{Flag|United States}} | A member of the family Orbitestellidae. | |
| Orthosurcula ethani
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | | |
| Oxyacrum gracilis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Pagonia
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species P. spiralis. | |
| Palaeorhaphis? laevis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Paleofusus
| Gen. et 3 sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species P. bellus, P. parvicrassus and P. elegantissima. | |
| Papuliscala? acus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Possibly a species of Papuliscala. | |
| Papuliscala keani
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | | |
| Parasassia vargai
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A member of the family Cymatiidae. | |
| Parthenina pacaudi
| Nom. nov | Valid | Landau & Micali | Miocene | | {{Flag|France}} | A species of Parthenina; a replacement name for Pyrgulina (Parthenina) dollfusi Cossmann (1921). | |
| Parvisipho? spiralis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Paskentana dixoni
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A member of Abyssochrysoidea belonging to the family Paskentanidae. | |
| Paskentana xenophontosi
| Sp. nov | Valid | Kaim et al. | Late Cretaceous (Cenomanian to Turonian) | | {{Flag|Cyprus}} | A member of Abyssochrysoidea belonging to the family Paskentanidae. | |
| Perotrochus bermotiensis
| Sp. nov | Valid | Bose, Das & Mondal | Oligocene | | {{Flag|India}} | A species of Perotrochus. | |
| Perrona harzhauseri
| Sp. nov | Valid | Kovács & Vicián | Middle Miocene | Lajta Limestone Formation | {{Flag|Hungary}} | A species of Perrona. | |
| Perrona nemethi
| Sp. nov | Valid | Kovács & Vicián | Middle Miocene | Lajta Limestone Formation | {{Flag|Hungary}} | A species of Perrona. | |
| Personopsis merlei
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A species of Personopsis. | |
| Pfefferiola
| Nom. nov | Valid | Harzhauser & Neubauer | Eocene | | {{Flag|Italy}} | A member of the family Streptaxidae; a replacement name for Oppenheimiella Pfeffer (1930). | |
| Phos estotiensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Phos. | |
| Pleurotomella irminonvilla
| Nom. nov | Valid | Pacaud | Eocene (Bartonian) | | {{Flag|France}} | A species of Pleurotomella; a replacement name for Pleurotomella cossmanni Morellet & Morellet (1946). | |
| Pliciscala obliqua
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | | |
| Poirieria (Paziella) delera
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Polinices tehuacana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Polinices. | |
| Popenoeum tabulatum
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Pressulasphaera
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" pamphilus d'Orbigny (1850). | |
| Priscoficus johnfoxi
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Priscoficus nodulina
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Proceraedis
| Gen. et 2 sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species P. imbricataria and P. tegumenis. | |
| Prososthenia esuegirottii{{Cite journal|last1=Dominici |first1=S. |last2=Forli |first2=M. |title=Lower Pliocene molluscs from southern Tuscany (Italy) |year=2021 |journal=Bollettino della Società Paleontologica Italiana |volume=60 |issue=1 |pages=69–98 |doi=10.4435/BSPI.2021.04 |doi-broken-date=2024-11-20 }}
| Sp. nov | Valid | Dominici & Forli | | | {{Flag|Italy}} | A member of the family Hydrobiidae. | |
| Protoplex? zsoldosi
| Sp. nov | Valid | Vicián & Kovács | Eocene (Lutetian) | Csernye Formation | {{Flag|Hungary}} | A member of the family Cymatiidae. | |
| Proviviparus{{Cite journal|last1=Frese |first1=M. |last2=Ponder |first2=W. |title=Proviviparus talbragarensis gen. et sp. nov., the first viviparid snail from the Late Jurassic of Australia |year=2021 |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=45 |issue=3 |pages=344–353 |doi=10.1080/03115518.2021.1940276 |bibcode=2021Alch...45..344F |s2cid=238777174 }}
| Gen. et sp. nov | Valid | Frese & Ponder | | | {{Flag|Australia}} | A member of the family Viviparidae. Genus includes new species P. talbragarensis | |
| Pseudoananias
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" subgranulata Münster (1841). | |
| Pseudokatosira uhlhingleyorum{{Cite journal|last1=Gründel |first1=J. |last2=Davies |first2=P. G. |last3=Simonsen |first3=S. |year=2021 |title=The genus Pseudokatosira Nützel & Gründel, 2007 (Gastropoda: Caenogastropoda) in the European Jurassic |journal=Paläontologie, Stratigraphie, Fazies |volume=(26), Freiberger Forschungshefte, C 559 |pages=1–16 |url=https://tu-freiberg.de/sites/default/files/media/palaeontologie-stratigraphie-fazies-2397/c559_psf26_gruendel_et_al.pdf }}
| Sp. nov | Valid | Gründel, Davies & Simonsen | Early Jurassic (Pliensbachian) | | | | |
| Pseudolatirus acuelatus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Pseudolatirus. | |
| Pseudolatirus boettgeri{{Cite journal|last=Kovács |first=Z. |title=Middle Miocene Buccinoidea (Neogastropoda) assemblage from the Făget Basin (Romania) in the collection of the Hungarian Natural History Museum, Budapest |year=2021 |journal=Fragmenta Palaeontologica Hungarica |volume=37 |pages=65–99 |doi=10.17111/FragmPalHung.2021.37.65 |s2cid=252346302 |url=https://www.researchgate.net/publication/358264811 }}
| Nom. nov | Valid | Kovács | Miocene | | {{Flag|Romania}} | A member of the family Fasciolariidae; a replacement name for Fusus sublamellosus Boettger (1906). Originally described as a species of Pseudolatirus; Harzhauser, Landau & Vermeij (2024) tentatively placed it in the genus Takashius.{{Cite journal|last1=Harzhauser |first1=M. |last2=Landau |first2=B. M. |last3=Vermeij |first3=G. J. |year=2024 |title=The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea |journal=Zootaxa |volume=5470 |issue=1 |pages=1–92 |doi=10.11646/zootaxa.5470.1.1 }} | |
| Pseudosassia
| Gen. et comb. et 2 sp. nov | Valid | Vicián & Kovács | Paleocene–Miocene | | {{Flag|Austria}} {{Flag|Belgium}} {{Flag|Bulgaria}} {{Flag|Czech Republic}} {{Flag|Denmark}} {{Flag|France}} {{Flag|Germany}} {{Flag|Hungary}} {{Flag|Italy}} {{Flag|Netherlands}} {{Flag|Poland}} {{Flag|Romania}} {{Flag|Serbia}} {{Flag|Slovakia}} {{Flag|Turkey}} {{Flag|Ukraine}} {{Flag|United Kingdom}} | A member of the family Cymatiidae. The type species is "Triton" flandricum Koninck (1838); genus also includes new species P. gurdoni and P. traceyi. | |
| Pseudoterebellum{{Cite journal|last1=Maxwell |first1=S. J. |last2=Rymer |first2=T. L. |last3=Congdon |first3=B. C. |title=Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae) |year=2021 |journal=Zootaxa |volume=4990 |issue=3 |pages=401–453 |doi=10.11646/zootaxa.4990.3.1 |pmid=34186750 |s2cid=235686526 }}
| Gen. et comb. nov | Valid | Maxwell, Rymer & Congdon | Eocene | Ghazij Shales | {{Flag|Indonesia}} {{Flag|Pakistan}} {{Flag|Saudi Arabia}} | A member of the family Seraphsidae. The type species is "Terebellum" diversiornatum Eames (1952); genus also includes "Terebellum" saudiensis Abbass (1972) and "Terebellum" olympiae Rolando (2001). | |
| Ptychatractus cataractus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Ptychatractus. | |
| Pulchraspira{{Cite journal|last1=Yu |first1=T. |last2=Salvador |first2=R. |last3=Jarzembowski |first3=E. A. |title=New terrestrial gastropods of Pupinidae and Diplommatinidae (Cyclophoroidea) from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104828 |doi=10.1016/j.cretres.2021.104828 |bibcode=2021CrRes.12404828Y |s2cid=233682256 }}
| Gen. et sp. nov | In press | Yu, Salvador & Jarzembowski | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | Genus includes new species P. teneristoma. | |
| Puposyrnola? confusa
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Possibly a species of Puposyrnola. | |
| Puposyrnola multibaca
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Puposyrnola. | |
| Puposyrnola obliqua
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Puposyrnola. | |
| Puposyrnola shirleyae
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Puposyrnola. | |
| Pusia aturensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Pusia. | |
| Pusia confunda
| Sp. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Romania}} | A species of Pusia. | |
| Pusia? falsitranssylvanica
| Nom. nov | Valid | Harzhauser & Landau | Miocene | Dej Formation | {{Flag|Romania}} | Possibly a species of Pusia; a replacement name for Mitra (Costellaria) recticosta var. transsylvanica Boettger (1902). | |
| Pusia subpyrenaica
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Pusia. | |
| Pusia virodunensis
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Pusia. | |
| Pustularia korneli
| Sp. nov | Valid | Fehse & Vicián | Miocene | | {{Flag|Indonesia}} | A species of Pustularia. | |
| Pyrgiscus elegans{{cite journal|last1=Tabanelli |first1=C. |last2=Bertaccini |first2=E. |last3=Bertamini |first3=R. |last4=Bongiardino |first4=C. |last5=Gardella |first5=F. |last6=Petracci |first6=P. |year=2021 |title=The malacofauna of the "spungone". Pyrgiscus elegans n. sp. (Mollusca: Gastropoda: Heterobranchia: Pyramidellidae) |journal=Quaderno di Studi e Notizie di Storia Naturale della Romagna |volume=54 |pages=31–37 |url=https://www.researchgate.net/publication/359866749 }}
| Sp. nov | Valid | Tabanelli et al. | Pliocene | | {{Flag|Italy}} | A species of Pyrgiscus. | |
| Pyrgiscus jaapmulderi
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Pyrgiscus. | |
| Pyrgulina marliesae
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Pyrgulina. | |
| Retusa (Cylichnina) praemoryi
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Retusa. | |
| Rhaphaulus zhuoi
| Sp. nov | In press | Yu, Salvador & Jarzembowski | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | | |
| Rinaldoella tornata
| Sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. | |
| Ringicula taenia
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | A species of Ringicula. | |
| Sansonia texana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Sassella
| Gen. et 2 sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species S. mixta and S. frosti. | |
| Scalptia nemethi{{Cite journal|last1=Kovács |first1=Z. |last2=Vicián |first2=Z. |title=New Cancellariidae (Neogastropoda) assemblages from the Middle Miocene of Hungary with description of a new species |year=2021 |journal=Central European Geology |volume=64 |issue=2 |pages=59–73 |doi=10.1556/24.2020.00006 |bibcode=2021CEJGl..64...59K |doi-access=free |hdl=10831/82983 |hdl-access=free }}
| Sp. nov | Valid | Kovács & Vicián | Miocene | | {{Flag|Hungary}} | A species of Scalptia. | |
| Scaphella kendrae{{Cite journal|last1=Petuch |first1=E. J. |last2=Berschauer |first2=D. P. |title=New fossil scaphelline volutes from the Pliocene of southern Florida |year=2021 |journal=The Festivus |volume=53 |issue=2 |pages=101–108 |doi=10.54173/F532101 |s2cid=240548109 |doi-access=free }}
| Sp. nov | Valid | Petuch & Berschauer | | | {{Flag|United States}} | A species of Scaphella. | |
| Schizogonium undae
| Sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. | |
| Seraphs kaindraperi
| Sp. nov | Valid | Maxwell, Rymer & Congdon | Eocene (Priabonian) | | {{Flag|Jamaica}} | A member of the family Seraphsidae. | |
| Similalvania
| Gen. et comb. nov | Valid | Chirli & Forli | Pliocene | | {{Flag|Italy}} | A member of the family Rissoidae. The type species is "Alvania" magistra Chirli (2006) | |
| Solariella microstriata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Solariella. | |
| Solariorbis planulatis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Squamoworthenia
| Gen. et comb. nov | Valid | Mazaev | Carboniferous | | {{Flag|Australia}} {{Flag|Germany}} {{Flag|Kazakhstan}} | Genus includes S. duponti (Holzapfel, 1889) and "Worthenia" crenilunula Yoo (1994). | |
| Stellaria striatissima
| Sp. nov | Valid | Raven | Miocene | | {{Flag|Malaysia}} | A species of Stellaria. | |
| Striacingulum
| Gen. et comb. nov | Valid | Karapunar & Nützel | | | | A member of Pleurotomariida. The type species is "Pleurotomaria" cancellatocingulata Klipstein (1844). | |
| Striaticostatum figus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Stuckenbergispira dombarensis
| Sp. nov | Valid | Mazaev | Early Carboniferous | | {{Flag|Kazakhstan}} | | |
| Sveltella novus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Sveltella. | |
| Sveltia sofiae{{Cite journal |last1=Landau |first1=B. M. |last2=da Silva |first2=C. M. |title=The genus Sveltia (Gastropoda, Cancellariidae) in the Atlantic Pliocene of Iberia with a new species from the Cenozoic Mondego Basin of Portugal |year=2021 |journal=Journal of Paleontology |volume=96 |pages=143–151 |doi=10.1017/jpa.2021.83 |s2cid=246488911 }}
| Sp. nov | Valid | Landau & da Silva | | | {{Flag|Portugal}} | A species of Sveltia. | |
| Syrnola debrae
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | A species of Syrnola. | |
| Taioma? argilla
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Taioma calxa
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Taranis occulta
| Sp. nov | Valid | Brunetti & Cresti | Pliocene | | {{Flag|Italy}} | A species of Taranis. | |
| Teinostoma regularis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Teinostoma. | |
| Terebellum humilispirum
| Sp. nov | Valid | Raven | Miocene | | {{Flag|Malaysia}} | A species of Terebellum. | |
| Terebra (Mirula) mexia
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Terebra. | |
| Terebrifusus megahani
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | | |
| Thylacodes devriesi{{Cite journal|last1=Sanfilippo |first1=R. |last2=Kočí |first2=T. |last3=Bosio |first3=G. |last4=Collareta |first4=A. |last5=Ekrt |first5=B. |last6=Malinverno |first6=E. |last7=Di Celma |first7=C. |last8=Urbina |first8=M. |last9=Bianucci |first9=G. |year=2021 |title=An investigation of vermetid reefs from the Miocene of Peru, with the description of a new species |journal=Journal of South American Earth Sciences |volume=108 |pages=Article 103233 |doi=10.1016/j.jsames.2021.103233 |bibcode=2021JSAES.10803233S |s2cid=233920362 }}
| Sp. nov | In press | Sanfilippo et al. | | | {{Flag|Peru}} | A species of Thylacodes. | |
| Tornatellaea globosa
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Tornatellaea linifera
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Tripsycha strapplei
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Tritia miniridibunda
| Sp. nov | Valid | Lozouet | Miocene | | {{Flag|France}} | A species of Tritia. | |
| Tritia praebasteroti
| Sp. nov | Valid | Lozouet | Oligocene (Chattian) | | {{Flag|France}} | A species of Tritia. | |
| Tritiaria formosa
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Trivia disjuncta
| Sp. nov | Valid | Fehse | Late Oligocene | | {{Flag|France}} | A species of Trivia. | |
| Trivia simplex
| Sp. nov | Valid | Fehse | Late Oligocene | | {{Flag|France}} | A species of Trivia. | |
| Trona colossus
| Sp. nov | Valid | Groves & Landau | Early–middle Miocene | Baitoa Formation | {{Flag|Dominican Republic}} | A species of Trona. | |
| Truncatella jiaozhouensis{{Cite journal|last1=Yu |first1=T. |last2=Salvador |first2=R. B. |last3=Wang |first3=H. |last4=Fang |first4=Y. |last5=Neubauer |first5=T. A. |last6=Li |first6=S. |last7=Zhang |first7=H. |last8=Wan |first8=X. |title=A latest Cretaceous gastropod fauna from the Jiaolai Basin of East Asia |year=2021 |journal=Cretaceous Research |volume=121 |pages=Article 104736 |doi=10.1016/j.cretres.2020.104736 |bibcode=2021CrRes.12104736Y |s2cid=233547414 }}
| Sp. nov | In press | Yu et al. | Late Cretaceous | Jiaolai Basin | {{Flag|China}} | A species of Truncatella. | |
| Tumidosipho
| Gen. et sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | Genus includes new species T. tehuacana. | |
| Turbo (Marmarostoma) ishidai{{Cite journal |last1=Tomida |first1=S. |last2=Sano |first2=H. |last3=Kase |first3=T. |title=An Additional New Species of Turbo (Marmarostoma) (Gastropoda: Turbinidae) from the Middle Miocene of the Izu Peninsula, Central Japan |year=2021 |journal=Paleontological Research |volume=25 |issue=4 |pages=375–378 |doi=10.2517/2021PR006 |s2cid=238861250 }}
| Sp. nov | Valid | Tomida, Sano & Kase | Miocene | Ena Limestone | {{Flag|Japan}} | A species of Turbo. | |
| Turbonilla bincki
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla bongiardinoi
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla crovatoi
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla malacitana
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla mauroi
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla plioalboranensis
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turbonilla velerinensis
| Sp. nov | Valid | Landau & Micali | Pliocene | | {{Flag|Spain}} | A species of Turbonilla. | |
| Turricula frosti
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Turricula. | |
| Turricula imbricata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Turricula. | |
| "Turritella" calax
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Umbilia furneauxensis{{Cite journal|last1=Southgate |first1=P. C. |last2=Militz |first2=T. A. |last3=Roberts |first3=M. |title=A new species of Umbilia Jousseaume, 1884 (Mollusca: Cypraeidae) from the Australian Pliocene |year=2021 |journal=Molluscan Research |volume=41 |issue=3 |pages=214–221 |doi=10.1080/13235818.2021.1962588 |s2cid=238231852 }}
| Sp. nov | Valid | Southgate, Militz & Roberts | | | {{Flag|Australia}} | A species of Umbilia. | |
| Unitas duomillias
| Sp. nov | Valid | Garvie | Eocene | Cook Mountain Formation | {{Flag|United States}} | | |
| Unitas euani
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Unitas petiti
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Valvata jiaolaiensis
| Sp. nov | In press | Yu et al. | Late Cretaceous | Jiaolai Basin | {{Flag|China}} | A species of Valvata. | |
| Vanikoropsis bashforthi{{Cite journal|last1=Schnetler |first1=K. I. |last2=Nielsen |first2=M. S. |title=On the genus Vanikoropsis Meek, 1876 (Gastropoda, Caenogastropoda) from the Paleocene of Denmark and West Greenland with descriptions of three new species |year=2021 |journal=Bulletin of the Geological Society of Denmark |volume=69 |pages=215–232 |doi=10.37570/bgsd-2021-69-09 |s2cid=245577704 |doi-access=free }}
| Sp. nov | Valid | Schnetler & Nielsen | Paleocene (Danian) | | {{Flag|Greenland}} | Possibly a member of the family Vanikoridae. | |
| Vanikoropsis jakobseni
| Sp. nov | Valid | Schnetler & Nielsen | Paleocene (Danian) | Agatdal Formation | {{Flag|Greenland}} | Possibly a member of the family Vanikoridae. | |
| Vanikoropsis mortenseni
| Sp. nov | Valid | Schnetler & Nielsen | Paleocene (Selandian) | Kerteminde Marl | {{Flag|Denmark}} | Possibly a member of the family Vanikoridae. | |
| Varicospira bekenuensis
| Sp. nov | Valid | Raven | Miocene | | {{Flag|Malaysia}} | A species of Varicospira. | |
| Varicospira reticulata
| Sp. nov | Valid | Raven | Miocene | | {{Flag|Malaysia}} | A species of Varicospira. | |
| Vermicularia? columella
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Vexillum transalpinum
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Hungary}} {{Flag|Romania}} {{Flag|Turkey}} | A species of Vexillum. | |
| Vitrinella discus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Vitrinella. | |
| Vitrinella praelaevis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Vitrinella. | |
| Volutifusus dougsheltoni
| Sp. nov | Valid | Petuch & Berschauer | Pliocene (Piacenzian) | | {{Flag|United States}} | A species of Volutifusus. | |
| Volutifusus kissimmeensis
| Sp. nov | Valid | Petuch & Berschauer | Pliocene (Piacenzian) | Tamiami Formation | {{Flag|United States}} | A species of Volutifusus. | |
| Volutomorpha ultima
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Wormsina
| Gen. et comb. nov | Valid | Harzhauser & Landau | Miocene | | {{Flag|Romania}} | A member of the family Mitridae. The type species is "Mitra (Cylindra)" transsylvanica Hoernes & Auinger (1880). | |
| Wortheniella klipsteini
| Sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. | |
| Wortheniella paolofedelei
| Sp. nov | Valid | Karapunar & Nützel | Late Triassic | San Cassiano Formation | {{Flag|Italy}} | A member of Pleurotomariida. | |
| Zafrona sphaerocorrugata
| Sp. nov | Valid | Harzhauser & Landau | Miocene | Baden Formation | {{Flag|Austria}} {{Flag|Bosnia and Herzegovina}} {{Flag|Bulgaria}} {{Flag|Hungary}} {{Flag|Romania}} {{Flag|Ukraine}} | A species of Zafrona. | |
| Zebinella tehuacana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Zebinella. | |
| Zonaria vokesae
| Sp. nov | Valid | Groves & Landau | Early–middle Miocene | Baitoa Formation | {{Flag|Dominican Republic}} | A species of Zonaria. | |
Bivalves
=New taxa=
| class="wikitable sortable" align="center" width="100%" |
| Name
! Novelty ! Status ! Authors ! Age ! Type locality ! Country ! Notes ! Images |
|---|
| Acar beadata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Acar. | |
| Amphidonte mirkamalovi{{Cite journal |last1=Metelkin |first1=E. K. |last2=Kosenko |first2=I. N. |title=Aptian-Albian oysters from the southwestern spurs of the Gissar Ridge. Part 1. Genus Amphidonte Fischer de Waldheim |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=608–617 |doi=10.1134/S0031030121060083 |s2cid=245010332 |url=https://www.elibrary.ru/item.asp?id=46621791 |url-access=subscription }}
| Sp. nov | Valid | Metelkin & Kosenko | Early Cretaceous (Aptian) | | | An oyster. | |
| Arcopsis subrhombus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Arcopsis tumulus
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Arkhanguelskya{{Cite journal|last1=Cooper |first1=M. R. |last2=Leanza |first2=H. A. |title=On the Buchotrigoniidae (Cretaceous Trigoniida), their palaeobiogeography, evolution and classification |year=2021 |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=301 |issue=2 |pages=119–134 |doi=10.1127/njgpa/2021/0999 |s2cid=238736681 }}
| Gen. et comb. nov | Valid | Cooper & Leanza | Cretaceous | | | A member of the family Buchotrigoniidae. Genus includes "Trigonia" weberi Arkhangelsky (1916). | |
| Baluchicardia? eoplata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Bengtsonigonia{{Cite journal|last1=Ceccolini |first1=F. |last2=Cianferoni |first2=F. |year=2021 |title=Five replacement names for fossil bivalve genera (Mollusca: Bivalvia) |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=302 |issue=2 |pages=231–234 |doi=10.1127/njgpa/2021/1025|issn=0077-7749 |s2cid=243903591 }}
| Nom. nov | Valid | Ceccolini & Cianferoni | Early Cretaceous | Riachuelo Formation | {{Flag|Brazil}} | A replacement name for Bengtsonella Cooper & Leanza (2019). | |
| Bohemigonia
| Gen. et sp. et comb. nov | Valid | Cooper | | | {{Flag|Czech Republic}} | A member of the family Megatrigoniidae. The type species is B. frici; genus also includes "Trigonia" debuchii Geinitz (1840). | |
| Caryocorbula? praefigera
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Chama ferrata{{Cite journal |last=Berezovsky |first=A. A. |title=New species of Chama (Bivalvia) from the middle Eocene of southern Ukraine |year=2021 |journal=Paleontological Journal |volume=55 |issue=4 |pages=378–383 |doi=10.1134/S0031030121040031 |s2cid=237378531 |url=https://elibrary.ru/item.asp?id=46227213 |url-access=subscription }}
| Sp. nov | Valid | Berezovsky | Eocene | | {{Flag|Ukraine}} | A species of Chama. | |
| Chama subcalcarata
| Sp. nov | Valid | Berezovsky | Eocene | | {{Flag|Ukraine}} | A species of Chama. | |
| Corbulamella palaeocenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Crassatellina minuta{{Cite journal|last1=Jaitly |first1=A. K. |last2=Kumar |first2=S. |last3=Pandey |first3=B. |last4=Gautam |first4=J. P. |last5=Tiwari |first5=D. N. |year=2021 |title=Turonian (Late Cretaceous) Heterodont Subfamilies: Lucininae, Eriphylinae and Opinae (Bivalvia: Veneroida) from the Narmada Basin, central India |journal=Journal of the Palaeontological Society of India |volume=66 |issue=2 |pages=336–345 |doi=10.1177/0971102320210217 |url=https://www.researchgate.net/publication/357478605 }}
| Sp. nov | Valid | Jaitly et al. | Late Cretaceous (Turonian) | Nodular Limestone Formation | {{Flag|India}} | A member of the family Lucinidae belonging to the subfamily Eriphylinae. | |
| Crassatellina paleocaenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Cuspidaria texana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Cuspidaria. | |
| Diplodonta frosti
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Diplodonta paleocaenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Dorbigonia
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | {{Flag|France}} | A member of the family Megatrigoniidae. Genus includes "Trigonia" disparilis d'Orbigny (1844). | |
| Elianegonia
| Nom. nov | Valid | Ceccolini & Cianferoni | Late Cretaceous | | {{Flag|Madagascar}} | A replacement name for Elianella Cooper (2015). | |
| Eomiltha strangensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Eurydesma sacculum snapperensis{{cite book |last=Waterhouse |first=J. B. |chapter=Species of the bivalve Eurydesma Morris in east Australia |year=2021 |title=Macro-faunal biozones from the early marine Permian of East Australia and New Zealand, with description of some Australian faunas |pages=57–103 |series=Earthwise |volume=19 |url=https://gsnz.org.nz/assets/Uploads/Shop/Products/Earthwise/Earthwise-19-Macro-Faunal-Biozones-from-the-Early-Marine-Permian-of-East-Australia.pdf }}
| Ssp. nov | Valid | Waterhouse | Permian | Wandrawandian Formation | {{Flag|Australia}} | A bivalve belonging to the group Pectinida and the family Eurydesmidae. | |
| Freneixicardia picturata{{Cite journal |last=Berezovsky |first=A. A. |title=New species of Cardiidae (Bivalvia) from the Eocene of Ukraine |year=2021 |journal=Paleontological Journal |volume=55 |issue=1 |pages=18–28 |doi=10.1134/S0031030121010044 |s2cid=232070780 |url=https://www.elibrary.ru/item.asp?id=44492491 |access-date=2021-01-12|url-access=subscription }}
| Sp. nov | Valid | Berezovsky | | | {{Flag|Ukraine}} | A cockle. | |
| Frenguelliella eopacifica{{Cite journal |last1=Echevarría |first1=J. |last2=Damborenea |first2=S. E. |last3=Manceñido |first3=M. O. |title=Early Jurassic Trigoniida (Bivalvia) from Argentina |year=2021 |journal=Journal of Paleontology |volume=95 |issue=Supplement S85 |pages=1–55 |doi=10.1017/jpa.2021.4|bibcode=2021JPal...95..763B |issn=0022-3360 |s2cid=238715559 |url=https://zenodo.org/record/4391425 }}
| Sp. nov | Valid | Echevarría, Damborenea & Manceñido | | | {{Flag|Argentina}} | A member of Trigoniida belonging to the family Frenguelliellidae. | |
| Glycymeris laevigata
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Glycymeris. | |
| Grammatodon superstes
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Gregariella frosti
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Gregariella. | |
| Himenourella{{Cite journal|last=Cooper |first=M. |title=On the Apiotrigoniinae (Bivalvia, Trigoniida): their palaeobiogeography, evolution and classification |year=2021 |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=300 |issue=2 |pages=145–174 |doi=10.1127/njgpa/2021/0982 |s2cid=242113854 }}
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | {{Flag|Japan}} {{Flag|Russia}} | A member of the family Megatrigoniidae. Genus includes "Trigonia" jimbonis Strand (1928), "Trigonia" sawatai Yehara (1923), "Apiotrigonia" subjimboi Miroljubov in Vereshchagin et al. (1965) and "Trigonia" subovalis Jimbo (1894). | |
| Kolymia rutskovi{{Cite journal |last1=Biakov |first1=A. S. |last2=Danukalova |first2=M. K. |last3=Kuzmichev |first3=A. B. |last4=Sobolev |first4=E. S. |title=Permian bivalves of the Pronchishchev Ridge (North Siberia): new data on taxonomic composition, biostratigraphy, and biogeographic relationships |year=2021 |journal=Paleontological Journal |volume=55 |issue=3 |pages=259–271 |doi=10.1134/S0031030121030084 |s2cid=235966259 |url=https://elibrary.ru/item.asp?id=45690930 |url-access=subscription }}
| Sp. nov | Valid | Biakov in Biakov et al. | | | {{Flag|Russia}} | | |
| Kushmurunskya
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | {{Flag|Kazakhstan}} | A member of the family Megatrigoniidae. Genus includes "Trigonia" gladkowskii Shcheglova-Borodina (1960). | |
| Kyzylkumgonia
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | | A member of the family Megatrigoniidae. Genus includes "Trigonia" syrdariensis Arkhangelsky (1916), "Trigonia" turcmenensis Arkhangelsky (1916) and "Megatrigonia" tagamensis Belyakova (1980). | |
| Loxocardium marmoreum
| Sp. nov | Valid | Berezovsky | Eocene | | {{Flag|Ukraine}} | A cockle. | |
| Lucina crassiplica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Lucina. | |
| Lucina multivittae
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Lucina. | |
| Lucina tehuacanensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Lucina. | |
| Maedagonia
| Gen. et comb. et nom. nov | Valid | Cooper | Cretaceous | | {{Flag|Japan}} {{Flag|Russia}} | A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" mikasaensis Tashiro (1979), "Apiotrigonia" ashizawaensis Saito (1962), "Apiotrigonia" crassoradiata Nakano (1957), "Apiotrigonia" futabaensis Maeda & Kawabe (1967), "Apiotrigonia" hetonaiana Tashiro (1978), "Apiotrigonia" hironoensis Maeda & Kawabe (1967), "Apiotrigonia" jimboi Nakano (1957), "Apiotrigonia" obliquecostata Nakano (1957), "Apiotrigonia" obsoleta Nakano (1957), "Apiotrigonia" orikiensis Maeda & Kawabe (1967), "Apiotrigonia" undulosa Nakano (1957) and "Apiotrigonia" utoensis Tashiro (1972), as well as M. yabei (a replacement name for Trigonia subovalis var. minor Yabe & Nagao, 1925). | |
| Martesia mazanticus{{Cite journal |last1=Castañeda-Posadas |first1=C. |last2=Trujillo-Hernández |first2=A. |last3=Zúñiga-Mijangos |first3=L. A. |title=Especie nueva del género Martesia (Bivalvia: Pholadidae) incluida en el ámbar de Chiapas como indicador de un ambiente estuarino |year=2021 |journal=Revista Mexicana de Biodiversidad |volume=92 |pages=e923600 |doi=10.22201/ib.20078706e.2021.92.3600 |url=http://revista.ib.unam.mx/index.php/bio/article/view/3600 |doi-access=free }}
| Sp. nov | Valid | Castañeda-Posadas, Trujillo-Hernández & Zúñiga-Mijangos | | | {{Flag|Mexico}} | A species of Martesia. | |
| Matsudagonia
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | {{Flag|Japan}} | A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" dubia Tashiro (1979). | |
| Moerickella
| Gen. nov | Valid | Echevarría, Damborenea & Manceñido | Early Jurassic | | {{Flag|Argentina}} | A member of Trigoniida, most likely the oldest member of the family Myophorellidae. | |
| Naiadites devonicus{{Cite journal |last1=Scholze |first1=F. |last2=Gess |first2=R. W. |title=Late Devonian non-marine Naiadites devonicus nov. sp. (Bivalvia: Pteriomorphia) from the Waterloo Farm Lagerstätte in South Africa |year=2021 |journal=Geobios |volume=69 |pages=55–67 |doi=10.1016/j.geobios.2021.07.002 |bibcode=2021Geobi..69...55S |s2cid=238797443 }}
| Sp. nov | Valid | Scholze & Gess | | | {{Flag|South Africa}} | |
| Opis reticulata
| Sp. nov | Valid | Jaitly et al. | Late Cretaceous (Turonian) | Nodular Limestone Formation | {{Flag|India}} | A member of the family Lucinidae belonging to the subfamily Opinae. | |
| Ostrea jibananandai{{Cite journal|last1=Halder |first1=K. |last2=Mitra |first2=A. |title=Facultative monogamy in an early Eocene brooding oyster and its evolutionary implications |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=3 |pages=647–662 |doi=10.4202/app.00863.2020 |doi-access=free }}
| Sp. nov | Valid | Halder & Mitra | | Cambay Shale | {{Flag|India}} | A species of Ostrea. | |
| Palaeolignopholas{{Cite journal|last1=Bolotov |first1=I. N. |last2=Aksenova |first2=O. V. |last3=Vikhrev |first3=I. V. |last4=Konopleva |first4=E. S. |last5=Chapurina |first5=Y. E. |last6=Kondakov |first6=A. V. |title=A new fossil piddock (Bivalvia: Pholadidae) may indicate estuarine to freshwater environments near Cretaceous amber-producing forests in Myanmar |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 6646 |doi=10.1038/s41598-021-86241-y |pmid=33758318 |pmc=7988128 |bibcode=2021NatSR..11.6646B }}
| Gen. et sp. nov | Valid | Bolotov et al. | Late Cretaceous (Cenomanian) | Burmese amber | {{Flag|Myanmar}} | A member of the family Pholadidae belonging to the subfamily Martesiinae. The type species is P. kachinensis. | |
| Pandora? paleocenica
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Parmicorbula? sublaevis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Perezigonia
| Nom. nov | Valid | Ceccolini & Cianferoni | Middle Jurassic | | {{Flag|Chile}} | A replacement name for Perezella Cooper (2015). | |
| Pojetaia robsonae{{Cite journal|last1=Peel |first1=J. S. |last2=Skovsted |first2=C. B. |year=2021 |title=Lower Cambrian (Series 2) small shelly fossils from along Nares Strait (Nunavut and Greenland; Laurentia) |journal=Canadian Journal of Earth Sciences |volume=58 |issue=6 |pages=495–504 |doi=10.1139/cjes-2020-0167 |bibcode=2021CaJES..58..495P |s2cid=236545098 |url=http://urn.kb.se/resolve?urn=urn:nbn:se:nrm:diva-4310 }}
| Sp. nov | Valid | Peel & Skovsted | | Kane Basin Formation | {{Flag|Canada}} | | |
| Porterius? strangensis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Poultoniella
| Gen. et comb. nov | Valid | Cooper | Cretaceous | | {{Flag|Canada}} | A member of the family Megatrigoniidae. Genus includes "Trigonia" newcombei Packard (1921). | |
| Promyophorella basoaltorum
| Sp. nov | Valid | Echevarría, Damborenea & Manceñido | Early Jurassic | | {{Flag|Argentina}} | A member of Trigoniida belonging to the family Myophorellidae. | |
| Promyophorella? sanjuanina
| Sp. nov | Valid | Echevarría, Damborenea & Manceñido | Early Jurassic | | {{Flag|Argentina}} | A member of Trigoniida belonging to the family Myophorellidae. | |
| Pseudapiotrigonia
| Gen. et comb. nov | Valid | Cooper | | | {{Flag|United States}} | A member of the family Vaugoniidae. Genus includes "Trigonia" sturgisensis Whitfield & Hovey (1906), as well as "Vaugonia" utahensis Imlay (1964). | |
| Pseudovaugonia
| Gen. nov | Valid | Echevarría, Damborenea & Manceñido | Early Jurassic | | {{Flag|Argentina}} | A member of Trigoniida belonging to the family Myophorellidae. | |
| Pteromeris nodulina
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Pteromeris. | |
| Repmanigonia
| Nom. nov | Valid | Ceccolini & Cianferoni | Jurassic | | {{Flag|Russia}} | A replacement name for Repmania Kolesnikov (1980). | |
| Schedocardia imperfecta
| Sp. nov | Valid | Berezovsky | Eocene | | {{Flag|Ukraine}} | A cockle. | |
| Sinoliaoningia
| Nom. nov | Valid | Ceccolini & Cianferoni | Late Triassic | Yangcaogou Formation | {{Flag|China}} | A replacement name for Liaoningia Yu & Dong (1993). | |
| Sokolovskya
| Gen. et comb. nov | Valid | Cooper | | | | A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" levinae Pojarkova (1990) and "Apiotrigonia" khoresmensis Belyakova (1965). | |
| Sonoragonia
| Gen. et comb. et nom. nov | Valid | Cooper | Early Cretaceous | | {{Flag|United States}} | A member of the family Megatrigoniidae. Genus includes "Trigonia" cragini Stoyanow (1949), as well as S. scotti (a replacement name for Trigonia kitchini Stoyanow, 1949). | |
| Sphenotomorpha orientalis{{Cite journal |last=Dalenz Farjat |first=A. |title=Silurian and Devonian Modiomorphidae bivalves from Bolivia |year=2021 |journal=Ameghiniana |volume=58 |issue=3 |pages=242–271 |doi=10.5710/AMGH.07.01.2021.3400 |s2cid=234229534 }}
| Sp. nov | Valid | Dalenz Farjat | Devonian (Pragian) | Icla Formation | {{Flag|Bolivia}} | A member of Modiomorphida belonging to the family Modiomorphidae. | |
| Sunetta crassatelliformis{{Cite journal|last1=Fukuda |first1=H. |last2=Ishida |first2=S. |last3=Watanabe |first3=T. |last4=Yoshimatsu |first4=S. |last5=Haga |first5=T. |title=The bivalve genus Sunetta Link, 1807 (Heterodonta: Veneridae) of Japan and the neighbouring waters – a taxonomic revision with the descriptions of three new species |year=2021 |journal=Molluscan Research |volume=41 |issue=2 |pages=107–171 |doi=10.1080/13235818.2021.1937016 |s2cid=236520379 }}
| Sp. nov | Valid | Haga & Fukuda in Fukuda et al. | Early Pleistocene | | {{Flag|Japan}} | A species of Sunetta. | |
| Sunetta nomurai
| Sp. nov | Valid | Haga & Fukuda in Fukuda et al. | Pleistocene | | {{Flag|Taiwan}} | A species of Sunetta. | |
| Syndesmyella pliominor{{Cite journal|last1=Brunetti |first1=M. M. |last2=Cresti |first2=M. |title=Syndesmyella pliominor n. sp. (Bivalvia: Semelidae) per il Pliocene spagnolo |year=2021 |journal=Bollettino Malacologico |volume=57 |issue=1 |pages=92–96 }}
| Sp. nov | Valid | Brunetti & Cresti | Early Pliocene | | {{Flag|Spain}} | A member of the family Semelidae. | |
| Tellina brevis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | A species of Tellina. | |
| Terrypoultoniella{{Cite journal |last1=Echevarría |first1=J. |last2=Damborenea |first2=S. E. |last3=Manceñido |first3=M. O. |title=Early Jurassic Trigoniida (Bivalvia) from Argentina - Addendum |year=2021 |journal=Journal of Paleontology |volume=95 |issue=Supplement S85 |page=56 |doi=10.1017/jpa.2021.97 |bibcode=2021JPal...95S..56E |s2cid=244230211 }}
| Gen. et sp. nov | Valid | Echevarría, Damborenea & Manceñido | Early Jurassic (Pliensbachian–Toarcian) | | {{Flag|Argentina}} {{Flag|Portugal}}? | A member of Trigoniida belonging to the family Frenguelliellidae. The type species is "Poultoniella" jaworskii Echevarría, Damborenea & Manceñido (2021); genus might also include "Trigonia" malladae Choffat (1885). | |
| Tindaria hamuroi{{Cite journal |last1=Amano |first1=K. |last2=Haga |first2=T. |title=A new Miocene Tindaria (Bivalvia: Tindariidae) from central Honshu, Japan |year=2021 |journal=The Nautilus |volume=135 |issue=3–4 |pages=84–88 |url=https://www.researchgate.net/publication/357033393 }}
| Sp. nov | Valid | Amano & Haga | Miocene | Higashibessho Formation | {{Flag|Japan}} | A member of the family Tindariidae. | |
| Titomaya{{Cite journal |last1=Alvarez |first1=M. J. |last2=del Río |first2=C. J. |title=A new veneroid genus (Mollusca: Bivalvia) of the Danian of Patagonia (Chubut Province, Argentina) |year=2021 |journal=Ameghiniana |volume=58 |issue=2 |pages=112–121 |doi=10.5710/AMGH.04.01.2021.3377|issn=0002-7014 |s2cid=234152656 |url=https://www.ameghiniana.org.ar/index.php/ameghiniana/article/view/3377 |url-access=subscription }}
| Gen. et comb. et sp. nov | Valid | Alvarez & del Río | | | {{Flag|Argentina}} | A member of Venerida belonging to the group Veneroidea. Genus includes "Meretrix" chalcedonica, as well as new species T. longobucca. | |
| Tivelina texana
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Tivelina trigonalis
| Sp. nov | Valid | Garvie | Paleocene (Danian) | Kincaid Formation | {{Flag|United States}} | | |
| Unionites kobyumensis{{Cite journal |last1=Biakov |first1=A. S. |last2=Kutygin |first2=R. V. |title=First record of the genus Unionites Wissmann (Bivalvia) in the Boreal Permian of northeast Russia |year=2021 |journal=Paleontological Journal |volume=55 |issue=4 |pages=372–377 |doi=10.1134/S0031030121040055 |s2cid=237377742 |url=https://elibrary.ru/item.asp?id=46227212 |url-access=subscription }}
| Sp. nov | Valid | Biakov in Biakov & Kutygin | Late Permian | | {{Flag|Russia}} | | |
| Vaccinites alencasteri{{Cite journal|last=Mitchell |first=S. F. |title=Rudist bivalves (Hippuritoidea) from the Clifton Limestone (Lower Campanian) of western Jamaica and a reassessment of the genus Vaccinites in the Americas |year=2021 |journal=Carnets de Géologie |volume=21 |issue=14 |pages=315–341 |doi=10.2110/carnets.2021.2114 |s2cid=235802666 }}
| Sp. nov | Valid | Mitchell | Late Cretaceous (probably late Turonian or early Coniacian) | Cuautla Formation | {{Flag|Mexico}} | A rudist bivalve. | |
| Vaccinites temazcali
| Sp. nov | Valid | Mitchell | Late Cretaceous (Campanian) | | {{Flag|Mexico}} | A rudist bivalve. | |
| Villamilia
| Gen. et comb. nov | Valid | Cooper & Leanza | Cretaceous | | {{Flag|Colombia}} | A member of the family Buchotrigoniidae. Genus includes "Buchotrigonia" etayoi Villamil (1992), "Trigonia" humboldtii Buch (1839), "Buchotrigonia" kauffmani Villamil (1992), "Buchotrigonia" perezi Villamil (1992) and "Vaugonia niranohamensis" santamariae Geyer (1973; raised to the rank of the species V. santamariae). | |
| Weyla santuccii{{Cite journal |last1=Damborenea |first1=S. |last2=Blodgett |first2=R. B. |last3=Hodges |first3=M. S. |last4=Hodges |first4=C. L. |title=Weyla santuccii n. sp. (Bivalvia, Pectinidae) from the Pogibshi Formation, south-central Alaska, the oldest species known of the genus in western North America |year=2021 |journal=New Mexico Museum of Natural History and Science Bulletin |volume=82 |pages=57–70 }}
| Sp. nov | Valid | Damborenea et al. | | | {{Flag|United States}} | A scallop. | |
| Whitfieldiella luceae
| Sp. nov | Valid | Mitchell | Late Cretaceous (Campanian) | Clifton Limestone | {{Flag|Jamaica}} {{Flag|United States}} | A rudist bivalve. | |
| Yunannia{{Cite journal|last1=Zhang |first1=R. |last2=Niu |first2=Z. |last3=Li |first3=C. |last4=Wang |first4=Z. |last5=Song |first5=F. |last6=He |first6=Y. |last7=Yang |first7=W. |last8=Lin |first8=X. |year=2021 |title=Yunannioidea—a new superfamily of Pteriomorphids (Bivalvia) from the Middle Ordovician, Yunkai Area, western Guangdong, South China |journal=Acta Palaeontologica Sinica |volume=60 |issue=3 |pages=Article 2020026 |doi=10.19800/j.cnki.aps.2020026 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210302 }}
| Gen. et 2 sp. nov | Valid | Zhang & Niu in Zhang et al. | | | {{Flag|China}} | A member of Pteriomorphia of uncertain phylogenetic placement. The type species is Y. gankengensis; genus also includes Y. yunkaiensis. | |
=Research=
- A study on the evolution and possible causes of the extinction of alatoconchid bivalves is published by Chen et al. (2021).{{Cite journal|last1=Chen|first1=F. |last2=Xue |first2=W. |last3=Yan |first3=J. |last4=Meng |first4=Q. |title=The implications of the giant bivalve family Alatoconchidae for the end-Guadalupian (Middle Permian) extinction event |year=2021 |journal=Geological Journal |volume=56 |issue=12 |pages=6073–6087 |doi=10.1002/gj.4151 |s2cid=234815123 }}
- A study on the arcoid bivalve diversity during the Eocene doubthouse interval of global climate cooling is published by Hickman (2021).{{Cite journal|last=Hickman |first=C. S. |title=Arcoid bivalve biodiversity during Eocene doubthouse cooling: Contrasting the active Cascadia Margin coldspot with the intracratonic Paris Basin hotspot |year=2021 |journal=PaleoBios |volume=38 |pages=ucmp_paleobios_55236 |doi=10.5070/P938055236 |s2cid=244410822 |url=https://escholarship.org/uc/item/0670875m |doi-access=free }}
Other molluscs
General research
- A study on the relationship between temperature and diversity in the fossil record of marine molluscs over the last 145 million years is published by Boag, Gearty & Stockey (2021).{{Cite journal|last1=Boag |first1=T. H. |last2=Gearty |first2=W. |last3=Stockey |first3=R. G. |title=Metabolic tradeoffs control biodiversity gradients through geological time |year=2021 |journal=Current Biology |volume=31 |issue=13 |pages=2906–2913.e3 |doi=10.1016/j.cub.2021.04.021 |pmid=33961786 |s2cid=233998188 |doi-access=free }}
- A study on the relationship between the species richness of shallow-marine molluscs, the number of ecological functional groups, and oceanic temperature in the fossil record of New Zealand over the last ~40 million years is published by Womack et al. (2021).{{Cite journal|last1=Womack |first1=T. M. |last2=Crampton |first2=J. S. |last3=Hannah |first3=M. J. |last4=Collins |first4=K. S. |title=A positive relationship between functional redundancy and temperature in Cenozoic marine ecosystems |year=2021 |journal=Science |volume=373 |issue=6558 |pages=1027–1029 |doi=10.1126/science.abf8732 |pmid=34446605 |bibcode=2021Sci...373.1027W |s2cid=237308691 }}