Lagerstätte#Types

{{Short description|Sedimentary deposit with well-preserved extraordinary fossils}}

{{Use dmy dates|date=April 2020}}

File:Opabinia_smithsonian.JPG from Burgess Shale Lagerstätte (Middle Cambrian) ]]

{{Paleontology}}

A Fossil-Lagerstätte ({{IPA|de|ˈlaːɡɐˌʃtɛtə|lang}}, from Lager 'storage, lair' Stätte 'place'; plural Lagerstätten) is a sedimentary deposit that preserves an exceptionally high amount of palaeontological information.{{cite journal| last=Shields |first=Graham |title=What are Lagerstätten |journal=Lethaia |year=1998 |volume=31 |pages=124 |url=https://www.idunn.no/doi/pdf/10.1111/j.1502-3931.1998.tb00498.x}} Konzentrat-Lagerstätten preserve a high concentration of fossils, while Konservat-Lagerstätten offer exceptional fossil preservation, sometimes including preserved soft tissues. Konservat-Lagerstätten may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. Fossil-Lagerstätten spans geological time from the Neoproterozoic era to the present.

Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Ordovician Soom Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Triassic Madygen Formation, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, & Agua Nueva formations and the Tanis Fossil Site, the Eocene Fur Formation, Green River Formation, Messel Formation & Monte Bolca, the Miocene Foulden Maar and Ashfall Fossil Beds, the Pliocene Gray Fossil Site, and the Pleistocene Naracoorte Caves & La Brea Tar Pits.

Types

Palaeontologists distinguish two kinds:The term was originally coined by Adolf Seilacher in: {{cite journal

| author = Seilacher, A.

| year = 1970

| title = Begriff und Bedeutung der Fossil-Lagerstätten: Neues Jahrbuch fur Geologie und Paläontologie

| journal = Monatshefte

| volume = 1970

| pages = 34–39

| language=de

}}The term was redefined by Julien Kimmig and James D. Schiffbauer in: {{cite journal |last=Kimmig |first=Julien |author2=James D. Schiffbauer |date=25 April 2024 |title=A modern definition of Fossil-Lagerstätten |url=https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(24)00086-7 |journal=Trends in Ecology and Evolution |volume=39 |issue=6 |pages=621–624 |doi=10.1016/j.tree.2024.04.004 |pmid=38670863 |url-access=subscription}}

  1. Konzentrat-Lagerstätten (concentration Lagerstätten) are deposits with a particular "concentration" of disarticulated organic hard parts, such as a bone bed. These Lagerstätten are less spectacular than the more famous Konservat-Lagerstätten. Their contents invariably display a large degree of time averaging, as the accumulation of bones in the absence of other sediment takes some time. Deposits with a high concentration of fossils that represent an in situ community, such as reefs or oyster beds, are not considered Lagerstätten.
  2. Konservat-Lagerstätten (conservation Lagerstätten) are deposits known for the exceptional preservation of fossilized organisms or traces. The individual taphonomy of the fossils varies with the sites. Conservation Lagerstätten are crucial in elucidating important moments in the history and evolution of life. For example, the Burgess Shale of British Columbia is associated with the Cambrian explosion, and the Solnhofen limestone with the earliest known bird, Archaeopteryx.

File:Archaeopteryx_lithographica_(Berlin_specimen).jpg specimen from Solnhofen Lagerstätte (Upper Jurassic)]]

File:Echinochimaera meltoni (fossil fish) (Bear Gulch Limestone, Upper Mississippian; Montana, USA) (33784748875).jpg from the Bear Gulch Limestone (Middle Carboniferous) ]]

Preservation

Konservat-Lagerstätten preserve lightly sclerotized and soft-bodied organisms or traces of organisms that are not otherwise preserved in the usual shelly and bony fossil record; thus, they offer more complete records of ancient biodiversity and behavior and enable some reconstruction of the palaeoecology of ancient aquatic communities. In 1986, Simon Conway Morris calculated only about 14% of genera in the Burgess Shale had possessed biomineralized tissues in life. The affinities of the shelly elements of conodonts were mysterious until the associated soft tissues were discovered near Edinburgh, Scotland, in the Granton Lower Oil Shale of the Carboniferous.Briggs et al. 1983; Aldridge et al. 1993.{{full citation needed|date=January 2022}} Information from the broader range of organisms found in Lagerstätten have contributed to recent phylogenetic reconstructions of some major metazoan groups. Lagerstätten seem to be temporally autocorrelated, perhaps because global environmental factors such as climate might affect their deposition.{{Cite journal | last1 = Retallack | first1 = G. J. | doi = 10.1016/j.palaeo.2011.04.023 | title = Exceptional fossil preservation during CO2 greenhouse crises? | journal = Palaeogeography, Palaeoclimatology, Palaeoecology | year = 2011 | volume=307 | issue = 1–4 | pages=59–74| bibcode = 2011PPP...307...59R }}

A number of taphonomic pathways may produce Konservat-Lagerstätten:{{Cite journal |last1=Kimmig |first1=Julien |last2=Schiffbauer |first2=James D. |date=2024 |title=A modern definition of Fossil-Lagerstätten |url=https://linkinghub.elsevier.com/retrieve/pii/S0169534724000867 |journal=Trends in Ecology & Evolution |volume=39 |issue=7 |pages=621–624 |language=en |doi=10.1016/j.tree.2024.04.004|pmid=38670863 |url-access=subscription }}

The identification of a fossil site as a Konservat-Lagerstätte may be based on a number of different factors which constitute "exceptional preservation". These may include the completeness of specimens, soft tissue preservation, fine-scale detail, taxonomic richness, distinctive taphonomic pathways (often multiple at the same site), the extent of the fossil layer in time and space, and particular sediment facies encouraging preservation.

Notable Lagerstätten

{{Dynamic list}}{{See also|List of fossil sites}}

The world's major Lagerstätten include:

=Precambrian=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Francevillian B Formation

|2140-2080 Ma

|Franceville, Gabon

|Possibly preserves the earliest macroscopic eukaryotes,{{cite journal |last1=Albani |first1=Abderrazak El |last2=Bengtson |first2=Stefan |last3=Canfield |first3=Donald E. |last4=Bekker |first4=Andrey |last5=Macchiarelli |first5=Roberto |last6=Mazurier |first6=Arnaud |last7=Hammarlund |first7=Emma U. |last8=Boulvais |first8=Philippe |last9=Dupuy |first9=Jean-Jacques |last10=Fontaine |first10=Claude |last11=Fürsich |first11=Franz T. |last12=Gauthier-Lafaye |first12=François |last13=Janvier |first13=Philippe |last14=Javaux |first14=Emmanuelle |last15=Ossa |first15=Frantz Ossa |last16=Pierson-Wickmann |first16=Anne-Catherine |last17=Riboulleau |first17=Armelle |last18=Sardini |first18=Paul |last19=Vachard |first19=Daniel |last20=Whitehouse |first20=Martin |last21=Meunier |first21=Alain |title=Large colonial organisms with coordinated growth in oxygenated environments 2.1 Gyr ago |journal=Nature |date=July 2010 |volume=466 |issue=7302 |pages=100–104 |doi=10.1038/nature09166|pmid=20596019 }}{{cite journal |last1=El Albani |first1=A. |last2=Konhauser |first2=K.O. |last3=Somogyi |first3=A. |last4=Ngwalghoubou Ikouanga |first4=J. |last5=Lamboux |first5=A. |last6=Blichert-Toft |first6=J. |last7=Chi-Fru |first7=E. |last8=Fontaine |first8=C. |last9=Mazurier |first9=A. |last10=Riboulleau |first10=A. |last11=Pierson-Wickmann |first11=A.-C. |last12=Albarède |first12=F. |title=A search for life in Palaeoproterozoic marine sediments using Zn isotopes and geochemistry |journal=Earth and Planetary Science Letters |date=June 2023 |volume=612 |pages=118169 |doi=10.1016/j.epsl.2023.118169}} however these may instead be pseudofossils.{{cite journal |last1=Fakhraee |first1=Mojtaba |last2=Tarhan |first2=Lidya G. |last3=Reinhard |first3=Christopher T. |last4=Crowe |first4=Sean A. |last5=Lyons |first5=Timothy W. |last6=Planavsky |first6=Noah J. |title=Earth's surface oxygenation and the rise of eukaryotic life: Relationships to the Lomagundi positive carbon isotope excursion revisited |journal=Earth-Science Reviews |date=May 2023 |volume=240 |pages=104398 |doi=10.1016/j.earscirev.2023.104398}}{{Cite journal |last1=Javaux |first1=Emmanuelle J. |last2=Lepot |first2=Kevin |date=January 2018 |title=The Paleoproterozoic fossil record: Implications for the evolution of the biosphere during Earth's middle-age |journal=Earth-Science Reviews |language=en |volume=176 |pages=68–86 |doi=10.1016/j.earscirev.2017.10.001 |bibcode=2018ESRv..176...68J |s2cid=37069547 |quote="The identity of the [Francevillian biota] macrostructures remains unknown and their biogenicity is questionable"|doi-access=free |hdl=20.500.12210/62416 |hdl-access=free }}

|File:Gabonionta I.jpg

Chuanlinggou Formation

|1630 Ma{{cite journal |last1=Miao |first1=Lanyun |last2=Yin |first2=Zongjun |last3=Knoll |first3=Andrew H. |last4=Qu |first4=Yuangao |last5=Zhu |first5=Maoyan |title=1.63-billion-year-old multicellular eukaryotes from the Chuanlinggou Formation in North China |journal=Science Advances |date=26 January 2024 |volume=10 |issue=4 |pages=eadk3208 |doi=10.1126/sciadv.adk3208|pmid=38266082 |pmc=10807817 }}

|Yan Mountains, North China

|Preserves the first definitive multicellular eukaryotes in the form of Qingshania, Tawuia and Changchengia.{{cite journal |last1=Liu |first1=Jingqi |last2=Zhang |first2=Yang |last3=Shi |first3=Xiaoying |last4=Chen |first4=Anfeng |last5=Tang |first5=Dongjie |last6=Yang |first6=Tinglu |title=Macroscopic fossils from the Chuanlinggou Formation of North China: evidence for an earlier origin of multicellular algae in the late Palaeoproterozoic |journal=Palaeontology |date=November 2023 |volume=66 |issue=6 |doi=10.1111/pala.12685}}

|File:Changchengia sp 1.png

Chitrakoot Formation

|1600 Ma

|Vindhya Range, India

|Preserves earliest red algae, Rafatazmia and Ramathallus{{cite journal |last1=Bengtson |first1=Stefan |last2=Sallstedt |first2=Therese |last3=Belivanova |first3=Veneta |last4=Whitehouse |first4=Martin |title=Three-dimensional preservation of cellular and subcellular structures suggests 1.6 billion-year-old crown-group red algae |journal=PLOS Biology |date=14 March 2017 |volume=15 |issue=3 |pages=e2000735 |doi=10.1371/journal.pbio.2000735|doi-access=free |pmid=28291791 |pmc=5349422 }}

|File:Ramathallus lobatus.png

Gaoyuzhuang Formation

|1588.8±6.5 Ma{{cite journal |last1=Chen |first1=Kai |last2=Yang |first2=Chuan |last3=Miao |first3=Lanyun |last4=Zhao |first4=Fangchen |last5=Zhu |first5=Maoyan |title=New SIMS U–Pb zircon age on the macroscopic multicellular eukaryotes from the early Mesoproterozoic Gaoyuzhuang Formation, North China |journal=Geological Magazine |date=2024 |volume=161 |doi=10.1017/S0016756824000220}}

|Yan Mountains, North China

|Preserves some of the first algae such as Tuanshanzia and Grandilingulata,{{cite journal |last1=Chen |first1=Kai |last2=Miao |first2=Lanyun |last3=Zhao |first3=Fangchen |last4=Zhu |first4=Maoyan |title=Carbonaceous macrofossils from the early Mesoproterozoic Gaoyuzhuang Formation in the Yanshan Range, North China |journal=Precambrian Research |date=July 2023 |volume=392 |pages=107074 |doi=10.1016/j.precamres.2023.107074 |language=en}} alongside a diverse microbiota including possible fungi.{{cite journal |last1=Shi |first1=Min |last2=Feng |first2=Qinglai |last3=Khan |first3=Maliha Zareen |last4=Zhu |first4=Shixing |title=An eukaryote-bearing microbiota from the early mesoproterozoic Gaoyuzhuang Formation, Tianjin, China and its significance |journal=Precambrian Research |date=December 2017 |volume=303 |pages=709–726 |doi=10.1016/j.precamres.2017.09.013}}

|File:Grandilingulata qianxiensis.png

Volyn biota

|1500 Ma

|Zhytomyr Oblast, Ukraine

|Possibly one of the first fossil sites of the deep biosphere.{{cite journal |last1=Franz |first1=Gerhard |last2=Lyckberg |first2=Peter |last3=Khomenko |first3=Vladimir |last4=Chournousenko |first4=Vsevolod |last5=Schulz |first5=Hans-Martin |last6=Mahlstedt |first6=Nicolaj |last7=Wirth |first7=Richard |last8=Glodny |first8=Johannes |last9=Gernert |first9=Ulrich |last10=Nissen |first10=Jörg |title=Fossilization of Precambrian microfossils in the Volyn pegmatite, Ukraine |journal=Biogeosciences |date=31 March 2022 |volume=19 |issue=6 |pages=1795–1811 |doi=10.5194/bg-19-1795-2022|doi-access=free }}{{cite journal |last1=Franz |first1=Gerhard |last2=Khomenko |first2=Vladimir |last3=Lyckberg |first3=Peter |last4=Chournousenko |first4=Vsevolod |last5=Struck |first5=Ulrich |last6=Gernert |first6=Ulrich |last7=Nissen |first7=Jörg |title=The Volyn biota (Ukraine) – indications of 1.5 Gyr old eukaryotes in 3D preservation, a spotlight on the "boring billion" |journal=Biogeosciences |date=24 May 2023 |volume=20 |issue=10 |pages=1901–1924 |doi=10.5194/bg-20-1901-2023|doi-access=free }}

|File:Volyn biota - Franz et al., 2023, fig. 3j.png

Hunting Formation

|1200 Ma

|Somerset Island, Canada

|Shallow-water Mesoproterozoic deposits, containing multiple genera of algae{{cite journal |last1=Butterfield |first1=Nicholas J. |title=Bangiomorpha pubescens n. gen., n. sp.: implications for the evolution of sex, multicellularity, and the Mesoproterozoic/Neoproterozoic radiation of eukaryotes |journal=Paleobiology |date=2000 |volume=26 |issue=3 |pages=386–404 |doi=10.1666/0094-8373(2000)026<0386:BPNGNS>2.0.CO;2}}{{cite journal |last1=Butterfield |first1=Nicholas J |title=Paleobiology of the late Mesoproterozoic (ca. 1200 Ma) Hunting Formation, Somerset Island, arctic Canada |journal=Precambrian Research |date=October 2001 |volume=111 |issue=1–4 |pages=235–256 |doi=10.1016/S0301-9268(01)00162-0}} including Bangiomorpha.

|File: Bangiomorpha pubescens ; clustered uniseriate filaments with basal holdfasts (Lower Hunting Formation, Stenium Period, Somerset Island, Canadian Arctic Islands).png

Nonesuch Formation

|1083-1070 Ma

|Michigan, USA

|An oxygenated Mesoproterozoic lake{{Cite journal |last1=Slotznick |first1=Sarah P. |last2=Swanson-Hysell |first2=Nicholas L. |last3=Zhang |first3=Yiming |last4=Clayton |first4=Katherine E. |last5=Wellman |first5=Charles H. |last6=Tosca |first6=Nicholas J. |last7=Strother |first7=Paul K. |date=22 August 2023 |title=Reconstructing the paleoenvironment of an oxygenated Mesoproterozoic shoreline and its record of life |url=https://pubs.geoscienceworld.org/gsabulletin/article/doi/10.1130/B36634.1/627730/Reconstructing-the-paleoenvironment-of-an |journal=Geological Society of America Bulletin |language=en |doi=10.1130/B36634.1 |issn=0016-7606 |access-date=21 May 2024 |via=GeoScienceWorld}} containing exceptionally preserved limnic microbes.{{cite journal |last1=Strother |first1=Paul K. |last2=Wellman |first2=Charles H. |date=30 November 2020 |title=The Nonesuch Formation Lagerstätte: a rare window into freshwater life one billion years ago |journal=Journal of the Geological Society |volume=178 |issue=2 |pages=1–12 |doi=10.1144/jgs2020-133 |s2cid=229003508 |doi-access=free }}

|

Lakhanda Group

|

1030-1000 Ma

|

Uchur-Maya Depression, Russia

|A site preserving a Mesoproterozoic community dominated by anaerobic bacteria.{{Cite journal |last1=Duda |first1=Jan-Peter |last2=König |first2=Hannah |last3=Reinhardt |first3=Manuel |last4=Shuvalova |first4=Julia |last5=Parkhaev |first5=Pavel |date=8 December 2021 |title=Molecular fossils within bitumens and kerogens from the ~ 1 Ga Lakhanda Lagerstätte (Siberia, Russia) and their significance for understanding early eukaryote evolution |journal=PalZ |language=en |volume=95 |issue=4 |pages=577–592 |doi=10.1007/s12542-021-00593-4 |issn=0031-0220 |doi-access=free }} The lagerstätte contains evidence of trophic interactions from the Boring Billion.{{cite journal |last1=Shuvalova |first1=J. V. |last2=Nagovitsin |first2=K. E. |last3=Parkhaev |first3=P. Yu. |date=26 February 2021 |title=Evidences of the Oldest Trophic Interactions in the Riphean Biota (Lakhanda Lagerstätte, Southeastern Siberia) |url=https://link.springer.com/article/10.1134/S0012496621010105 |journal=Doklady Biological Sciences |volume=496 |issue=1 |pages=34–40 |doi=10.1134/S0012496621010105 |pmid=33635488 |s2cid=254411264 |access-date=30 April 2023|url-access=subscription }}{{cite journal |last1=Podkovyrov |first1=Victor N. |date=September 2009 |title=Mesoproterozoic Lakhanda Lagerstätte, Siberia: Paleoecology and taphonomy of the microbiota |url=https://www.sciencedirect.com/science/article/abs/pii/S0301926809000965 |journal=Precambrian Research |volume=173 |issue=1–4 |pages=146–153 |doi=10.1016/j.precamres.2009.04.006 |bibcode=2009PreR..173..146P |access-date=30 April 2023|url-access=subscription }}

|

Bitter Springs

|

1000–850 Ma

|

South Australia

|

Preserved fossils include cyanobacteria microfossils.

|

File:Inzeria intia (stromatolite) (Neoproterozoic, ~800 Ma; Bitter Springs, Australia) 1 (41559714260).jpg]]

Diabaig Formation

|994 ± 48 Ma{{Cite journal |last1=Turnbull |first1=Matthew J. M. |last2=Whitehouse |first2=Martin J. |last3=Moorbath |first3=Stephen |date=November 1996 |title=New isotopic age determinations for the Torridonian, NW Scotland |url=https://www.lyellcollection.org/doi/10.1144/gsjgs.153.6.0955 |journal=Journal of the Geological Society |language=en |volume=153 |issue=6 |pages=955–964 |doi=10.1144/gsjgs.153.6.0955 |issn=0016-7649 |access-date=21 May 2024|url-access=subscription }}

|Scotland

|A freshwater environment preserving phosphatic microfossils,{{Cite journal |last1=Battison |first1=Leila |last2=Brasier |first2=Martin D. |date=February 2012 |title=Remarkably preserved prokaryote and eukaryote microfossils within 1Ga-old lake phosphates of the Torridon Group, NW Scotland |url=https://linkinghub.elsevier.com/retrieve/pii/S0301926811002749 |journal=Precambrian Research |language=en |volume=196-197 |pages=204–217 |doi=10.1016/j.precamres.2011.12.012 |access-date=7 July 2024 |via=Elsevier Science Direct|url-access=subscription }} which represent some of the oldest known non-marine eukaryotes.{{Cite journal |last1=Nielson |first1=Grace C. |last2=Stüeken |first2=Eva E. |last3=Prave |first3=Anthony R. |date=February 2024 |title=Estuaries house Earth's oldest known non-marine eukaryotes |url=https://linkinghub.elsevier.com/retrieve/pii/S0301926823003182 |journal=Precambrian Research |language=en |volume=401 |pages=107278 |doi=10.1016/j.precamres.2023.107278 |access-date=21 May 2024 |via=Elsevier Science Direct|hdl=10023/28949 |hdl-access=free }}

|

Dolores Creek Formation

|950 Ma

|Yukon, Canada

|An Early Tonian site containing pyritised macroalgal fossils.{{cite journal |last1=Maloney |first1=Katie M. |last2=Schiffbauer |first2=James D. |last3=Halverson |first3=Galen P. |last4=Xiao |first4=Shuhai |last5=Laflamme |first5=Marc |date=13 April 2022 |title=Preservation of early Tonian macroalgal fossils from the Dolores Creek Formation, Yukon |journal=Scientific Reports |volume=12 |issue=1 |page=6222 |doi=10.1038/s41598-022-10223-x |pmid=35418588 |pmc=9007953 |bibcode=2022NatSR..12.6222M }}

Svanbergfjellet Formation

|

791.1±4.9 Ma{{cite journal |last1=Mughal |first1=Sanaa |last2=Millikin |first2=Alexie E. G. |last3=Zhang |first3=Tianran |last4=Gibson |first4=Timothy M. |last5=Rooney |first5=Alan D. |last6=Tosca |first6=Nicholas J. |last7=Bergmann |first7=Kristin D. |last8=Strauss |first8=Justin V. |last9=Anderson |first9=Ross P. |title=The Svanbergfjellet Formation: eukaryotic life in a Tonian Sea |journal=Journal of the Geological Society |date=5 May 2025 |volume=182 |issue=3 |doi=10.1144/jgs2024-083|url=https://figshare.com/articles/journal_contribution/28151958 }}

|

Svalbard

|A site preserving some of the earliest green algae, such as Proterocladus, alongside various enigmatic eukaryotes like Pseudotawuia.{{cite book |last1=Butterfield |first1=Nicholas J. |last2=Knoll |first2=Andrew H. |last3=Swett |first3=Keene |title=Paleobiology of the Neoproterozoic Svanbergfjellet Formation, Spitsbergen |series=Fossils and Strata |date=15 July 1994 |volume=34 |pages=1–84 |doi=10.18261/8200376494-1994-01|isbn=82-00-37649-4 }}

|

Chichkan Lagerstätte

|

775 Ma

|

Kazakhstan

|A site from the transition between the prokaryote-dominated biota of the Early Neoproterozoic and the eukaryote-dominated biota of the Late Neoproterozoic and Phanerozoic.{{cite journal |last1=Sergeev |first1=Vladimir N. |last2=Schopf |first2=J. William |title=Taxonomy, paleoecology and biostratigraphy of the late Neoproterozoic Chichkan microbiota of South Kazakhstan: The marine biosphere on the eve of metazoan radiation |date=1 May 2010 |url=https://pubs.geoscienceworld.org/jpaleontol/article-abstract/84/3/363/108791/TAXONOMY-PALEOECOLOGY-AND-BIOSTRATIGRAPHY-OF-THE |journal=Journal of Paleontology |volume=84 |issue=3 |pages=363–401 |doi=10.1666/09-133.1 |bibcode=2010JPal...84..363S |s2cid=140668863 |access-date=30 April 2023|url-access=subscription }}

|

Doushantuo Formation

|

600–555 Ma

|

Guizhou Province, China

|

Spans the poorly understood interval between the end of the Cryogenian period and the late Ediacaran Avalon explosion.

|File:Ediacaran embryo-like fossil.jpg-like fossil]]

Portfjeld Formation

|570 Ma

|North Greenland

|A Middle Ediacaran biota from the continent of Laurentia exhibiting Doushantuo-type preservation.{{cite journal |last1=Willman |first1=Sebastian |last2=Peel |first2=John S. |last3=Ineson |first3=Jon R. |last4=Schovsbo |first4=Niels H. |last5=Rugen |first5=Elias J. |last6=Frei |first6=Robert |date=6 November 2020 |title=Ediacaran Doushantuo-type biota discovered in Laurentia |journal=Communications Biology |volume=3 |issue=1 |page=647 |doi=10.1038/s42003-020-01381-7 |pmid=33159138 |pmc=7648037 }}

Mistaken Point

|

565 Ma

|

Newfoundland, Canada

|

This site contains one of the most diverse and well-preserved collections of Precambrian fossils.

|

File:Ediacaran fossils Mistaken Point Newfoundland.jpg ediacaran organisms]]

Itajai Biota

|563 Ma

|Brazil

|An Ediacaran lagerstatte preserved by volcanism.{{Cite journal |last1=Becker-Kerber |first1=Bruno |last2=El Albani |first2=Abderrazak |last3=Konhauser |first3=Kurt |last4=Abd Elmola |first4=Ahmed |last5=Fontaine |first5=Claude |last6=Paim |first6=Paulo S. G. |last7=Mazurier |first7=Arnaud |last8=Prado |first8=Gustavo M. E. M. |last9=Galante |first9=Douglas |last10=Kerber |first10=Pedro B. |last11=da Rosa |first11=Ana L. Z. |last12=Fairchild |first12=Thomas R. |last13=Meunier |first13=Alain |last14=Pacheco |first14=Mírian L. A. F. |date=3 March 2021 |title=The role of volcanic-derived clays in the preservation of Ediacaran biota from the Itajaí Basin (ca. 563 Ma, Brazil) |journal=Scientific Reports |language=en |volume=11 |issue=1 |page=5013 |doi=10.1038/s41598-021-84433-0 |issn=2045-2322 |pmc=7930025 |pmid=33658558 }}

|

Ediacara Hills

|

555 Ma

|

South Australia

|

The type location the Ediacaran period, and has preserved a significant amount of fossils from that time.

|

File:Spriggina floundersi - MUSE.jpg, a worm-like organism]]

Shibantan Lagerstätte

|551-543 Ma

|Hubei, China

|A terminal Ediacaran fossil assemblage preserving life forms living just before the Proterozoic-Phanerozoic transition.{{cite journal |last1=Xiao |first1=Shuhai |last2=Chen |first2=Zhe |last3=Pang |first3=Ke |last4=Zhou |first4=Chuanming |last5=Yuan |first5=Xunlai |date=12 October 2020 |title=The Shibantan Lagerstätte: insights into the Proterozoic–Phanerozoic transition |url=https://www.lyellcollection.org/doi/full/10.1144/jgs2020-135 |journal=Journal of the Geological Society |volume=178 |pages=1–11 |doi=10.1144/jgs2020-135 |s2cid=225242787 |access-date=30 April 2023|url-access=subscription }}

|

Gaojiashan Lagerstätte

|551-541 Ma

|Shaanxi, China

|A lagerstätte documenting tube growth patterns of Cloudina.{{cite journal |last1=Cai |first1=Yaoping |last2=Hua |first2=Hong |last3=Schiffbauer |first3=James D. |last4=Sun |first4=Bo |last5=Yuan |first5=Xunlai |date=April 2014 |title=Tube growth patterns and microbial mat-related lifestyles in the Ediacaran fossil Cloudina, Gaojiashan Lagerstätte, South China |url=https://www.sciencedirect.com/science/article/abs/pii/S1342937X1300052X |journal=Gondwana Research |volume=25 |issue=3 |pages=1008–1018 |doi=10.1016/j.gr.2012.12.027 |bibcode=2014GondR..25.1008C |access-date=18 June 2023|url-access=subscription }}

|

Jiucheng Member

|551-543 Ma

|Yunnan, China

|A latest Ediacaran macrofossil biota dominated by giant, unbranching thallophytes.{{Cite journal |last1=Feng |first1=Tang |last2=Chongyu |first2=Yin |last3=Pengju |first3=Liu |last4=Linzhi |first4=Gao |last5=Wenyan |first5=Zhang |date=7 September 2010 |title=A New Diverse Macrofossil Lagerstätte from the Uppermost Ediacaran of Southwestern China |url=https://onlinelibrary.wiley.com/doi/10.1111/j.1755-6724.2008.tb00709.x |journal=Acta Geologica Sinica - English Edition |language=en |volume=82 |issue=6 |pages=1095–1103 |doi=10.1111/j.1755-6724.2008.tb00709.x |issn=1000-9515 |access-date=21 May 2024|url-access=subscription }}

|

Khatyspyt Lagerstätte

|

544 Ma

|

Yakutia, Russia

|

A Late Ediacaran lagerstätte preserving an Avalon-type biota.{{cite journal |last1=Duda |first1=Jan-Peter |last2=Love |first2=Gordon D. |last3=Rogov |first3=Vladimir I. |last4=Melnyk |first4=Dmitry S. |last5=Blumenberg |first5=Martin |last6=Grazhdankin |first6=Dmitriy V. |date=3 September 2020 |title=Understanding the geobiology of the terminal Ediacaran Khatyspyt Lagerstätte (Arctic Siberia, Russia) |journal=Geobiology |volume=18 |issue=6 |pages=643–662 |doi=10.1111/gbi.12412 |pmid=32881267 |bibcode=2020Gbio...18..643D |s2cid=221477265 |doi-access=free }}

|

Bernashivka open pit

|? (Upper Vendian)

|Vinnytsia Oblast, Ukraine

|A Late Ediacaran lagerstätte with numerous soft-bodied animals, algae, microfossils, bacteria, and fungi, comprising a number of different geological formations.{{Cite journal |last1=National Museum of Natural History, National Academy of Sciences of Ukraine (Kyiv, Ukraine) |last2=Grytsenko |first2=Volodymyr |date=2020-10-02 |title=Diversity of the Vendian fossils of Podillia (Western Ukraine) |url=http://museumkiev.org/public/visnyk/19_2020/gb1903-grytsenko.html |journal=Geo&Bio |volume=2020 |issue=19 |pages=3–19 |doi=10.15407/gb1903|doi-access=free }}

|

=Cambrian=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Zhangjiagou

|Fortunian

|Shaanxi, China

|A lagerstätte from the earliest Cambrian notable for its fossils of cnidarians,{{Cite journal |last1=Shao |first1=Tiequan |last2=Tang |first2=Hanhua |last3=Liu |first3=Yunhuan |last4=Waloszek |first4=Dieter |last5=Maas |first5=Andreas |last6=Zhang |first6=Huaqiao |date=March 2018 |title=Diversity of cnidarians and cycloneuralians in the Fortunian (early Cambrian) Kuanchuanpu Formation at Zhangjiagou, South China |url=https://www.cambridge.org/core/product/identifier/S0022336017000944/type/journal_article |journal=Journal of Paleontology |language=en |volume=92 |issue=2 |pages=115–129 |doi=10.1017/jpa.2017.94 |issn=0022-3360 |access-date=14 May 2024 |via=Cambridge Core|url-access=subscription }} cycloneuralians,{{Cite journal |last1=Shao |first1=T. Q. |last2=Qin |first2=J. C. |last3=Shao |first3=Y. |last4=Liu |first4=Y. H. |last5=Waloszek |first5=D. |last6=Maas |first6=A. |last7=Duan |first7=B. C. |last8=Wang |first8=Q. |last9=Xu |first9=Y. |last10=Zhang |first10=H. Q. |date=October 2020 |title=New macrobenthic cycloneuralians from the Fortunian (lowermost Cambrian) of South China |url=https://linkinghub.elsevier.com/retrieve/pii/S0301926819301366 |journal=Precambrian Research |language=en |volume=349 |pages=105413 |doi=10.1016/j.precamres.2019.105413 |access-date=14 May 2024 |via=Elsevier Science Direct|url-access=subscription }}{{Cite journal |last1=Liu |first1=Yunhuan |last2=Qin |first2=Jiachen |last3=Wang |first3=Qi |last4=Maas |first4=Andreas |last5=Duan |first5=Baichuan |last6=Zhang |first6=Yanan |last7=Zhang |first7=Hu |last8=Shao |first8=Tiequan |last9=Zhang |first9=Huaqiao |date=19 October 2018 |editor-last=Zhang |editor-first=Xi-Guang |title=New armoured scalidophorans (Ecdysozoa, Cycloneuralia) from the Cambrian Fortunian Zhangjiagou Lagerstätte, South China |url=https://onlinelibrary.wiley.com/doi/10.1002/spp2.1239 |journal=Papers in Palaeontology |language=en |volume=5 |issue=2 |pages=241–260 |doi=10.1002/spp2.1239 |issn=2056-2799 |access-date=14 May 2024 |via=Wiley Online Library|url-access=subscription }} and the basal ecdysozoan Saccorhytus coronarius.{{Cite journal |last1=Han |first1=Jian |last2=Morris |first2=Simon Conway |last3=Ou |first3=Qiang |last4=Shu |first4=Degan |last5=Huang |first5=Hai |date=9 February 2017 |title=Meiofaunal deuterostomes from the basal Cambrian of Shaanxi (China) |url=https://www.nature.com/articles/nature21072 |journal=Nature |language=en |volume=542 |issue=7640 |pages=228–231 |doi=10.1038/nature21072 |pmid=28135722 |issn=0028-0836 |access-date=14 May 2024|url-access=subscription }}

|File:Saccorhytus_microscopic.jpg of Saccorhytus as it may have been seen through the lens of a microscope, living among grains of sand.]]

Maotianshan Shales (Chengjiang)

|

518 Ma

|

Yunnan, China

|

The preservation of an extremely diverse faunal assemblage renders the Maotianshan Shales the world's most important formation for understanding the evolution of early multi-cellular life. Microscopic animals like Yicaris are preserved here, showing the presence of an Orsten-type deposit within the formation.{{cite journal |last1=Zhang |first1=Xi-guang |last2=Siveter |first2=David J. |last3=Waloszek |first3=Dieter |last4=Maas |first4=Andreas |title=An epipodite-bearing crown-group crustacean from the Lower Cambrian |journal=Nature |date=4 October 2007 |volume=449 |issue=7162 |pages=595–598 |doi=10.1038/nature06138|pmid=17914395 }}

|

File:Haikouichthys4.png, a primitive craniate]]

Qingjiang biota

|

518 Ma

|

Hubei, China

|

This site is particularly notable due to both the large proportion of new taxa represented (approximately 53% of the specimens), and the notable volume of soft-body tissue preservation.

|

File:Duplapex.png, a hymenocarine arthropod]]

Sirius Passet

|

523-518 Ma

|

Greenland

|

A site known for its fauna, and that they were most likely preserved by a death mask. It is a part of the larger Buen Formation, and has a fauna similar to the Maotianshan shales.

|

File:Kerygmachela kierkegaardi. MGUH 32054.jpg, a dinocaridid-like lobopodian]]

Xiaoshiba Lagerstätte

|Cambrian Stage 3

|Yunnan, China

|A site known for its detailed preservation of Early Cambrian macroalgae,{{Cite journal |last1=Lan |first1=Tian |last2=Yang |first2=Jie |last3=Zhang |first3=Xi-guang |last4=Hou |first4=Jin-bo |date=15 June 2018 |title=A new macroalgal assemblage from the Xiaoshiba Biota (Cambrian Series 2, Stage 3) of southern China |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018217308969 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=499 |pages=35–44 |doi=10.1016/j.palaeo.2018.02.029 |issn=0031-0182 |access-date=13 September 2024 |via=Elsevier Science Direct|url-access=subscription }} alongside fossils of Omnidens.{{cite journal |last1=Li |first1=Wei |last2=Yang |first2=Jie |last3=Yang |first3=Xiaoyu |last4=Dhungana |first4=Alavya |last5=Wang |first5=Yu |last6=Zhang |first6=Xiguang |last7=Smith |first7=Martin R. |title=Omnidens appendages and the origin of radiodont mouthparts |journal=Papers in Palaeontology |date=November 2024 |volume=10 |issue=6 |doi=10.1002/spp2.1600}}

|File:Omnidens qiongqii fossil.jpg

Poleta Formation

|519-518 Ma

|Nevada, USA

|The middle member of the formation preserves the Indian Springs Lagerstätte, one of the oldest such sites from former Laurentia. This site preserves a diversity of mineralized organisms such as trilobites and brachiopods, but also non-mineralized remains such as sponges, algae, and soft-bodied arthropods.{{Cite journal |last1=English |first1=Adam M. |last2=Babcock |first2=Loren E. |date=2010-09-01 |title=Census of the Indian Springs Lagerstätte, Poleta Formation (Cambrian), western Nevada, USA |url=https://www.sciencedirect.com/science/article/pii/S0031018210003287 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=295 |issue=1 |pages=236–244 |doi=10.1016/j.palaeo.2010.05.041 |issn=0031-0182|url-access=subscription }}

|File:Nevadia weeksi CRF.jpg]]

Sinsk Algal Lens

|

518 Ma

|

Yakutia, Russia

|

One of the oldest known Cambrian lagerstätten. The fauna of this site is unique, as it seems that they were adapted to living in dysaerobic conditions.{{cite journal |last1=Ivantsov |first1=Andrey Yu. |last2=Zhuravlev |first2=Andrey Yu. |last3=Leguta |first3=Anton V. |last4=Krassilov |first4=Valentin A. |last5=Melnikova |first5=Lyudmila M. |last6=Ushatinskaya |first6=Galina T. |date=2 May 2005 |title=Palaeoecology of the Early Cambrian Sinsk biota from the Siberian Platform |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018204005784#! |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=220 |issue=1–2 |pages=69–88 |doi=10.1016/j.palaeo.2004.01.022 |bibcode=2005PPP...220...69I |access-date=12 November 2022|url-access=subscription }}

|

File:20191215 Siberion lenaicus.png]]

Guanshan Biota

|

Cambrian Stage 4

|

Eastern Yunnan, China

|

A relatively diverse lagerstätte within the Wulongqing Formation, between the more famous Maotianshan Shales and Burgess Shale in age, alongside having both taxa from the previous two formations and completely new genera/species preserved here. It is also unusual for being quite shallow and having a brachiopod-dominated fauna.{{cite journal |last1=Chen |first1=Feiyang |last2=Brock |first2=Glenn A. |last3=Zhang |first3=Zhiliang |last4=Laing |first4=Brittany |last5=Ren |first5=Xinyi |last6=Zhang |first6=Zhifei |title=Brachiopod-dominated communities and depositional environment of the Guanshan Konservat-Lagerstätte, eastern Yunnan, China |journal=Journal of the Geological Society |date=January 2021 |volume=178 |issue=1 |doi=10.1144/jgs2020-043|doi-access=free }}

|

File:S41467-020-16332-3 Fig1.webp

Cranbrook Lagerstätte

|Cambrian Stage 4

|British Columbia

|One of the oldest Burgess Shale-type biotas of North America.{{Cite journal |last1=Caron |first1=Jean-Bernard |last2=Webster |first2=Mark |last3=Briggs |first3=Derek E. G. |last4=Pari |first4=Giovanni |last5=Santucci |first5=Guy |last6=Mángano |first6=M. Gabriela |last7=Izquierdo-López |first7=Alejandro |last8=Streng |first8=Michael |last9=Gaines |first9=Robert R. |date=13 December 2023 |title=The lower Cambrian Cranbrook Lagerstätte of British Columbia |url=https://www.lyellcollection.org/doi/10.1144/jgs2023-106 |journal=Journal of the Geological Society |language=en |volume=181 |issue=1 |doi=10.1144/jgs2023-106 |issn=0016-7649 |access-date=7 July 2024 |via=Lyell Collection Geological Society Publications|url-access=subscription }}

|

Tatelt Formation

|515 Ma

|High Atlas, Morocco

|A layer in this formation has produced some of the most well-preserved trilobites ever discovered, with preserved internal organs, feeding structures, and articulated appendages. The trilobites were likely rapidly buried and preserved by a volcanic eruption.{{Cite web |last=Weisberger |first=Mindy |date=2024-07-09 |title='Prehistoric Pompeii' reveals 515 million-year-old sea bugs' anatomy in pristine 3D |url=https://www.cnn.com/2024/07/09/science/trilobite-fossils-discovery-volcanic-ash/index.html |access-date=2024-07-13 |website=CNN |language=en}}

|File:Gigoutella-MHNM-HA-TT-CA-1A ventral fossil.jpg]]

Emu Bay Shale

|

513 Ma

|

South Australia

|

Noted soft tissue mineralization, most often of blocky apatite or fibrous calcium carbonate, including the oldest phosphatized muscle tissue.

|

File:Oestokerkus megacholix reconstruction.png, a megacheiran arthropod]]

Parker Slate

|513-511 Ma

|Vermont, US

|A Burgess Shale-type biota with rare but exceptionally preserved soft-bodied animals. The earliest Burgess Shale-type biota to be described, being documented 25 years before the Burgess Shale itself.{{Cite journal |last1=Pari |first1=Giovanni |last2=Briggs |first2=Derek E.G. |last3=Gaines |first3=Robert R. |date=2021-02-22 |title=The Parker Quarry Lagerstätte of Vermont—The first reported Burgess Shale–type fauna rediscovered |url=https://doi.org/10.1130/G48422.1 |journal=Geology |volume=49 |issue=6 |pages=693–697 |doi=10.1130/g48422.1 |issn=0091-7613|url-access=subscription }}

|

Kaili Formation

|

513–501 Ma

|

Guizhou, China

|

The middle part of the Kaili Formation, the Oryctocephalus indicus Zone, contains a Burgess Shale-type lagerstätte with many well-preserved fossils known collectively as the Kaili Biota.

|

File:Rotadiscus guizhouensis (fossil eldonioid) (Kaili Formation, Middle Cambrian; quarry between Balang & Dunzhou, Guizhou Province, China).jpg a cambroernid animal]]

Murero Lagerstätte

|511-503 Ma

|

Spain

|

Thanks to the paleontological content, mainly trilobites, fourteen biozones have been established, the most precise biozonation for this time interval in the world. It also records in detail the so-called Valdemiedes event, the mass extinction episode at the end of the Lower Cambrian.Gámez Vintaned, J. A.; Liñán, E. y Gozalo, R. (2013) «Los trilobites cámbricos de la Biota de Murero (Zaragoza, España)». Cuadernos de Paleontología Aragonesa, 7: 5-27

|

File:Eccaparadoxides mediterraneus - Murero, Zaragoza - Museo Ciencias Naturales Universidad Zaragoza.jpg trilobites showing sexual dimorphism]]

Blackberry Hill

|

~510–500 Ma

|

Central Wisconsin, US

|This site preserves some of the oldest evidence of multicellular life walking out of the ocean, and onto dry land (in the form of large mollusks and euthycarcinoid arthropods). Other notable fossils include stranded scyphozoans, and some of the oldest true crustaceans (in the form of phyllocarids).

|

File:Mosineia macnaughtoni.jpg, a euthycarcinoid arthropod]]

Henson Gletscher Formation

|Wuliuan

|North Greenland

|A phosphatised lagerstätte preserving hatching priapulid larvae, pentastomids{{cite journal |last1=Peel |first1=John S. |title=The oldest tongue worm: a stem-group pentastomid arthropod from the early middle Cambrian (Wuliuan Stage) of North Greenland (Laurentia) |journal=GFF |date=3 April 2022 |volume=144 |issue=2 |pages=97–105 |doi=10.1080/11035897.2022.2064543}} and abundant bradoriid and phosphatocopid arthropods.{{Cite journal |last1=Department of Earth Sciences (Palaeobiology), Uppsala University, Sweden. |last2=Peel |first2=John S. |date=7 June 2023 |title=A phosphatised fossil Lagerstätte from the middle Cambrian (Wuliuan Stage) of North Greenland (Laurentia) |url=https://2dgf.dk/publikationer/bulletin/bulletin-volume-72-2023/#3 |journal=Bulletin of the Geological Society of Denmark |volume=72 |pages=101–122 |doi=10.37570/bgsd-2023-72-03 |access-date=7 July 2024|doi-access=free }}

|

Burgess Shale

|

508 Ma

|

British Columbia, Canada

|

One of the most famous fossil localities in the world. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints.

|

File:Aria et al. 2020 Anomalocaris canadensis head ROMIP 51212.png, a predatory radiodont]]

Duchesnay Formation

|

~505 Ma?

|

Southeastern British Columbia

|

Similar to the Burgess Shale above (both formations even have similar depositional environments next to escarpments), but seemingly lower-diversity and slightly younger. Several organisms are shared between both formations, although the Duchesnay Formation preserves several new genera. Brine pools are also preserved.{{cite journal |last1=Johnston |first1=Kimberley J. |last2=Johnston |first2=Paul A. |last3=Powell |first3=Wayne G. |title=A new, Middle Cambrian, Burgess Shale-type biota, Bolaspidella Zone, Chancellor Basin, southeastern British Columbia |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |date=June 2009 |volume=277 |issue=1–2 |pages=106–126 |doi=10.1016/j.palaeo.2009.02.015}}

|

Spence Shale

|

507 Ma

|

Northeastern Utah, Southeastern Idaho, US

|

A site known for its abundant Cambrian trilobites and the preservation of Burgess Shale-type fossils. The type locality for this site is Spence Gulch in southeastern Idaho.

|

File:Hyoliths01.JPG, most likely lophophorates]]

Linyi Lagerstätte

|

504 Ma

|

Shandong, China

|

A lagerstätte recognised for its exceptional preservation of arthropod limbs, intestines, and eyes.{{Cite journal |last1=Sun |first1=Z. |last2=Zhao |first2=F. |last3=Zeng |first3=H. |last4=Luo |first4=C. |last5=Van Iten |first5=H. |last6=Zhu |first6=M. |year=2022 |title=The middle Cambrian Linyi Lagerstätte from the North China Craton: a new window on the Cambrian evolutionary fauna |journal=National Science Review |url=https://academic.oup.com/nsr/article/9/7/nwac069/6563905 |volume=9 |issue=7 |pages=nwac069 |doi=10.1093/nsr/nwac069 |pmid=35832778 |pmc=9273334 |access-date=2 April 2023}}

|File:Life on the platform margin of the Miaolingian sea, North China.png

Ravens Throat River Lagerstätte

|Drumian

|Northwest Territories

|A Burgess Shale-type biota coeval in age with the more famous Wheeler Shale and Marjum Formation.{{Cite journal |last1=Kimmig |first1=Julien |last2=Pratt |first2=Brian R. |date=29 July 2015 |title=Taphonomy of the middle Cambrian (Drumian) Ravens Throat River Lagerstätte, Rockslide Formation, Mackenzie Mountains, Northwest Territories, Canada |url=https://www.idunn.no/doi/10.1111/let.12135 |journal=Lethaia |language=en |volume=49 |issue=2 |pages=150–169 |doi=10.1111/let.12135 |issn=0024-1164|url-access=subscription }}

|

Wheeler Shale (House Range)

|

504 Ma

|

Western Utah, US

|

A world-famous locality known for its prolific agnostid and Elrathia kingii trilobite remains. Varied soft bodied organisms are also locally preserved, including Naraoia, Wiwaxia and Hallucigenia.

|

File:Elrathia kingii growth series.jpg sp. trilobites]]

Marjum Formation

|

502 Ma

|

Western Utah, US

|

A site known for its occasional preservation of soft-bodied tissue, and diverse assemblage.

|

File:Branchiocaris KUMIP204797.png a pelagic hymenocarine]]

Weeks Formation

|

500 Ma

|

Western Utah, US

|

A site that is dominated by trilobites and brachiopods, but also comprising various soft-bodied organisms, such as Falcatamacaris.

|

File:Beckwithia collection Paul Hille.jpg, an aglaspidid trilobite relative]]

Kinnekulle Orsten and Alum Shale

|

500 Ma

|

Sweden

|

The Orsten sites reveals the oldest well-documented benthic meiofauna in the fossil record. Fossils such as microfossils of arthropods like free-living pentastomids are known. Multiple "Orsten-type" lagerstätten are also known from other countries.

|

File:Cambropachycope.png, a stem-group mandibulate arthropod]]

=Ordovician=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Fezouata Formation

|

about 485 Ma

|

Draa Valley, Morocco

|

It was deposited in a marine environment, and is known for its exceptionally preserved fossils, filling an important preservational window beyond the earlier and more common Cambrian Burgess shale-type deposits.

|

File:20191205 Aegirocassis benmoulai Aegirocassis benmoulae.png, a giant hurdiid radiodont]]

Cabrieres biota

|Floian

|Montagne Noire, France

|A polar marine ecosystem from the Early Ordovician that likely served as a refuge from the high temperatures of the epoch.{{Cite journal |last1=Saleh |first1=Farid |last2=Lustri |first2=Lorenzo |last3=Gueriau |first3=Pierre |last4=Potin |first4=Gaëtan J.-M. |last5=Pérez-Peris |first5=Francesc |last6=Laibl |first6=Lukáš |last7=Jamart |first7=Valentin |last8=Vite |first8=Antoine |last9=Antcliffe |first9=Jonathan B. |last10=Daley |first10=Allison C. |last11=Nohejlová |first11=Martina |last12=Dupichaud |first12=Christophe |last13=Schöder |first13=Sebastian |last14=Bérard |first14=Emilie |last15=Lynch |first15=Sinéad |last16=Drage |first16=Harriet B. |last17=Vauchier |first17=Romain |last18=Vidal |first18=Muriel |last19=Monceret |first19=Eric |last20=Monceret |first20=Sylvie |last21=Lefebvre |first21=Bertrand |date=9 February 2024 |title=The Cabrières Biota (France) provides insights into Ordovician polar ecosystems |journal=Nature Ecology & Evolution |language=en |volume=8 |issue=4 |pages=651–662 |doi=10.1038/s41559-024-02331-w |pmid=38337049 |pmc=11009115 |issn=2397-334X }}

|

Liexi fauna

|

About 470 Ma (early-middle Floian)

|

Hunan Province, China

|Preserves Early Ordovician fauna with soft tissue, includes not only Cambrian relics but also taxa originated during Ordovician.{{Cite journal |last1=Fang |first1=Xiang |last2=Mao |first2=Yingyan |last3=Liu |first3=Qi |last4=Yuan |first4=Wenwei |last5=Chen |first5=Zhongyang |last6=Wu |first6=Rongchang |last7=Li |first7=Lixia |last8=Zhang |first8=Yuchen |last9=Ma |first9=Junye |last10=Wang |first10=Wenhui |last11=Zhan |first11=Renbin |last12=Peng |first12=Shanchi |last13=Zhang |first13=Yuandong |last14=Huang |first14=Diying |date=2022-07-13 |title=The Liexi fauna: a new Lagerstätte from the Lower Ordovician of South China |journal=Proceedings of the Royal Society B: Biological Sciences |language=en |volume=289 |issue=1978 |doi=10.1098/rspb.2022.1027 |issn=0962-8452 |pmc=9277276 |pmid=35858062}}

|

Castle Bank

|

About 461 Ma

|

Llandrindod Wells, Wales

|

A unique environment deposited during the middle Ordovician that possibly shows iconic groups from Cambrian lagerstättes, like Opabiniids and Megacheirans, survived for longer than what was thought.

|

File:20221119 Mieridduryn bonniae diagrammatic reconstruction.png, an opabiniid-like panarthropod]]

Douglas Dam Member{{Cite journal |last1=Young |first1=Graham |last2=Rudkin |first2=David |last3=Dobrzanski |first3=Edward |last4=Robson |first4=Sean |last5=Cuggy |first5=Michael |last6=Demski |first6=Matthew |last7=Thompson |first7=Deborah |date=2012 |title=Great Canadian Lagerstätten 3. Late Ordovician Konservat-Lagerstätten in Manitoba |url=https://www.erudit.org/en/journals/geocan/2012-v39-n4-geocan39_4/geocan39_4ser01/ |journal=Geoscience Canada |language=en |volume=39 |issue=4 |pages=201–213 |issn=0315-0941}}

|

460 Ma

|

Tennessee, US

|Low-diversity assemblage of arthropod fossils, which are preserved well because of volcanic ash.

|

File:20200606_Chasmataspis_laurencii.png, a horseshoe crab-like chasmataspid arthropod]]

Winneshiek Shale

|

460 Ma

|

Decorah, Iowa

|

A Middle Ordovician site confined to a large impact Crater that is known for exceptionally exquisite preservation of conodonts, bivalved arthropods, and the earliest eurypterids in the fossil record.{{cite journal |last1=Liu |first1=Huaibao P. |last2=Bergström |first2=Stig M. |last3=Witzke |first3=Brian J. |last4=Briggs |first4=Derek E. G. |last5=McKay |first5=Robert M. |last6=Ferretti |first6=Annalisa |date=28 February 2017 |title=Exceptionally preserved conodont apparatuses with giant elements from the Middle Ordovician Winneshiek Konservat-Lagerstätte, Iowa, USA |journal=Journal of Paleontology |volume=91 |issue=3 |pages=493–511 |doi=10.1017/jpa.2016.155 |bibcode=2017JPal...91..493L |s2cid=132698401 |doi-access=free }}

|File:Eurypterids Pentecopterus Vertical.jpg, the oldest known eurypterid]]

Beecher's Trilobite Bed

|

460? Ma

|

New York, US

|

Noted exceptionally preserved trilobites with soft tissue preserved by pyrite replacement.
Pyritisation allows the use of X-rays to study fine detail of preserved soft body parts.

|

File:Triarthrus.jpg trilobites with pyritized soft-tissue]]

Harding Sandstone

|

? (Sandbian)

|

Colorado, US

|Although preservation is not excellent, this lagoonal site provides early vertebrate fossils such as Astraspis and Eriptychius.

|

File:Overview_of_Eriptychius_americanus_PF_1795.png

Walcott-Rust Quarry

|

about 455? Ma

|

New York, US

|

This site is an excellent example of an obrution (rapid burial or "smothered") Lagerstätte.
Unique preservation of trilobite appendages resulted from early cementation of the surrounding rock and spar filling of the interior cavity of the appendages.

|

File:Isotelus gigas fossil trilobite (Ordovician; Walcott-Rust site, New York State, USA) (16763508314).jpg, a large asaphid trilobite]]

Big hill Lagerstätte

|about 450? Mya

|Michigan, US

|A site known for its preservation of soft-bodied medusae (jellyfish), as well as linguloid brachiopods, algae, and arthropods (namely chasmataspidids, leperditid ostracods, and eurypterids).

|File:20200605 Hoplitaspis hiawathai.png, chasmataspid arthropod]]

Brechin Lagerstätte

|

450 Ma

|

Ontario, Canada

|

Known for preserving one of the most diverse crinoid fauna of the Katian.{{Cite journal|author1=Selina R. Cole |author2=David F. Wright |author3=William I. Ausich |date=1 May 2019 |title=Phylogenetic community paleoecology of one of the earliest complex crinoid faunas (Brechin Lagerstätte, Ordovician) |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=521 |pages=82–98 |doi=10.1016/j.palaeo.2019.02.006 |bibcode=2019PPP...521...82C |s2cid=135129430 |doi-access=free }}

|File:Tomlinsonus.png, a marrellomorph]]

Soom Shale

|

450? Ma

|

South Africa

|

Known for its remarkable preservation of soft-tissue in fossil material. Deposited in still waters, the unit lacks bioturbation, perhaps indicating anoxic conditions.

|

File:Promissum NT small.jpg, a conodont known from rare soft-tissues]]

Tafilalt Biota

|

? (Sandbian-middle Katian)

|

Tafilalt, Morocco

|

Known from range of non-biomineralised and soft-bodied organisms in polar environment.{{Cite journal |last1=García-Bellido |first1=Diego C. |last2=Gutiérrez-Marco |first2=Juan Carlos |date=2022-10-29 |title=Polar gigantism and remarkable taxonomic longevity in new palaeoscolecid worms from the Late Ordovician Tafilalt Lagerstätte of Morocco |url=http://dx.doi.org/10.1080/08912963.2022.2131404 |journal=Historical Biology |volume=35 |issue=11 |pages=2011–2021 |doi=10.1080/08912963.2022.2131404 |s2cid=253297071 |issn=0891-2963|url-access=subscription }}

|

File:Eldonia_berbera.jpg" barvera, an eldoniid]]

Cat Head Member

|

? (middle Katian)

|

Manitoba, Canada

|Fossils like algae, conulariids and trilobites are known from this site.

|

Vauréal Formation{{cite journal |last1=Knaust |first1=Dirk |last2=Desrochers |first2=André |title=Exceptionally preserved soft-bodied assemblage in Ordovician carbonates of Anticosti Island, eastern Canada |journal=Gondwana Research |date=July 2019 |volume=71 |pages=117–128 |doi=10.1016/j.gr.2019.01.016}}

|

Katian

|

Anticosti Island, Quebec

|Preserves earliest known acoelomorphs, nemerteans, nematodes and flatworms in the fossil record

|

Georgian Bay Formation

|

449-445.6 Ma

|

Manitoulin District, Canada

|Low-diversity assemblage of arthropod fossils.

|

File:Orcanopterus.png, a middle-sized waeringopterid eurypterid]]

William Lake (Stony Mountain Formation)

|

445 Ma

|

Manitoba, Canada

|Well-preserved fossils like jellyfish, xiphosurans, sea spiders are known from this site, it is important since many of the fossils are unknown in other Ordovician sites.

| rowspan="2" |

File:20200813_Lunataspis_aurora.png aurora, an early xiphosuran known in two sites]]

Airport Cove

|

445 Ma

|

Manitoba, Canada

|Fossils like eurypterids, algae and xiphosurans are preserved in this site.

=Silurian=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Kalana Lagerstätte

|

~440 Ma
Aeronian

|

Estonia

|

Known for well preserved fossils of algae and crinoids,{{cite journal |last1=Ausich |first1=William I. |last2=Wilson |first2=Mark A. |last3=Tinn |first3=Oive |date=14 May 2019 |title=Kalana Lagerstätte crinoids: Early Silurian (Llandovery) of central Estonia |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/kalana-lagerstatte-crinoids-early-silurian-llandovery-of-central-estonia/B6F78AF4B4DC8CD22133091426A77E27 |journal=Journal of Paleontology |volume=94 |issue=1 |pages=131–144 |doi=10.1017/jpa.2019.27 |s2cid=181399467 |access-date=27 April 2023|url-access=subscription }} along with an osteostracan fossilised via an extremely unusual carbonaceous mode of preservation that was previously unknown among vertebrates.{{cite journal |last1=Tinn |first1=Oive |last2=Lang |first2=Liisa |last3=Märss |first3=Tiiu |last4=Vahur |first4=Signu |last5=Kirsimäe |first5=Kalle |date=6 May 2022 |title=A demineralized osteostracan fossil from the Silurian Kalana Lagerstätte of Estonia: revealing its internal anatomy and uncovering a unique type of fossilization |journal=Lethaia |volume=55 |issue=1 |pages=1–13 |doi=10.1111/let.12452 |doi-access=free }}

|File:Kalanaspis delectabilis.jpg, the earliest known osteostracan]]

Chongqing Lagerstätte (Huixingshao Formation){{Cite journal |last1=Zhu |first1=You-an |last2=Li |first2=Qiang |last3=Lu |first3=Jing |last4=Chen |first4=Yang |last5=Wang |first5=Jianhua |last6=Gai |first6=Zhikun |last7=Zhao |first7=Wenjin |last8=Wei |first8=Guangbiao |last9=Yu |first9=Yilun |last10=Ahlberg |first10=Per E. |last11=Zhu |first11=Min |date=2022 |title=The oldest complete jawed vertebrates from the early Silurian of China |url=https://www.nature.com/articles/s41586-022-05136-8 |journal=Nature |language=en |volume=609 |issue=7929 |pages=954–958 |doi=10.1038/s41586-022-05136-8 |pmid=36171378 |bibcode=2022Natur.609..954Z |s2cid=252569910 |issn=1476-4687|url-access=subscription }}

|

436 Ma

|

Chongqing, China

|This site preserved complete fossils of earliest jawed vertebrates, as well as some galeaspids and eurypterids.

|

File:Xiushanosteus.jpg, the earliest placoderm]]

Waukesha Biota (Brandon Bridge Formation)

|

~435 Ma
Early Silurian

|

Wisconsin, US

|

Well-studied site known for the exceptional preservation of its diverse, soft-bodied and lightly skeletonized fauna, includes many major taxa found nowhere else in strata of similar age. It was one of the first fossil sites with soft bodied preservation known to science.

|

File:20211029 Parioscorpio venator.png, an enigmatic arthropod]]

Herefordshire Lagerstätte (Coalbrookdale Formation)

|

~430 Ma

|

Herefordshire, UK

|

Known for the well-preserved fossils of various invertebrate animals many of which are in their three-dimensional structures. Fossils are preserved within volcanic ash, because of that sometimes this site has been compared to Pompeii.{{Cite web |last=BBC |title=Fossils found in 425 million year old 'Pompeii' |url=https://www.bbc.co.uk/herefordandworcester/content/articles/2009/06/01/herefordshire_fossils_feature.shtml |access-date=2023-10-07 |website=www.bbc.co.uk |language=en-gb}} Some of the fossils are regarded as earliest evidences and evolutionary origin of some of the major groups of modern animals.

|

File:20200810 Offacolus kingi.png, a euchelicerate]]

Eramosa Lagerstätte

|

~425 Ma

|

Ontario & New York (state)

|

Known for preservation of both hard and soft bodied organisms in great detail, including early scorpions, eurypterids, agnathan vertebrates, and several other species.

|File:Eramoscorpius_fossil.png, an early scorpion]]

Bertie Group

|

422.9-416 Ma

|

Ontario & New York State

|

This limestone have produced thousands of fossil eurypterids, such as giant Acutiramus and well-known Eurypterus, as well as other fauna like scorpions and fish.

|File:Nerepisacanthus.png, an acanthodian]]

Tonoloway Formation{{Cite journal |last1=Vrazo |first1=Matthew B. |last2=Trop |first2=Jeffrey M. |last3=Brett |first3=Carlton E. |date=2014 |title=A New Eurypterid Lagerstätte from the Upper Silurian of Pennsylvania |url=https://www.jstor.org/stable/43683811 |journal=PALAIOS |volume=29 |issue=7/8 |pages=431–448 |doi=10.2110/palo.2014.003 |jstor=43683811 |bibcode=2014Palai..29..431V |s2cid=30066085 |issn=0883-1351|url-access=subscription }}

|

~420 Ma

|

Pennsylvania, US

|Known from exceptionally preserved mass assemblage of Eurypterus, the most abundant eurypterid in the fossil record.

|File:Eurypterus_Paleoart.jpg

Rochester Shale

|

415 Ma{{Citation |last=Taylor |first=Wendy L. |title=Middle Silurian Rochester Shale of Western New York, USA, and Southern Ontario, Canada |date=1999-10-28 |url=http://dx.doi.org/10.1017/cbo9780511626159.012 |work=Fossil Crinoids |pages=87–92 |access-date=2023-10-09 |publisher=Cambridge University Press|doi=10.1017/cbo9780511626159.012 |isbn=9780521450249 |url-access=subscription }}

|

New York, US and Ontario, Canada

|Echinoderms (such as crinoids) and trilobites are known from Lewiston Member in this shale.

|File:Arctinurus_boltoni_(Rochester_Shale,_Middle_Silurian;_Middleport,_New_York_State,_USA)_(15273506362).jpg, one of the trilobites from this site]]

=Devonian=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Ditton Group

|Lochkovian

|Wales, UK

|A wildfire-derived lagerstätte containing pyrolysis products of ancient wildfires.{{Cite journal |last1=Glasspool |first1=I.J. |last2=Edwards |first2=D. |last3=Axe |first3=L. |date=December 2006 |title=Charcoal in the Early Devonian: A wildfire-derived Konservat–Lagerstätte |url=https://www.sciencedirect.com/science/article/abs/pii/S0034666706001369 |journal=Review of Palaeobotany and Palynology |language=en |volume=142 |issue=3–4 |pages=131–136 |doi=10.1016/j.revpalbo.2006.03.021 |access-date=8 April 2025 |via=Elsevier Science Direct|url-access=subscription }}{{Citation |last1=Morris |first1=Jennifer L. |title=The Advantages and Frustrations of a Plant Lagerstätte as Illustrated by a New Taxon From the Lower Devonian of the Welsh Borderland, UK |date=2018 |work=Transformative Paleobotany |pages=49–67 |url=https://linkinghub.elsevier.com/retrieve/pii/B9780128130124000048 |access-date=2025-04-13 |publisher=Elsevier |language=en |doi=10.1016/b978-0-12-813012-4.00004-8 |isbn=978-0-12-813012-4 |last2=Edwards |first2=Dianne |last3=Richardson |first3=John B.|url-access=subscription }}

|

Rhynie chert400 MaScotland, UK

|The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material (euthycarcinoids, branchiopods, arachnids, hexapods, etc) preserved in place by an overlying volcanic deposit and hot springs. As well as one of the first known fully terrestrial ecosystems.

|File:Asteroxylon_life_restoration.jpg, an early vascular plant related to lycopods]]

Waxweiler Lagerstätte (Klerf Formation)409-392 MaEifel, Germany

|Waxweiler Lagerstätte is known from well-preserved fossils of chelicerates, giant claw of Jaekelopterus rhenaniae shows the largest arthropod ever known.

|File:20210106_Jaekelopterus_rhenaniae.png

Heckelmann Mill395 MaRhineland-Palatinate, Germany

|Heckelmann Mill preserves well preserved rhinocaridid archaeostracan phyllocarids,{{cite journal |last1=Poschmann |first1=Markus J. |date=3 February 2021 |title=A new phyllocarid (Crustacea, Archaeostraca) from the Early Devonian (late Emsian) Heckelmann Mill Fossil-Lagerstätte (Lahn Syncline, Rhineland-Palatinate, SW-Germany) |url=https://link.springer.com/article/10.1007/s12542-020-00535-6 |journal=PalZ |volume=95 |pages=27–36 |doi=10.1007/s12542-020-00535-6 |s2cid=231793893 |access-date=29 April 2023|url-access=subscription }} along with exceptionally abundant crinoid holdfasts from the late Emsian.{{cite journal |last1=Poschmann |first1=Markus J. |date=15 July 2019 |title=Pelmatozoan "rooting grounds" from the Early Devonian (late Emsian) Heckelmann Mill Fossil-Lagerstätte (Lahn Syncline, Rhineland-Palatinate, SW-Germany): morphological, palaeoenvironmental and taphonomic aspects |url=https://link.springer.com/article/10.1007/s12542-019-00454-1 |journal=PalZ |volume=94 |issue=2 |pages=311–325 |doi=10.1007/s12542-019-00454-1 |s2cid=198137119 |access-date=29 April 2023|url-access=subscription }}

Hunsrück Slates (Bundenbach)390 MaRheinland-Pfalz, Germany

|The Hunsrück slates are one of the few marine Devonian lagerstätte having soft tissue preservation, and in many cases fossils are coated by a pyritic surface layer.

|File:20210708 Schinderhannes bartelsi diagrammatic reconstruction.png, the youngest known radiodont]]

Gogo Formation380 Ma (Frasnian)Western Australia

|The fossils of the Gogo Formation display three-dimensional soft-tissue preservation of tissues as fragile as nerves and embryos with umbilical cords. Over fifty species of fish have been described from the formation, and arthropods.

|File:Materpiscis.png, a ptyctodontid placoderm fish that is the oldest vertebrate known to give live birth]]

Miguasha National Park (Escuminac Formation)370 MaQuébec, Canada

|Some of the fish, fauna, and spore fossils found at Miguasha are rare and ancient species. For example, Eusthenopteron is sarcopterygian that shares characters with early tetrapods.

|File:Scaumenacia_curta_&_Bothriolepis_canadensis.JPG and antiarch placoderm Bothriolepis]]

Kowala Lagerstätte~368 MaŚwiętokrzyskie Voivodeship, Poland

|A Late Devonian site known for its fossils of non-biomineralised algae and arthropods.{{cite journal |last1=Zatoń |first1=Michał |last2=Filipiak |first2=Paweł |last3=Rakociński |first3=Michał |last4=Krawczyński |first4=Wojciech |date=25 March 2014 |title=Kowala Lagerstätte: Late Devonian arthropods and non-biomineralized algae from Poland |url=https://onlinelibrary.wiley.com/doi/abs/10.1111/let.12062 |journal=Lethaia |volume=47 |issue=3 |pages=352–364 |doi=10.1111/let.12062 |access-date=29 June 2023|url-access=subscription }}

|

Maïder Basin368 Ma (for Thylacocephalan Layer)Anti-Atlas, Morocco

|Thylacocephalan Layer and Hangenberg Black Shale in this basin provides well-preserved fossils of Famennian fauna, including chondrichthyans and placoderms that preserved soft tissues.{{Cite journal |last1=Frey |first1=Linda |last2=Pohle |first2=Alexander |last3=Rücklin |first3=Martin |last4=Klug |first4=Christian |date=2020 |title=Fossil-Lagerstätten, palaeoecology and preservation of invertebrates and vertebrates from the Devonian in the eastern Anti-Atlas, Morocco |url=https://www.idunn.no/doi/10.1111/let.12354 |journal=Lethaia |language=en |volume=53 |issue=2 |pages=242–266 |doi=10.1111/let.12354 |s2cid=210627752 |issn=0024-1164|url-access=subscription }}

|File:Amazichthys.jpg, pelagic placoderm with preserved soft tissue]]

Strud{{Cite journal |last1=Denayer |first1=Julien |last2=Prestianni |first2=Cyrille |last3=Gueriau |first3=Pierre |last4=Olive |first4=Sébastien |last5=Clément |first5=Gaël |date=2016 |title=Stratigraphy and depositional environments of the Late Famennian (Late Devonian) of Southern Belgium and characterization of the Strud locality |url=https://www.cambridge.org/core/journals/geological-magazine/article/abs/stratigraphy-and-depositional-environments-of-the-late-famennian-late-devonian-of-southern-belgium-and-characterization-of-the-strud-locality/BAFD866A0B080EAFA24836F73F437EBD |journal=Geological Magazine |language=en |volume=153 |issue=1 |pages=112–127 |doi=10.1017/S001675681500031X |bibcode=2016GeoM..153..112D |s2cid=129070535 |issn=0016-7568|url-access=subscription }}? (Late Famennian)Namur Province, Belgium

|Mainly juvenile placoderms are known, suggesting this site would be nursery site of placoderms.{{Cite journal |last1=Olive |first1=Sébastien |last2=Clément |first2=Gaël |last3=Daeschler |first3=Edward B. |last4=Dupret |first4=Vincent |date=2016-08-23 |title=Placoderm Assemblage from the Tetrapod-Bearing Locality of Strud (Belgium, Upper Famennian) Provides Evidence for a Fish Nursery |journal=PLOS ONE |language=en |volume=11 |issue=8 |pages=e0161540 |doi=10.1371/journal.pone.0161540 |issn=1932-6203 |pmc=4994939 |pmid=27552196 |bibcode=2016PLoSO..1161540O |doi-access=free }} Various biota like tetrapods, arthropods and plants are also known, Strudiella from this site may be the earliest insect, but its affinity is disputed.

|File:Strudops_goldenbergi.jpg, the earliest known notostracan.]]

Canowindra, New South Wales (Mandagery Sandstone)360 MaAustralia

|An accidentally discovered lagerstätte known for its exceptional preservation of Sarcopterygian and Placoderm fish.

|File:Mandageria_skull.png, a lobe-finned fish that is the NSW State Fossil Emblem in Australia]]

Waterloo Farm Lagerstätte (Witpoort Formation)360 MaSouth Africa

|Important site that providing the only record of a high latitude (near polar) coastal ecosystem, overturning numerous assumptions about high latitude conditions during the latest Devonian.

|File:Priscomyzon_riniensis02.jpg, the oldest known genus of lamprey]]

=Carboniferous=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Granton Shrimp Bed? (Dinantian)Firth of Forth, Scotland

|Dominated by well-preserved crustacean fossils, this site provided first body fossil of Clydagnathus which solved long-lasted mystery of conodont fossils.

|

East Kirkton Quarry{{Cite book |last1=Fraser |first1=Nicholas C. |url=https://books.google.com/books?id=JysFngEACAAJ |title=Terrestrial Conservation Lagerstätten: Windows Into the Evolution of Life on Land |last2=Sues |first2=Hans-Dieter |date=2017 |publisher=Dunedin Academic Press |isbn=978-1-78046-014-7 |language=en}}335 MaWest Lothian, Scotland

|This site has produced numerous well-preserved fossils of early tetrapods like temnospondyls or reptiliomorphs, and large arthropods like scorpions or eurypterids.

|File:Silvanerpeton1DB.jpg, a possible reptiliomorph]]

Bear Gulch Limestone324 MaMontana, US

|A limestone-rich geological lens in central Montana. It is renowned for its unusual and ecologically diverse fossil composition of chondrichthyans, the group of cartilaginous fish containing modern sharks, rays, and chimaeras. Other animals like brachiopods, ray finned fish, arthropods, and the possible mollusk Typhloesus are also known from the site.

|File:Falcatus_falcatus_01.JPG, a holocephalian which males had large fin spine]]

Wamsutta Formation

|~320-318 Ma

|Massachusetts

|A subhumid alluvial fan deposit that preserves ichnofossils, plants, invertebrates, and vertebrates.{{Cite journal |last1=Knecht |first1=Richard J. |last2=Benner |first2=Jacob S. |last3=Swain |first3=Anshuman |last4=Azevedo-Schmidt |first4=Lauren |last5=Cleal |first5=Christopher J. |last6=Labandeira |first6=Conrad C. |last7=Engel |first7=Michael S. |last8=Dunlop |first8=Jason A. |last9=Selden |first9=Paul A. |last10=Eble |first10=Cortland F. |last11=Renczkowski |first11=Mark D. |last12=Wheeler |first12=Dillon A. |last13=Funderburk |first13=Mataeus M. |last14=Emma |first14=Steve L. |last15=Knoll |first15=Andrew H. |last16=Pierce |first16=Naomi E. |date=9 September 2024 |title=Early Pennsylvanian Lagerstätte reveals a diverse ecosystem on a subhumid, alluvial fan |journal=Nature Communications |language=en |volume=15 |issue=1 |pages=7876 |doi=10.1038/s41467-024-52181-0 |pmid=39251605 |issn=2041-1723 |pmc=11383953 }}

|

Bickershaw{{Cite journal |last1=Anderson |first1=Lyall I. |last2=Dunlop |first2=Jason A. |last3=Horrocks |first3=Carl A. |last4=Winkelmann |first4=Heather M. |last5=Eagar |first5=R. M. C. |date=1997 |title=Exceptionally preserved fossils from Bickershaw, Lancashire UK (Upper Carboniferous, Westphalian A (Langsettian)) |url=https://onlinelibrary.wiley.com/doi/10.1002/(SICI)1099-1034(199709)32:33.0.CO;2-6 |journal=Geological Journal |language=en |volume=32 |issue=3 |pages=197–210 |doi=10.1002/(SICI)1099-1034(199709)32:3<197::AID-GJ739>3.0.CO;2-6|url-access=subscription }}? (Langsettian)Lancashire, England

|This locality contains exceptionally preserved fossils within nodules. Arthropods have greater diversity, many of which are aquatic ones that lived in brackish environment.

|File:Valloisella_lievinensis.JPG, a xiphosuran]]

Joggins Fossil Cliffs (Joggins Formation)315 MaNova Scotia, Canada

|A fossil site that preserves a diverse terrestrial ecosystem consisting of plants like lycopsids, giant arthropods, fish, and the oldest known sauropsid, Hylonomus.

|File:Hylonomus_lyelli_-_MUSE.jpg

Castlecomer fauna

|315-307

|Kilkenny, Ireland

|A konservat-lagerstätte with a high number of well-preserved spinicaudatan clam shrimp.{{Cite journal |last1=Orr |first1=Patrick J. |last2=Briggs |first2=Derek E. G. |last3=Parkes |first3=Matthew A. |date=1996 |title=The 'Castlecomer Fauna': A New Konservat-Lagerstätte from the Upper Carboniferous of Ireland |url=https://www.jstor.org/stable/30002319 |journal=Irish Journal of Earth Sciences |volume=15 |pages=93–106 |jstor=30002319 |issn=0790-1763}}{{Cite journal |last1=Orr |first1=P. J. |last2=Briggs |first2=D. E.G. |last3=Kearns |first3=S. L. |date=2008-05-01 |title=Taphonomy of Exceptionally Preserved Crustaceans from the Upper Carboniferous of Southeastern Ireland |url=https://pubs.geoscienceworld.org/palaios/article/23/5/298-312/145985 |journal=PALAIOS |language=en |volume=23 |issue=5 |pages=298–312 |doi=10.2110/palo.2007.p07-015r |issn=0883-1351|url-access=subscription }}

|

Linton Diamond Coal Mine{{Cite web|url=https://gsa.confex.com/gsa/2008NE/finalprogram/abstract_135299.htm|title=Direct Evidence of Food Chains at the Linton Lagerstatte|website=gsa.confex.com}}{{Cite thesis |title=Patterns and drivers of tetrapod diversity and biogeography in the late Palaeozoic and early Mesozoic |url=https://etheses.bham.ac.uk/id/eprint/9904/ |publisher=University of Birmingham |date=2020-07-01 |degree=d_ph |language=English |first=Emma M. |last=Dunne}}

|310 Ma

|Ohio, US

|A site known for its number of prehistoric tetrapods, like the lepospondyl Diceratosaurus.{{Cite journal |last1=Hook |first1=Robert W. |last2=Baird |first2=Donald |date=1986-06-19 |title=The Diamond Coal Mine of Linton, Ohio, and its Pennsylvanian-age vertebrates |url=http://www.tandfonline.com/doi/abs/10.1080/02724634.1986.10011609 |journal=Journal of Vertebrate Paleontology |language=en |volume=6 |issue=2 |pages=174–190 |doi=10.1080/02724634.1986.10011609 |bibcode=1986JVPal...6..174H |issn=0272-4634|url-access=subscription }}

|File:Diceratosaurus.jpg

Mazon Creek310 MaIllinois, US

|A conservation lagerstätte found near Morris, in Grundy County, Illinois. The fossils from this site are preserved in ironstone concretions with exceptional detail. The fossils were preserved in a large delta system that covered much of the area. The state fossil of Illinois, the enigmatic animal Tullimonstrum, is only known from these deposits.

|File:Tullimonstrum_gregarium_343.JPG

Buckhorn Asphalt Quarry

|~310 Ma

|Oklahoma, USA

|A quarry of the Boggy Formation known for its exceptionally rich orthocerid assemblage.{{Cite journal |last1=Niko |first1=Shuji |last2=Seuss |first2=Barbara |last3=Mapes |first3=Royal H. |date=1 January 2018 |title=Desmoinesian (Middle Pennsylvanian) Orthocerid Cephalopods from the Buckhorn Asphalt Lagerstätte in Oklahoma, Midcontinent North America |url=http://www.bioone.org/doi/10.2517/2017PR008 |journal=Paleontological Research |language=en |volume=22 |issue=1 |pages=20–36 |doi=10.2517/2017PR008 |issn=1342-8144 |access-date=12 May 2024 |via=BioOne Digital Library|url-access=subscription }}

|

Kinney Brick Quarry (Atrasado Formation)around 307 MaNew Mexico, US

|This site is known from rich fish fossils with preserved soft tissues, that lived in lagoonal environment. Dozens of fish genera are known, ranging from chondrichthyans like ctenacanths and hybodonts, to actinopterygians and sarcopterygians.{{Cite journal |last1=M. Hodnett |first1=John-Paul |last2=Lucas |first2=Spencer G. |title=Review Of The Late Pennsylvanian Fish Assemblage From The Kinney Brick Quarry, New Mexico |url=https://www.researchgate.net/publication/350890018 |journal=New Mexico Museum of Natural History and Science Bulletin |volume=84 |pages=359–390}}

|File:Dracopristis_hoffmanorum.png, a ctenacanth]]

Lontras Shale (Campáleo Outcrop){{Cite journal |last=Cardoso |first=Alexandre Ribeiro |last2=Schmidt |first2=Jaques Soares |last3=Bernardes |first3=Eduardo Müller |last4=Maraschin |first4=Anderson José |last5=de Coelho Andrade |first5=Débora |last6=de Medeiros Albano |first6=Filipe |last7=Rübensam |first7=Gabriel |last8=Barp |first8=Gustavo Bombardelli |last9=Kich |first9=Juliana Nichele |last10=Rodrigues |first10=Naira Poerner |last11=de Oliveira |first11=Yasmin Felix |last12=Zielinski |first12=João Pedro Tauscheck |last13=Weinschütz |first13=Luiz Carlos |last14=Vazquez |first14=Joselito Cabral |last15=Vecchia |first15=Felipe Dalla |date=2025-02-01 |title=Organic matter sources, accumulation and diagenesis of the Lontras Shale Lagerstätte (Paraná Basin, southern Brazil) |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018224006503 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=659 |pages=112661 |doi=10.1016/j.palaeo.2024.112661 |issn=0031-0182|url-access=subscription }}

|Gzhelian

|Santa Catarina, Brazil

|A fungal and palynological lagerstätte from Gondwana during Late Palaeozoic Ice Age.{{Cite journal |last1=Boardman |first1=Daiana Rockenbach |last2=Souza |first2=Paulo A. |last3=Scomazzon |first3=Ana Karina |last4=Félix |first4=Cristina Moreira |last5=Mori |first5=Ana Luisa Outa |last6=Weinschütz |first6=Luiz Carlos |date=January 2024 |title=Palynological analysis (biochronostratigraphy and paleoenvironments) from a fossil-Lagerstätte section within the Late Paleozoic Ice Age unit of the Paraná Basin (Itararé Group), Western Gondwana |url=https://linkinghub.elsevier.com/retrieve/pii/S0037073823002154 |journal=Sedimentary Geology |language=en |volume=459 |pages=106543 |doi=10.1016/j.sedgeo.2023.106543 |access-date=8 June 2024 |via=Elsevier Science Direct|url-access=subscription }}

|

Montceau-les-Mines300 MaFrance

|Exceptional preservation of Late Carboniferous fossil biota are known, including various vertebrates and arthropods, as well as plants.{{cite journal| last1=Perrier |first1=V. | last2=Charbonnier |first2=S. |year=2014 |title=The Montceau-les-Mines Lagerstätte (Late Carboniferous, France) |journal=Comptes Rendus Palevol |volume=13 | issue=5 |pages=353–67 | doi=10.1016/j.crpv.2014.03.002|bibcode=2014CRPal..13..353P }}{{Cite journal |last1=Heyler |first1=Daniel |last2=Poplin |first2=Cecile M. |date=1988 |title=The Fossils of Montceau-les-Mines |url=https://www.jstor.org/stable/24989233 |journal=Scientific American |volume=259 |issue=3 |pages=104–111 |doi=10.1038/scientificamerican0988-104 |jstor=24989233 |bibcode=1988SciAm.259c.104H |issn=0036-8733|url-access=subscription }}

|File:Idmonarachne_brasieri_reconstruction.jpg, an arachnid that is related to spiders]]

Hamilton Quarry300 MaKansas, US

|This site is known for its diverse assemblage of unusually well-preserved marine, euryhaline, freshwater, flying, and terrestrial fossils (invertebrates, vertebrates, and plants). This extraordinary mix of fossils suggests it was once an estuary.

|File:Spinoaequalis_schultzei_reconstruction.jpg, a semi-aquatic diapsid]]

Carrizo Arroyo? (Latest Gzhelian to earliest Asselian)New Mexico, US

|This site is known from exceptional preservation of arthropod fossils, mainly insects.{{Cite journal |title=NMGS 2016 Guidebook Details |url=https://nmgs.nmt.edu/publications/guidebooks/details.cfml?ID=131434 |access-date=2023-12-18 |website=New Mexico Geological Society |language=en |doi=10.56577/ffc-67.377|url-access=subscription }}

|

=Permian=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Taiyuan Formation298 MaInner Mongolia, China

|Known from exceptionally well-preserved plant fossils in volcanic ash.{{Cite journal |last1=Schmitz |first1=Mark D. |last2=Pfefferkorn |first2=Hermann W. |last3=Shen |first3=Shu-Zhong |last4=Wang |first4=Jun |date=2021-11-01 |title=A volcanic tuff near the Carboniferous–Permian boundary, Taiyuan Formation, North China: Radioisotopic dating and global correlation |url=https://linkinghub.elsevier.com/retrieve/pii/S0034666719303343 |journal=Review of Palaeobotany and Palynology |series=Wuda Tuff Flora: A Permian peat-forming T0 fossil plant assemblage from Wuda Coalfield, Inner Mongolia |volume=294 |pages=104244 |doi=10.1016/j.revpalbo.2020.104244 |issn=0034-6667|url-access=subscription }}{{Cite journal |last1=Wang |first1=Jun |last2=Hilton |first2=Jason |last3=Pfefferkorn |first3=Hermann W. |last4=Wang |first4=Shijun |last5=Zhang |first5=Yi |last6=Bek |first6=Jiri |last7=Pšenička |first7=Josef |last8=Seyfullah |first8=Leyla J. |last9=Dilcher |first9=David |date=2021-03-16 |title=Ancient noeggerathialean reveals the seed plant sister group diversified alongside the primary seed plant radiation |journal=Proceedings of the National Academy of Sciences |language=en |volume=118 |issue=11 |doi=10.1073/pnas.2013442118 |doi-access=free |issn=0027-8424 |pmc=7980368 |pmid=33836571}}

|

Meisenheim Formation{{Cite journal |last=Schoch |first=Rainer R. |date=2021-03-23 |title=Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships |url=https://fr.copernicus.org/articles/24/49/2021/ |journal=Fossil Record |language=English |volume=24 |issue=1 |pages=49–64 |doi=10.5194/fr-24-49-2021 |doi-access=free |issn=2193-0066}}?(Asselian to early Sakmarian){{Cite journal |last1=Poschmann |first1=Markus J. |last2=Nel |first2=André |last3=Raisch |first3=Manfred |date=2024-03-01 |title=Diversity and variability of grylloblattidan insects (Grylloblattida) from the early Permian Meisenheim Formation of the Saar-Nahe Basin (SW-Germany) |url=https://doi.org/10.1007/s12542-023-00672-8 |journal=PalZ |language=en |volume=98 |issue=1 |pages=67–84 |doi=10.1007/s12542-023-00672-8 |issn=1867-6812|url-access=subscription }}Lebach, Germany

|This site is well-known for the rich occurrence of fauna lived in large freshwater lakes, including fish, temnospondyls, insects and others.

|File:Orthacanthus Vienna.jpg, a large-sized xenacanth]]

Franchesse292 MaMassif Central, France

|A Sakmarian seymouriamorph lagerstätte from the Bourbon l'Archambault Basin in the French Massif Central containing hundreds of complete seymouriamorph specimens.{{cite journal |last1=Steyer |first1=J.-Sébastien |last2=Sanchez |first2=Sophie |last3=Debriette |first3=Pierre J. |last4=Valli |first4=Andrea M. F. |last5=Escuille |first5=François |last6=Pohl |first6=Burkhard |last7=Dechambre |first7=Roger-Paul |last8=Vacant |first8=Renaud |last9=Spence |first9=Christopher |last10=De Ploëg |first10=Gaël |date=1 December 2012 |title=A new vertebrate Lagerstätte from the Lower Permian of France (Franchesse, Massif Central): palaeoenvironmental implications for the Bourbon-l'Archambault basin |url=https://pubs.geoscienceworld.org/sgf/bsgf/article-abstract/183/6/509/314022/A-new-vertebrate-Lagerstatte-from-the-Lower |journal=Bulletin de la Société Géologique de France |volume=183 |issue=6 |pages=509–515 |doi=10.2113/gssgfbull.183.6.509 |access-date=29 April 2023|url-access=subscription }}

|

Chemnitz petrified forest291 MaSaxony, Germany

|A petrified forest in Germany that is composed of Arthropitys bistriata, a type of Calamites, giant horsetails that are ancestors of modern horsetails, found on this location with never seen multiple branches. Many more plants and animals from this excavation are still in an ongoing research.{{cite journal |last1=Luthardt |first1=Ludwig |last2=Rößler |first2=Ronny |last3=Schneider |first3=Joerg W. |date=1 January 2016 |title=Palaeoclimatic and site-specific conditions in the early Permian fossil forest of Chemnitz—Sedimentological, geochemical and palaeobotanical evidence |url=https://www.sciencedirect.com/science/article/pii/S0031018215005775 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=441 |issue= |pages=627–652 |doi=10.1016/j.palaeo.2015.10.015 |bibcode=2016PPP...441..627L |access-date=8 June 2022|url-access=subscription }}

|File:Bundesarchiv Bild 183-C1113-0001-001, Chemnitz, Versteinerte Bäume.jpg

Mangrullo Formationabout 285–275 Ma (Artinskian)Uruguay

|This site is known for its abundant mesosaur fossils. It also contains the oldest known konservat-lagerstätte in South America, as well as the oldest known fossils of amniote embryos.{{Cite journal | last1 = Piñeiro | first1 = G. | last2 = Ramos | first2 = A. | last3 = Goso | first3 = C. S. | last4 = Scarabino | first4 = F. | last5 = Laurin | first5 = M. | title = Unusual Environmental Conditions Preserve a Permian Mesosaur-Bearing Konservat-Lagerstätte from Uruguay | doi = 10.4202/app.2010.0113 | journal = Acta Palaeontologica Polonica | volume = 57 | issue = 2 | pages = 299–318 | year = 2012 | doi-access = free }}

|File:Stereosternum.jpg, an aquatic mesosaurid reptile]]

Chekarda (Koshelevka Formation)about 283–273 MaPerm, Russia

|Over 260 species of insect species are described from this site as well as diverse taxa of plants, making it one of the most important Permian konservat-lagerstätten.{{Cite journal |last1=Khramov |first1=Alexander V. |last2=Naugolnykh |first2=Sergey V. |last3=Węgierek |first3=Piotr |date=2022-09-12 |title=Possible long-proboscid insect pollinators from the Early Permian of Russia |journal=Current Biology |volume=32 |issue=17 |pages=3815–3820.e2 |doi=10.1016/j.cub.2022.06.085 |issn=0960-9822|doi-access=free |pmid=35858616 }}

|File:Tshekardocoleidae larva PIN-1700-4747 fig3A.png beetle larva]]

Toploje Member273-264 MaPrince Charles Mountains, Antarctica

|This site preserves a high-latitude fauna in exceptional position before the large extinctions that happened later in the Permian.{{cite journal |last1=Slater |first1=Ben J. |last2=McLoughlin |first2=Stephen |last3=Hilton |first3=Jason |date=June 2015 |title=A high-latitude Gondwanan lagerstätte: The Permian permineralised peat biota of the Prince Charles Mountains, Antarctica |journal=Gondwana Research |volume=27 |issue=4 |pages=1446–1473 |doi=10.1016/j.gr.2014.01.004 |bibcode=2015GondR..27.1446S |doi-access=free }}

|

Onder Karoo266.9–264.28 MaKaroo Basin, South Africa

|A high latitude, cool-temperate lacustrine ecosystem preserving detailed plant and insect fossils.{{cite journal |last1=Prevec |first1=Rosemary |last2=Nel |first2=André |last3=Day |first3=Michael O. |last4=Muir |first4=Robert A. |last5=Matiwane |first5=Aviwe |last6=Kirkaldy |first6=Abigail P. |last7=Moyo |first7=Sydney |last8=Staniczek |first8=Arnold |last9=Cariglino |first9=Bárbara |last10=Maseko |first10=Zolile |last11=Kom |first11=Nokuthula |last12=Rubidge |first12=Bruce S. |last13=Garrouste |first13=Romain |last14=Holland |first14=Alexandra |last15=Barber-James |first15=Helen M. |date=30 October 2022 |title=South African Lagerstätte reveals middle Permian Gondwanan lakeshore ecosystem in exquisite detail |journal=Communications Biology |volume=5 |issue=1 |page=1154 |doi=10.1038/s42003-022-04132-y |pmid=36310243 |pmc=9618562 }}

|

Sakamena Group{{Cite journal |date=1989-11-06 |title=Mass extinctions among tetrapods and the quality of the fossil record |url=https://royalsocietypublishing.org/doi/10.1098/rstb.1989.0094 |journal=Philosophical Transactions of the Royal Society of London. B, Biological Sciences |language=en |volume=325 |issue=1228 |pages=369–386 |doi=10.1098/rstb.1989.0094 |pmid=2574883 |issn=0080-4622 |last1=Benton |first1=M. J. |bibcode=1989RSPTB.325..369B |url-access=subscription }}260–247 MaMadagascar

|Lower Sakamena Formation (Permian) and Middle Sakamena Formation (Triassic) contain fossils of animals lived around wetland environment, such as semi-aquatic and gliding neodiapsids.

|File:Claudiosaurus_germaini_-_Redpath_Museum_-_McGill_University_-_Montreal,_Canada_-_DSC07787.jpg, aquatic neodiapsid reptile]]

Kupferschiefer259–255 MaCentral Europe

|This site deposited in an open marine and shallow marine environment provides fossils of reptiles as well as many fish.

|File:Weigeltisaurus_reconstruction.png, gliding weigeltisaurid reptile]]

Huopu Lagerstätte~255 MaGuizhou, China

|A plant fossil site documenting floral dynamics between the end-Guadalupian and end-Permian extinction events.{{cite journal |last1=Neregato |first1=R. |last2=D'Apolito |first2=C. |last3=Glasspool |first3=I. J. |last4=Wang |first4=S.-J. |last5=Liu |first5=F. |last6=Windslow |first6=P. |last7=Lu |first7=J. |last8=Shao |first8=L. |last9=Hilton |first9=J. |date=15 May 2016 |title=Palynological constraints on the provenance and stratigraphic range of a Lopingian (Late Permian) inter-extinction floral lagerstätte from the Xuanwei Formation, Guizhou Province, China |url=https://www.sciencedirect.com/science/article/abs/pii/S0166516216302762 |journal=International Journal of Coal Geology |volume=162 |pages=139–150 |doi=10.1016/j.coal.2016.06.005 |bibcode=2016IJCG..162..139N |s2cid=132822933 |access-date=18 June 2023|url-access=subscription }}

|

=Triassic=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Guiyang biota{{cite journal |last1=Dai |first1=Xu |last2=Davies |first2=Joshua H.F.L. |last3=Yuan |first3=Zhiwei |last4=Brayard |first4=Arnaud |last5=Ovtcharova |first5=Maria |last6=Xu |first6=Guanghui |last7=Liu |first7=Xiaokang |last8=Smith |first8=Christopher P.A. |last9=Schweitzer |first9=Carrie E. |last10=Li |first10=Mingtao |last11=Perrot |first11=Morgann G. |last12=Jiang |first12=Shouyi |last13=Miao |first13=Luyi |last14=Cao |first14=Yiran |last15=Yan |first15=Jia |last16=Bai |first16=Ruoyu |last17=Wang |first17=Fengyu |last18=Guo |first18=Wei |last19=Song |first19=Huyue |last20=Tian |first20=Li |last21=Dal Corso |first21=Jacopo |last22=Liu |first22=Yuting |last23=Chu |first23=Daoliang |last24=Song |first24=Haijun |year=2023 |title=A Mesozoic fossil lagerstätte from 250.8 million years ago shows a modern-type marine ecosystem |journal=Science |volume=379 |issue=6632 |pages=567–572 |doi=10.1126/science.adf1622|pmid=36758082 |bibcode=2023Sci...379..567D |s2cid=256697946|url=https://u-bourgogne.hal.science/hal-04016004/file/Dai%20et%20al.%20MS%20Version%20HAL.pdf }}250.8 MaGuizhou Province, China

|The oldest known Mesozoic lagerstätte (Dienerian). It preserves taxa belonging to 12 classes and 19 orders, including several species of fish.

|File:Teffichthys elegans.jpg elegans, a perleidiform ray-finned fish]]

Paris biota{{cite journal |last1=Brayard |first1=Arnaud |last2=Krumenacker |first2=L. J. |last3=Botting |first3=Joseph P. |last4=Jenks |first4=James F. |last5=Bylund |first5=Kevin G. |last6=Fara |first6=Emmanuel |last7=Vennin |first7=Emmanuelle |last8=Olivier |first8=Nicolas |last9=Goudemand |first9=Nicolas |last10=Saucède |first10=Thomas |last11=Charbonnier |first11=Sylvain |last12=Romano |first12=Carlo |last13=Doguzhaeva |first13=Larisa |last14=Thuy |first14=Ben |last15=Hautmann |first15=Michael |last16=Stephen |first16=Daniel A. |last17=Thomazo |first17=Christophe |last18=Escarguel |first18=Gilles |title=Unexpected Early Triassic marine ecosystem and the rise of the Modern evolutionary fauna |journal=Science Advances |year=2017 |volume=3 |issue=2 |pages=e1602159 |doi=10.1126/sciadv.1602159 |pmid=28246643 |pmc=5310825 |bibcode=2017SciA....3E2159B |doi-access=free}}~249 MaIdaho, Nevada, USA

|This earliest Spathian aged assemblage preserves fossils belonging to 7 phyla and 20 orders, combining Paleozoic groups (e.g. leptomitid protomonaxonid sponges otherwise known from the early Paleozoic) with members of the Modern evolutionary fauna (e.g. gladius-bearing coleoids).

|File:Ankitokazocaris NT.jpg triassica is a thylacocephalan from the Paris biota]]

Jialingjiang Formation{{Cite journal |last1=Flannery Sutherland |first1=Joseph T. |last2=Moon |first2=Benjamin C. |last3=Stubbs |first3=Thomas L. |last4=Benton |first4=Michael J. |date=2019-02-27 |title=Does exceptional preservation distort our view of disparity in the fossil record? |journal=Proceedings of the Royal Society B: Biological Sciences |language=en |volume=286 |issue=1897 |pages=20190091 |doi=10.1098/rspb.2019.0091 |issn=0962-8452 |pmc=6408902 |pmid=30963850}}249.2–247.2 MaHubei Province, China

|This site preserved aquatic reptiles soon after Permian extinction. Hupehsuchians are exclusively known from here, and already got unique ecology like filter feeding.

|File:Eretmorhipis_complete.jpg, a hupehsuchian that potentially had platypus-like ecology]]

Nanlinghu Formation248 MaAnhui Province, China

|This site provides important fossils to show early evolution of ichthyosauriforms.

|File:Cartorhynchus_NT.jpg, a primitive ichthyosauriform]]

Petropavlovka Formation248 MaOrenburg Oblast, Russia

|A site known for preserving oligochaetes, whose fossil record is extremely sparse.{{cite journal |last1=Shcherbakov |first1=Dmitry |last2=Tarmo |first2=Timm |last3=Tzetlin |first3=Alexander B. |last4=Vinn |first4=Olev |last5=Zhuravlev |first5=Andrey |date=April 2020 |title=A probable oligochaete from an Early Triassic Lagerstätte of the southern Cis-Urals and its evolutionary implications |url=https://www.researchgate.net/publication/340790069 |journal=Acta Palaeontologica Polonica |volume=65 |issue=2 |pages=219–233 |doi=10.4202/app.00704.2019 |s2cid=219097612 |access-date=28 April 2023|doi-access=free }}

Zarzaïtine Formation

|Olenekian-Anisian

|In Amenas, Algeria

|A site with a high number of exceptionally well-preserved temnospondyl specimens, indicating of a seasonal climate with sudden droughts, with a freshwater ecosystem that could rapidly turn into a sebkha.{{Cite journal |date=2016 |title=A new Mastodonsauroid Temnospondyl from the Triassic of Algeria: Implications for the biostratigraphy and palaeoenvironments of the Zarzaïtine Series, northern Sahara |url=https://sciencepress.mnhn.fr/fr/periodiques/comptes-rendus-palevol/15/fasc8/new-mastodonsauroid-temnospondyl-triassic-algeria-implications-biostratigraphy-and-palaeoenvironments-zarzaitine-series-northern-sahara |journal=Comptes Rendus Palevol |language=fr |volume=15 |issue=8 |pages=918–926 |doi=10.1016/j.crpv.2015.09.005|doi-access=free |last1=Dahoumane |first1=Anissa |last2=Nedjari |first2=Ahmed |last3=Aït-Ouali |first3=Rachid |last4=Taquet |first4=Philippe |last5=Vacant |first5=Renaud |last6=Steyer |first6=Jean-Sébastien }}

|

Luoping Biota (Guanling Formation){{Cite journal |last1=Hu |first1=Shi-xue |last2=Zhang |first2=Qi-yue |last3=Chen |first3=Zhong-Qiang |last4=Zhou |first4=Chang-yong |last5=Lü |first5=Tao |last6=Xie |first6=Tao |last7=Wen |first7=Wen |last8=Huang |first8=Jin-yuan |last9=Benton |first9=Michael J. |date=2011-08-07 |title=The Luoping biota: exceptional preservation, and new evidence on the Triassic recovery from end-Permian mass extinction |journal=Proceedings of the Royal Society B: Biological Sciences |language=en |volume=278 |issue=1716 |pages=2274–2282 |doi=10.1098/rspb.2010.2235 |issn=0962-8452 |pmc=3119007 |pmid=21183583}}

| ~247-245 Ma

Yunnan, China

|Various marine animals are preserved in this site, showing how marine ecosystem recovered after Permian extinction.{{Cite journal |last1=Liu |first1=Jun |last2=Martin Sander |first2=P. |date=2019-01-01 |title=The Vossenveld Formation and biotic recovery from the Permo-Triassic extinction |url=https://natuurtijdschriften.nl/pub/1021530/ |journal=Staringia |language=en |volume=16 |issue=1 |pages=147–152 |issn=0165-2354}}

|File:Barracudasauroides panxianensis MHNT Chine.jpg, an ichthyosaur preserved in articulation from the Guanling Formation]]

Hawkesbury Sandstone{{Cite journal |last1=Bicknell |first1=Russell D. C. |last2=Smith |first2=Patrick M. |date=2021-10-02 |title=The first fossil scorpion from Australia |url=https://www.tandfonline.com/doi/abs/10.1080/03115518.2021.1983874 |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=45 |issue=4 |pages=419–422 |doi=10.1080/03115518.2021.1983874 |issn=0311-5518|url-access=subscription }}

| Anisian

Sydney, Australia

|This sandstone produced many of freshwater fish fossils, as well as arthropods.

|File:Austrolimulus fletcheri AM F38275.png, a xiphosuran.]]

Grès à Voltzia245 MaFrance

|A fossil site remarkable for its detailed myriapod specimens.{{cite journal |last1=Shear |first1=W. A. |last2=Selden |first2=Paul A. |last3=Gall |first3=Jean-Claude |date=June 2009 |title=Millipedes from the Grès à Voltzia, Triassic of France, with comments on Mesozoic millipedes (Diplopoda: Helminthomorpha: Eugnatha) |url=https://www.researchgate.net/publication/233580187 |journal=International Journal of Myriapodology |volume=2 |issue=1 |pages=1–13 |doi=10.1163/187525409X462395 |doi-broken-date=13 April 2025 |access-date=18 June 2023|hdl=1808/8337 |hdl-access=free }} It also contains the earliest known aphid fossils.{{Cite journal |last1=Szwedo |first1=Jacek |last2=Nel |first2=André |date=1 December 2011 |title=The Oldest Aphid Insect from the Middle Triassic of the Vosges, France |journal=Acta Palaeontologica Polonica |language=en |volume=56 |issue=4 |pages=757–766 |doi=10.4202/app.2010.0034 |issn=0567-7920 |doi-access=free }}

Fossil Hill Member

|? (Anisian)

|Nevada, US

|One of many Anisian marine lagerstatte, the Fossil Hill Member represents an open-ocean environment with a well-preserved fauna largely dominated almost entirely by ichthyosaurs.

|File:Thalattoarchon saurophagis.jpg]]

Vossenveld Formation

|? (Anisian)

|Winterswijk,Netherlands

|An exposure of this Muschelkalk formation in the Winterswijk quarry has a diverse assemblage of well-preserved marine reptiles, amphibians, fishes, and plants. It is the only marginal marine assemblage recorded from the earlier Triassic.

|

Strelovec Formation

|? (Anisian)

|Slovenia

|A formation with well-preserved Triassic horseshoe crabs.{{Cite journal |last1=Bicknell |first1=Russell D. C. |last2=Zalohar |first2=Jure |last3=Miklavc |first3=Primoz |last4=Celarc |first4=Bogomir |last5=Kriznar |first5=Matija |last6=Hitij |first6=Tomaz |date=2021 |title=Revisiting horseshoe crab fossils from the Middle Triassic (Anisian) Strelovec Formation Konservat-Lagerstätte of Slovenia |url=https://rune.une.edu.au/web/handle/1959.11/37457 |journal=Palaeontologia Electronica |language=en |doi=10.26879/1168 |issn=1094-8074|doi-access=free }}

|

Saharonim Formation

|Late Anisian/Lower Ladinian

|Southern District, Israel

|One brachiopod-dominated horizon of this formation documents the rapid burial of a community of exclusively juvenile Coenothyris brachiopods and ten different bivalve genera.{{Cite journal |last=Feldman |first=Howard R. |date=2005 |title=Paleoecology, Taphonomy, and Biogeography of a Coenothyris Community (Brachiopoda, Terebratulida) from the Triassic (Upper Anisian–Lower Ladinian) of Israel |url=http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2005)479%5B0001%3APTABOA%5D2.0.CO%3B2 |journal=American Museum Novitates |language=en |issue=3479 |pages=1 |doi=10.1206/0003-0082(2005)479[0001:PTABOA]2.0.CO;2 |issn=0003-0082}}

|

Besano Formation242 MaAlps, Italy and Switzerland

|This formation is designated as a World Heritage Site, as it is famous for its preservation of Middle Triassic marine life including fish and aquatic reptiles.{{Cite journal |last1=Klug |first1=Christian |last2=Spiekman |first2=Stephan N. F. |last3=Bastiaans |first3=Dylan |last4=Scheffold |first4=Beat |last5=Scheyer |first5=Torsten M. |date=2024-03-04 |title=The marine conservation deposits of Monte San Giorgio (Switzerland, Italy): the prototype of Triassic black shale Lagerstätten |journal=Swiss Journal of Palaeontology |language=en |volume=143 |issue=1 |pages=11 |doi=10.1186/s13358-024-00308-7 |doi-access=free |issn=1664-2384 |pmc=10912274 |pmid=38450287}}

|File:Cymbospondylus buchseri skeleton 01.jpg, one of the many marine reptile specimens preserved in articulation from the Besano Formation]]

Pelsa-Vazzoler Lagerstätte

|Late Ladinian

|Northeastern Italy

|A site preserving a megadiverse mollusc fauna during the recovery from the Permian-Triassic extinction event.{{Cite journal |last1=Dominici |first1=Stefano |last2=Danise |first2=Silvia |last3=Tintori |first3=Andrea |date=15 August 2024 |title=A Middle Triassic Cassian-type fauna (Pelsa-Vazzoler Lagerstätte) and the adaptive radiation of the Modern evolutionary fauna |journal=Papers in Palaeontology |language=en |volume=10 |issue=4 |doi=10.1002/spp2.1579 |issn=2056-2799 |doi-access=free }}

|

Xingyi biota (Zhuganpo Formation)? (Upper Ladinian - Lower Carnian)Guizhou and Yunnan, China

|Previously considered as part of Falang Formation, this site yields many articulated skeletons of marine reptiles, as well as fish and invertebrates.

|File:Keichousaurus_hui_fossil.JPG, common sauropterygian from this site]]

Guanling biota (Xiaowa Formation)? (Carnian)Guizhou, China

|Like Xingi Biota, this site also yields well-preserved marine fauna, especially many species of thalattosaurs are known.

|File:Odontochelys semitestacea 433.jpg, an early pantestudine (relative of turtles)]]

Polzberg233 MaAustria

|A site from the Reingrabener Schiefer known for exceptional preservation of bromalites{{cite journal |last1=Lukeneder |first1=Alexander |last2=Surmik |first2=Dawid |last3=Gorzelak |first3=Przemysław |last4=Niedźwiedzki |first4=Robert |last5=Brachaniec |first5=Tomasz |last6=Salomon |first6=Mariusz A. |date=25 November 2020 |title=Bromalites from the Upper Triassic Polzberg section (Austria); insights into trophic interactions and food chains of the Polzberg palaeobiota |journal=Scientific Reports |volume=10 |issue=1 |page=20545 |doi=10.1038/s41598-020-77017-x |pmid=33239675 |pmc=7689505 }} and of cartilage,{{Cite journal|last1=Lukeneder |first1=P. |last2=Lukeneder |first2=A. |title=Mineralized belemnoid cephalic cartilage from the late Triassic Polzberg Konservat-Lagerstätte (Austria) |year=2022 |journal=PLOS ONE |volume=17 |issue=4 |pages=e0264595 |doi=10.1371/journal.pone.0264595 |pmid=35442996 |pmc=9020720 |bibcode=2022PLoSO..1764595L |doi-access=free }} deposited during the Carnian Pluvial Event.{{cite journal |last1=Lukeneder |first1=Alexander |last2=Lukeneder |first2=Petra |date=17 August 2021 |title=The Upper Triassic Polzberg palaeobiota from a marine Konservat-Lagerstätte deposited during the Carnian Pluvial Episode in Austria |journal=Scientific Reports |volume=11 |issue=1 |page=16644 |doi=10.1038/s41598-021-96052-w |pmid=34404880 |pmc=8370992 |bibcode=2021NatSR..1116644L }}

|File:Invertebrate members of the Polzberg palaeobiota.jpg

Madygen Formation230 MaKyrgyzstan

|The Madygen Formation is renowned for the preservation of more than 20,000 fossil insects, making it one of the richest Triassic lagerstätten in the world. Other vertebrate fossils as fish, amphibians, reptiles and synapsids have been recovered from the formation too, as well as minor fossil flora.

|File:Sharovipteryx mirabilis fossil.JPG of Sharovipteryx, a well-preserved gliding reptile]]

Cow Branch Formation230 MaVirginia, US

|This site preserves a wide variety of organisms (including Fish, reptiles, arachnids, and insects).

|File:Mecistotrachelos VMNH blog specimens.png, a gliding reptile distantly related to archosauromorphs, like crocodylians and dinosaurs]]

Alakir Çay

|? (Norian)

|southwest Turkey

|A konservat-lagerstätte with exceptionally well-preserved Triassic corals, retaining much of their original aragonite skeletons.{{Cite journal |last1=Frankowiak |first1=K. |last2=Mazur |first2=M. |last3=Gothmann |first3=A. M. |last4=Stolarski |first4=J. |date=2013-07-15 |title=Diagenetic Alteration of Triassic Coral from the Aragonite Konservat-Lagerstatte in Alakir Cay, Turkey: Implications for Geochemical Measurements |url=https://pubs.geoscienceworld.org/palaios/article/28/6/333-342/146328 |journal=PALAIOS |language=en |volume=28 |issue=6 |pages=333–342 |doi=10.2110/palo.2012.p12-116r |issn=0883-1351|url-access=subscription }}

|

=Jurassic=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Yuzhou Biota (Ziliujing Formation)

|~199 Ma

|Chongqing, China

|A fossil biota representing an extremely well-preserved lake ecosystem with freshwater mollusks, ray-finned fishes, lungfishes, sharks, and even pliosauroids, documenting one of the earliest well-preserved terrestrial ecosystems from Asia following the Triassic-Jurassic extinction event.{{Cite journal |last1=Ren |first1=Ting-Cong |last2=Ma |first2=Xin-Ying |last3=Wang |first3=Qing-Dong |last4=Xu |first4=Guang-Hui |date=2024-11-21 |title=An exceptionally preserved fossil assemblage from the early Jurassic of Chongqing (China) reveals a complex lacustrine ecosystem |journal=Scientific Reports |language=en |volume=14 |issue=1 |pages=26147 |doi=10.1038/s41598-024-77084-4 |pmid=39572595 |issn=2045-2322|pmc=11582640 }}

|File:Zigong Dinosaur Museum Bishanopliosaurus.jpg, a freshwater pliosaur]]

Osteno (Moltrasio Formation){{Cite journal |last1=Duffin |first1=Christopher J. |last2=Garassino |first2=Alessandro |last3=Pasini |first3=Giovanni |date=2023-05-19 |title=Squaloraja Riley 1833 (Holocephala: Squalorajidae) from the Lower Jurassic of Osteno Konservat-Lagerstätte (Como, NW Italy) |journal=Natural History Sciences |volume=10 |issue=1 |doi=10.4081/nhs.2023.642 |s2cid=258822179 |issn=2385-0922|doi-access=free }}~196-188 MaItaly

|Several kinds of marine biota such as fish, crustaceans, cephalopods, polychaetes, and nematodes have been recovered. This site is the only fossil deposit in Italy in which soft tissues are preserved other than Monte Bolca.

|File:Ostenocaris_vs_Ostenoteuthis_Osteno.png Ostenocaris and cephalopod Ostenoteuthis]]

Charmouth Mudstone Formation

|~196-188 Ma

|Lyme Regis and Charmouth (UK)

|Known for its amniote remains, specially complete specimens of the armoured dinosaur Scelidosaurus, but also plesiosaurs & ichthyosaurs.{{Cite journal |last=CONYBEARE |first=W. D. |date=1824 |title=XXI.—On the Discovery of an almost perfect Skeleton of the Plesiosaurus. |url=http://dx.doi.org/10.1144/transgslb.1.2.381 |journal=Transactions of the Geological Society of London |volume=1 |issue=2 |pages=381–389 |doi=10.1144/transgslb.1.2.381 |issn=2042-5295}}{{Cite journal |last1=Lomax |first1=Dean R. |last2=Massare |first2=Judy A. |date=2015-03-04 |title=A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K. |url=http://osf.io/89nhw |journal=Journal of Vertebrate Paleontology |language=en |volume=35 |issue=2 |pages=e903260 |bibcode=2015JVPal..35E3260L |doi=10.1080/02724634.2014.903260 |issn=0272-4634 |s2cid=85745787}}{{Cite journal |last=Norman |first=David B |date=2019-11-29 |title=Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: cranial anatomy |url=http://dx.doi.org/10.1093/zoolinnean/zlz074 |journal=Zoological Journal of the Linnean Society |volume=188 |issue=1 |pages=1–81 |doi=10.1093/zoolinnean/zlz074 |issn=0024-4082}} Large quantities of exceptional fish specimens and a diverse entomofauna are also known.{{Cite journal |last=Zeuner |first=F. E. |date=1962 |title=Fossil insects from the Lower Lias of Charmouth, Dorset |url=https://www.biodiversitylibrary.org/part/83511 |journal=Bulletin of the British Museum (Natural History), Geology |volume=7 |issue=1 |pages=155–171}}

|File:Scelidosaurus_harrisonii_(1).jpg]]

Rotzo Formation

|~192-184 Ma

|Italy

|Diverse fossils from a Carbonate archipelago, specially within plicatostylid reefs. It is mostly known for the exceptional preservation of organisms like Testate amoebae, and specially due to its amber-bearing layers and associated macroflora.{{Cite journal |last1=BASSI |first1=DAVIDE |last2=FUGAGNOLI |first2=ANNA |last3=POSENATO |first3=RENATO |last4=SCOTT |first4=DAVID B. |date=2008 |title=Testate Amoebae from the Early Jurassic of the Western Tethys, North-East Italy |journal=Palaeontology |volume=51 |issue=6 |pages=1335–1339 |bibcode=2008Palgy..51.1335B |doi=10.1111/j.1475-4983.2008.00817.x |issn=0031-0239 |s2cid=129670565 |doi-access=free}}{{cite journal |last1=Neri |first1=M. |last2=Roghi |first2=G. |last3=Ragazzi |first3=E. |last4=Papazzoni |first4=C. A. |date=2017 |title=First record of Pliensbachian (Lower Jurassic) amber and associated palynoflora from the Monti Lessini (northern Italy). |url=https://www.sciencedirect.com/science/article/pii/S0016699516301231 |journal=Geobios |volume=50 |issue=1 |pages=49–63 |bibcode=2017Geobi..50...49N |doi=10.1016/j.geobios.2016.10.001 |access-date=3 January 2022|url-access=subscription }}

|

Calcaires du Bou Dahar

|196-183 Ma

|Morocco

|It records the evolution of a carbonate platform.{{Cite journal |last1=MERINO-TOMÉ |first1=ÓSCAR |last2=PORTA |first2=GIOVANNA DELLA |last3=KENTER |first3=JEROEN A. M. |last4=VERWER |first4=KLAAS |last5=HARRIS |first5=PAUL (MITCH) |last6=ADAMS |first6=ERWIN W. |last7=PLAYTON |first7=TED |last8=CORROCHANO |first8=DIEGO |date=2011 |title=Sequence development in an isolated carbonate platform (Lower Jurassic, Djebel Bou Dahar, High Atlas, Morocco): influence of tectonics, eustacy and carbonate production |url=http://dx.doi.org/10.1111/j.1365-3091.2011.01232.x |journal=Sedimentology |volume=59 |issue=1 |pages=118–155 |doi=10.1111/j.1365-3091.2011.01232.x |issn=0037-0746|url-access=subscription }} It is known mostly for its rich coral bioherms and associated invertebrates.{{Cite journal |last1=Scheibner |first1=Christian |last2=Reijmer |first2=J. G. |date=1999 |title=Facies patterns within a Lower Jurassic upper slope to inner platform transect (Jbel Bou Dahar, Morocco) |url=http://dx.doi.org/10.1007/bf02537460 |journal=Facies |volume=41 |issue=1 |pages=55–80 |bibcode=1999Faci...41...55S |doi=10.1007/bf02537460 |issn=0172-9179|url-access=subscription }}{{Cite journal |last=Beauvais |first=L. |date=1986 |title=Monographie des Madreporaires du Jurassique Inferieur du Maroc (Monograph on Lower Jurassic Corals of Morocco) |url=https://www.schweizerbart.de/papers/pala/detail/A194/71204/Monographie_des_Madreporaires_du_Jurassique_inferieur_du_Maroc |journal=Palaeontographica Abteilung A |volume=194 |issue=3 |pages=1–68}}

|

Korsaröd (Djupadal Formation)

|183 Ma

|Skane, Sweden

|Know for it´s excepcionally preserved plant remains, specially Osmundastrum pulchellum, preserving intact calcified tissue with DNA and cells.{{cite journal |last1=Bomfleur |first1=B. |last2=McLoughlinVajda, V. (2014). Fossilized nuclei and chromosomes reveal 180 million years of genomic stasis in royal ferns. Science, 343(6177), 1376-1377 |first2=S. |last3=Vajda |first3=V. |date=2014 |title=Fossilized nuclei and chromosomes reveal 180 million years of genomic stasis in royal ferns |url=https://www.science.org/doi/full/10.1126/science.1249884 |journal=Science |volume=343 |issue=6177 |pages=1376–1377 |bibcode=2014Sci...343.1376B |doi=10.1126/science.1249884 |pmid=24653037 |s2cid=38248823 |access-date=30 July 2021}} It has also preserved its biotic interactions and even ongoing mitosis.{{cite journal |last1=McLoughlin |first1=S. |last2=Bomfleur |first2=B. |date=2016 |title=Biotic interactions in an exceptionally well preserved osmundaceous fern rhizome from the Early Jurassic of Sweden |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=464 |issue=1 |pages=86–96 |bibcode=2016PPP...464...86M |doi=10.1016/j.palaeo.2016.01.044 |doi-access=free}}{{cite journal |last1=Qu |first1=Y. |last2=McLoughlin |first2=M. |last3=van Zuilen |first3=M. A. |last4=Whitehouse |first4=M. |last5=Engdahl |first5=A. |last6=Vajda |first6=V. |date=2019 |title=Evidence for molecular structural variations in the cytoarchitectures of a Jurassic plant |journal=Geology |volume=47 |issue=4 |pages=325–329 |bibcode=2019Geo....47..325Q |doi=10.1130/g45725.1 |s2cid=84841297 |doi-access=free}}

|

Ya Ha Tinda183 MaAlberta, Canada

|A fossil site notable for containing abundant and extremely well-preserved vampire squid, being the largest concentration of vampire squid fossils outside the Tethys Ocean,{{cite journal |last1=Marroquín |first1=Selva M. |last2=Martindale |first2=Rowan C. |last3=Fuchs |first3=Dirk |date=7 February 2018 |title=New records of the late Pliensbachian to early Toarcian (Early Jurassic) gladius-bearing coleoid cephalopods from the Ya Ha Tinda Lagerstätte, Canada |url=https://onlinelibrary.wiley.com/doi/full/10.1002/spp2.1104 |journal=Papers in Palaeontology |volume=4 |issue=2 |pages=245–276 |doi=10.1002/spp2.1104 |s2cid=135120464 |access-date=8 April 2023|url-access=subscription }} and for being deposited during the Toarcian Oceanic Anoxic Event (TOAE).{{cite journal |last1=Martindale |first1=Rowan C. |last2=Aberhan |first2=Martin |date=15 July 2017 |title=Response of macrobenthic communities to the Toarcian Oceanic Anoxic Event in northeastern Panthalassa (Ya Ha Tinda, Alberta, Canada) |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018217300160 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=478 |pages=103–120 |doi=10.1016/j.palaeo.2017.01.009 |bibcode=2017PPP...478..103M |access-date=29 April 2023|url-access=subscription }}{{cite journal |last1=Martindale |first1=Rowan C. |last2=Them |first2=Theodore R. |last3=Gill |first3=Benjamin C. |last4=Marroquín |first4=Selva M. |last5=Knoll |first5=Andrew H. |date=1 July 2017 |title=A new Early Jurassic (ca. 183 Ma) fossil Lagerstätte from Ya Ha Tinda, Alberta, Canada |journal=Geology |volume=45 |issue=3 |pages=255–258 |doi=10.1130/G38808.1 |bibcode=2017Geo....45..255M |s2cid=56182151 |doi-access=free |hdl=10919/81874 |hdl-access=free }}{{cite journal |last1=Muscente |first1=A. D. |last2=Martindale |first2=Rowan C. |last3=Schiffbauer |first3=James D. |last4=Creighton |first4=Abby L. |last5=Bogan |first5=Brooke A. |date=4 November 2019 |title=Taphonomy of the Lower Jurassic Konservat-Lagerstätte at Ya Ha Tina (Alberta, Canada) and Its Significance for Exceptional Fossil Preservation During Oceanic Anoxic Events |url=https://pubs.geoscienceworld.org/sepm/palaios/article-abstract/34/11/515/574686/TAPHONOMY-OF-THE-LOWER-JURASSIC-KONSERVAT |journal=PALAIOS |volume=34 |issue=11 |pages=515–541 |doi=10.2110/palo.2019.050 |bibcode=2019Palai..34..515M |s2cid=208268686 |access-date=29 April 2023|url-access=subscription }}

|

Strawberry Bank183 MaSomerset, England

|A site from the TOAE documenting marine life during the recovery from the Triassic-Jurassic extinction event as well as the turmoil of the TOAE.{{cite journal |last1=Williams |first1=Matt |last2=Benton |first2=Michael James |last3=Ross |first3=Andrew |date=15 July 2015 |title=The Strawberry Bank Lagerstätte reveals insights into Early Jurassic life |url=https://pubs.geoscienceworld.org/jgs/article-abstract/172/6/683/144810/The-Strawberry-Bank-Lagerstatte-reveals-insights?redirectedFrom=fulltext |journal=Journal of the Geological Society |volume=172 |issue=6 |pages=683–692 |doi=10.1144/jgs2014-144 |bibcode=2015JGSoc.172..683W |hdl=1983/a7fc8bb4-889c-4fe9-b049-86d284141316 |s2cid=52971056 |access-date=18 June 2023|hdl-access=free }} The oldest pseudoplanktonic barnacles in the fossil record,{{cite journal |last1=Little |first1=Crispin T. S. |last2=Gale |first2=Andy |last3=Williams |first3=Matt |last4=Hammer |first4=Øyvind |last5=Fernandez |first5=Vincent |date=24 January 2023 |title=Bivalve-barnacle pseudoplanktonic colonisation of wood from the Toarcian, Lower Jurassic, Strawberry Bank Lagerstätte, Somerset, UK |url=https://www.app.pan.pl/article/item/app010182022.html |journal=Acta Palaeontologica Polonica |volume=68 |issue=1 |pages=133–142 |doi=10.4202/app.01018.2022 |s2cid=256258263 |access-date=18 June 2023|doi-access=free }} near-complete ichthyosaur skeletons,{{cite journal |last1=Srdic |first1=Alexander |last2=Beardmore |first2=Susan |last3=Lomax |first3=Dean R. |date=10 September 2019 |title=A rediscovered Lower Jurassic ichthyosaur skeleton possibly from the Strawberry Bank Lagerstätte, Somerset, UK |url=https://www.tandfonline.com/doi/abs/10.1080/08912963.2019.1663840 |journal=Historical Biology |volume=33 |issue=6 |pages=814–822 |doi=10.1080/08912963.2019.1663840 |s2cid=203358919 |access-date=18 June 2023}} and evidence of ichthyosaur niche partitioning are preserved at this site.{{cite journal |last1=Jamison-Todd |first1=Sarah |last2=Moon |first2=Benjamin C. |last3=Rowe |first3=Andre J. |last4=Williams |first4=Matt |last5=Benton |first5=Michael James |date=29 September 2022 |title=Dietary niche partitioning in Early Jurassic ichthyosaurs from Strawberry Bank |journal=Journal of Anatomy |volume=241 |issue=6 |pages=1409–1423 |doi=10.1111/joa.13744 |pmid=36175086 |pmc=9644957 }}

|

Grimmen Formation

|183 Ma

|Grimmen, Germany

|A coastal marine sequence, know specially for its fish fauna.{{cite journal |last1=Konwert |first1=M. |last2=Hörnig |first2=M. |date=2018 |title=Grimmenichthys ansorgei, gen. et sp. nov. (Teleostei, "Pholidophoriformes"), and other "pholidophoriform" fishes from the early Toarcian of Grimmen (Mecklenburg-Western Pomerania, Germany) |url=https://www.tandfonline.com/doi/abs/10.1080/02724634.2018.1451871 |journal=Journal of Vertebrate Paleontology |volume=38 |issue=3 |pages=1–16 |doi=10.1080/02724634.2018.1451872 |s2cid=90344418 |access-date=24 October 2021 |hdl-access=free |hdl=11336/84457}}{{cite journal |last1=Stumpf |first1=Sebastian |date=2017 |title=A synoptic review of the vertebrate fauna from the "Green Series"(Toarcian) of northeastern Germany with descriptions of new taxa: A contribution to the knowledge of Early Jurassic vertebrate palaeobiodiversity patterns |url=https://epub.ub.uni-greifswald.de/frontdoor/deliver/index/docId/1941/file/STUMPF2017DISS.pdf |journal=Greifswald University |type=PhD Thesis |pages=1–47 |access-date=9 September 2021}} The formation is also known for its rich entomofauna, including collections of up to 3000 specimens.{{cite journal |last1=Ansorge |first1=J. |date=2003 |title=Insects from the Lower Toarcian of Middle Europe and England |url=https://www.researchgate.net/publication/271444424 |journal=Proceedings of the Second Palaeoentomological Congress, Acta Zoologica Cracoviensia |volume=46 |issue=1 |pages=291–310 |access-date=30 July 2021}}

|File:Grimmen_fossil_invertebrates.png

Holzmaden/Posidonia Shale183 MaWürttemberg, Germany

|The Sachrang member is among the most important formations of the Toarcian boundary, due to the concentrations of exceptionally well-preserved complete skeletons of fossil marine fish and reptiles. It was also deposited during the TOAE.{{cite journal |last1=Ruebsam |first1=W. |last2=Schmid-Röhl |first2=A. |last3=Al-Husseini |first3=M. |title=Astronomical timescale for the early Toarcian (Early Jurassic) Posidonia Shale and global environmental changes |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |date=2023 |volume=623 |page=111619 |doi=10.1016/j.palaeo.2023.111619 |bibcode=2023PPP...62311619R |s2cid=258545235 |url=https://www.sciencedirect.com/science/article/pii/S0031018223002377 |access-date=10 July 2023|url-access=subscription }}{{cite journal |last1=Röhl |first1=Hans-Joachim |last2=Schmid-Röhl |first2=Annette |last3=Oschmann |first3=Wolfgang |last4=Frimmel |first4=Andreas |last5=Schwark |first5=Lorenz |date=1 January 2011 |title=The Posidonia Shale (Lower Toarcian) of SW-Germany: an oxygen-depleted ecosystem controlled by sea level and palaeoclimate |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018200001528 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=165 |issue=1–2 |pages=27–52 |doi=10.1016/S0031-0182(00)00152-8 |access-date=1 July 2023|url-access=subscription }}

|File:Ichtyosaurier Stenopterygius crassicostatus v. Huene mit 5 Embryonen 02.jpg Stenopterygius crassicostatus with preserved young]]

Cabeço da Ladeira

(Chão das Pias Formation)

|Late Bajocian

|Portugal

|A site known for exquisite preservation of microbial mats in a tidal flat.{{Cite journal |last1=Carvalho |first1=Carlos Neto de |last2=Pereira |first2=Bruno Claro |last3=Klompmaker |first3=Adiel |last4=Baucon |first4=Andrea |last5=Moita |first5=José António |last6=Pereira |first6=Pedro |last7=Machado |first7=Susana |last8=Belo |first8=João |last9=Carvalho |first9=Jorge M. F. |last10=Mergulhão |first10=Lia |date=3 May 2016 |title=Running crabs, walking crinoids, grazing gastropods: behavioral diversity and evolutionary implications in the Cabeço da Ladeira lagerstätte (Middle Jurassic, Portugal) |url=https://repositorioaberto.uab.pt/handle/10400.2/5285 |journal=Comunicações Geológicas |pages=39–54 |issn=0873-948X |access-date=8 June 2024}}

|File:Cabeço da Ladeira echinoderms.png

Monte Fallano? (Bajocian-Bathonian)Campania, Italy

|This Plattenkalk preserves fossils of terrestrial plants, crustaceans and fish.{{Cite journal |last1=Bravi |first1=Sergio |last2=Garassino |first2=Alessandro |last3=Bartiromo |first3=Antonello |last4=Audo |first4=Denis |last5=Charbonnier |first5=Sylvain |last6=Schweigert |first6=Günter |last7=Thévenard |first7=Frédéric |last8=Longobardi |first8=Cristiano |date=2014-04-01 |title=Middle Jurassic Monte Fallano Plattenkalk (Campania, southern Italy): first report on terrestrial plants, decapod crustaceans and fishes |url=https://www.schweizerbart.de/papers/njgpa/detail/272/82315/Middle_Jurassic_Monte_Fallano_Plattenkalk_Campania?af=crossref |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=272 |language=en |pages=79–107 |doi=10.1127/0077-7749/2014/0398|url-access=subscription }}

|

Christian Malford

|Callovian

|Wiltshire, England

|A site in the Oxford Clay Formation which preserves exceptionally detailed coleoid fossils.{{Cite journal |last1=Hart |first1=Malcolm B. |last2=De Jonghe |first2=Alex |last3=Page |first3=Kevin N. |last4=Price |first4=Gregory D. |last5=Smart |first5=Christopher W. |date=1 May 2016 |title=EXCEPTIONAL ACCUMULATIONS OF STATOLITHS IN ASSOCIATION WITH THE CHRISTIAN MALFORD LAGERSTÄTTE (CALLOVIAN, JURASSIC) IN WILTSHIRE, UNITED KINGDOM |url=https://pubs.geoscienceworld.org/palaios/article/31/5/203-220/325537 |journal=PALAIOS |language=en |volume=31 |issue=5 |pages=203–220 |doi=10.2110/palo.2015.066 |issn=0883-1351 |access-date=12 May 2024 |via=GeoScienceWorld|url-access=subscription }}

|

Mesa Chelonia{{citation

|journal=Naturwissenschaften

|title=An enormous Jurassic turtle bone bed from the Turpan Basin of Xinjiang, China

|first1=Oliver|last1= Wings

|first2=Márton|last2= Rabi

|first3=Jörg W.|last3= Schneider

|first4=Leonie|last4= Schwermann

|first5=Ge |last5=Sun

|first6=Chang-Fu|last6= Zhou

|first7=Walter G.|last7= Joyce

|volume=114|issue=11

|year=2012

|doi=10.1007/s00114-012-0974-5|pmid=23086389

|pages=925–35|bibcode=2012NW.....99..925W

|s2cid=17423081

}}

164.6 MaShanshan County, China

|This site is notable because it contains a large turtle bonebed, containing specimens of the genus Annemys. This bonebed contains up to an estimated 36 turtles per square meter.

|

La Voulte-sur-Rhône160 MyaArdèche, France

|La Voulte-sur-Rhône, in the Ardèche region of southwestern France, offers paleontologists an outstanding view of an undisturbed paleoecosystem that was preserved in fine detail. Notable finds include retinal structures in the eyes of thylacocephalan arthropods, and fossilized relatives of the modern day vampire squid, like Vampyronassa rhodanica.

|File:Rhomboteuthis lehmani.jpg]]

Shar Teeg Beds160-145 MyaGovi-Altay, Mongolia

|Many insect remains and some vertebrates like relatives of crocodilians are known from this site.{{Cite journal |last1=Ponomarenko |first1=A. G. |last2=Aristov |first2=D. S. |last3=Bashkuev |first3=A. S. |last4=Gubin |first4=Yu. M. |last5=Khramov |first5=A. V. |last6=Lukashevich |first6=E. D. |last7=Popov |first7=Yu. A. |last8=Pritykina |first8=L. N. |last9=Sinitsa |first9=S. M. |last10=Sinitshenkova |first10=N. D. |last11=Sukatsheva |first11=I. D. |last12=Vassilenko |first12=D. V. |last13=Yan |first13=E. V. |date=2014-12-01 |title=Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia |url=https://doi.org/10.1134/S0031030114140160 |journal=Paleontological Journal |language=en |volume=48 |issue=14 |pages=1573–1682 |doi=10.1134/S0031030114140160 |issn=1555-6174|url-access=subscription }}

|

Karabastau Formation155.7 MaKazakhstan

|This site is an important locality for insect fossils that has been studied since the early 20th century, alongside the rarer remains of vertebrates, including pterosaurs, salamanders, lizards and crocodiles.

|File:Sordes pilosus skeleton (cropped).JPG, small pterosaur with visible soft-tissues preserved]]

Tiaojishan Formation165-153 MaLiaoning Province, China

|It is known for its exceptionally preserved fossils, including those

of plants, insects and vertebrates. It is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks.

|File:Anchiornis-Beijing Museum of Natural History.jpg, small feathered anchiornithid dinosaur]]

La Casita Formation

|Kimmeridgian

|Coahuila, Mexico

|A marine konzentrat-lagerstätte deposited in a hemipelagic mud bottom during dysoxic conditions.{{Cite journal |last1=Zell |first1=Patrick |last2=Beckmann |first2=Seija |last3=Stinnesbeck |first3=Wolfgang |date=December 2014 |title=Age and depositional conditions of the marine vertebrate concentration Lagerstätte at Gomez Farías, southern Coahuila, Mexico |url=https://linkinghub.elsevier.com/retrieve/pii/S0895981114001114 |journal=Journal of South American Earth Sciences |language=en |volume=56 |pages=91–109 |doi=10.1016/j.jsames.2014.08.009 |access-date=8 June 2024 |via=Elsevier Science Direct|url-access=subscription }}

|

Talbragar fossil site{{Cite journal |last1=Frese |first1=Michael |last2=Gloy |first2=Gerda |last3=Oberprieler |first3=Rolf G. |last4=Gore |first4=Damian B. |date=2017-06-05 |title=Imaging of Jurassic fossils from the Talbragar Fish Bed using fluorescence, photoluminescence, and elemental and mineralogical mapping |journal=PLOS ONE |language=en |volume=12 |issue=6 |pages=e0179029 |doi=10.1371/journal.pone.0179029 |issn=1932-6203 |pmc=5459505 |pmid=28582427 |bibcode=2017PLoSO..1279029F |doi-access=free }}151 MaNew South Wales, Australia

|This bed is part of Purlawaugh Formation, and provided fauna like fish and insects that lived around the lake.

|File:Journal.pone.0179029.g007.tif, a teleost known from this site]]

Cleveland-Lloyd Dinosaur Quarry150 MaUtah, US

|Jurassic National Monument, at the site of the Cleveland-Lloyd Dinosaur Quarry, well known for containing the densest concentration of Jurassic dinosaur fossils ever found, is a paleontological site located near Cleveland, Utah, in the San Rafael Swell, a part of the geological layers known as the Morrison Formation. Up to 15,000 have been excavated from this site alone.

|File:Allosaurus Revised.jpg, the largest predator of the quarry]]

Canjuers Lagerstätte

|150 Ma

|France

|This site shows a high amount of biodiversity, including reptiles, invertebrates, fish, and other organisms.

|File:Canjuers_fossiles04.jpg, an aquatic rhynchocephalian]]

Agardhfjellet Formation

|150-140 Ma

|Spitsbergen, Norway

|The formation contains the Slottsmøya Member, a highly fossiliferous unit where many ichthyosaur and plesiosaur fossils have been found, as well as abundant and well preserved fossils of invertebrates.

|File:Pliosaurus_funkei21DB.jpg funkei, a large thalassophonean pliosaurid which was informally known as "Predator X"]]

Solnhofen Limestone149-148 MaBavaria, Germany

|This site is unique as it preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organisms such as sea jellies. The most familiar fossils of the Solnhofen Plattenkalk include the early feathered theropod dinosaur Archaeopteryx preserved in such detail that they are among the most famous and most beautiful fossils in the world.

|File:Archaeopteryx lithographica (Berlin specimen).jpg]]

Owadów–Brzezinki site

|~148 Ma

|Łódź Voivodeship, Poland

|A marine deposit of the Kcynia Formation similar to the Solnhofen Formation, with large numbers of preserved insect remains, numerous marine invertebrates, and vertebrates including fishes, marine reptiles, and pterosaurs.{{Cite journal |last1=Kin |first1=Adrian |last2=Gruszczyński |first2=Michał |last3=Martill |first3=David |last4=Marshall |first4=Jim D. |last5=Błażejowski |first5=Błażej |date=2013 |title=Palaeoenvironment and taphonomy of a Late Jurassic (Late Tithonian) Lagerstätte from central Poland |url=https://www.idunn.no/doi/10.1111/j.1502-3931.2012.00322.x |journal=Lethaia |language=en |volume=46 |issue=1 |pages=71–81 |doi=10.1111/j.1502-3931.2012.00322.x |issn=0024-1164|url-access=subscription }}{{Cite journal |last1=Błażejowski |first1=Błażej |last2=Weryński |first2=Łukasz |last3=Wierzbowski |first3=Andrzej |last4=Michalska |first4=Monika |last5=Hryniewicz |first5=Krzysztof |last6=Uchman |first6=Alfred |last7=Kugler |first7=Stanisław |last8=Bącal |first8=Paweł |last9=Hołda-Michalska |first9=Aleksandra |date=2023 |title=Summary of a decade of research at the Owadów–Brzezinki Lagerstätte (Tithonian, central Poland): A review and perspectives for the future |url=https://vjs.pgi.gov.pl/article/view/34082 |journal=Volumina Jurassica |language=en |volume=21 |pages=83–98 |issn=1731-3708}}{{Cite journal |last1=Weryński |first1=Łukasz |last2=Błażejowski |first2=Błażej |last3=Kędzierski |first3=Mariusz |date=2023 |title=A COMPARISON OF LATE JURASSIC PREDATORY ACTINOPTERYGII TEETH FROM OWADÓW-BRZEZINKI LÄGERSTATTE AND ITS PALAEOECOLOGICAL IMPLICATIONS |url=https://www.app.pan.pl/article/item/app010582023.html |journal=Acta Palaeontologica Polonica |volume=68 |doi=10.4202/app.01058.2023 |access-date=11 April 2025|doi-access=free }}

|File:Limulus darwini fossils.tif Crenatolimulus darwini]]

=Cretaceous=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Muzinho Shale

|latest Jurassic/earliest Cretaceous

|Parnaíba Basin, Brazil

|A black shale deposit containing articulated, three-dimensionally preserved fish skeletons.{{Cite journal |last1=Cardoso |first1=Alexandre Ribeiro |last2=Romero |first2=Guilherme Raffaeli |last3=Osés |first3=Gabriel Ladeira |last4=Nogueira |first4=Afonso César Rodrigues |date=2020-03-15 |title=Taphonomy of lacustrine fish fossils of the Parnaíba Basin, northeastern Brazil: Spatial and causative relations of Konservat Lagerstätten in West Gondwana during Jurassic-Cretaceous |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018219309009 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=542 |pages=109602 |doi=10.1016/j.palaeo.2020.109602 |issn=0031-0182|url-access=subscription }}

|

Angeac-Charente bonebed~141 MaCharente, France

|A lagerstätte preserving both vertebrate and invertebrate fossils from the poorly represented Berriasian stage known for its taphonomic and sedimentological ‘frozen scenes’.{{cite journal |last1=Rozada |first1=Lee |last2=Allain |first2=Ronan |last3=Vullo |first3=Romain |last4=Goedert |first4=Jean |last5=Augier |first5=Dominique |last6=Jean |first6=Amandine |last7=Marchal |first7=Jonathan |last8=Peyre de Fabrègues |first8=Claire |last9=Qvarnström |first9=Martin |last10=Royo-Torres |first10=Rafael |date=23 January 2021 |title=A Lower Cretaceous Lagerstätte from France: a taphonomic overview of the Angeac-Charente vertebrate assemblage |url=https://onlinelibrary.wiley.com/doi/abs/10.1111/let.12394 |journal=Lethaia |volume=54 |issue=2 |pages=141–165 |doi=10.1111/let.12394 |s2cid=234299378 |access-date=29 June 2023}}

|

El Montsec (La Pedrera de Rúbies Formation)~140-125 MaCatalonia, Spain

|Known from exceptional preservation of biota such as plants, fish, insects, crustaceans and even some tetrapods.{{Cite journal |last1=Gil-Delgado |first1=Alejandro |last2=Delclòs |first2=Xavier |last3=Sellés |first3=Albert |last4=Galobart |first4=Àngel |last5=Oms |first5=Oriol |date=2023-04-17 |title=The Early Cretaceous coastal lake Konservat-Lagerstätte of La Pedrera de Meià (Southern Pyrenees) |url=https://revistes.ub.edu/index.php/GEOACTA/article/view/41125 |journal=Geologica Acta |language=en |volume=21 |pages=1–XIII |doi=10.1344/GeologicaActa2023.21.3 |issn=1696-5728|hdl=2445/201820 |hdl-access=free }}

|File:Montsecosuchus.jpg, atoposaurid crocodylomorph]]

Lebanese amber

|~130-125 Ma (Barremian)

|Lebanon

|Preserves a high diversity of insects from the Early Cretaceous, and is among the oldest known fossilized amber to contain a significant number of preserved organisms.{{Cite journal |last1=Sánchez-García |first1=Alba |last2=Peñalver |first2=Enrique |last3=Delclòs |first3=Xavier |last4=Engel |first4=Michael S. |date=2019 |editor-last=Smith |editor-first=Andrew |title=Jumping bristletails (Insecta, Archaeognatha) from the Lower Cretaceous amber of Lebanon |url=https://onlinelibrary.wiley.com/doi/10.1002/spp2.1261 |journal=Papers in Palaeontology |language=en |volume=5 |issue=4 |pages=679–697 |doi=10.1002/spp2.1261 |hdl=2445/163385 |issn=2056-2799|hdl-access=free }} Includes many of the oldest known members of modern insect groups, and many of the youngest known members for extinct insect groups.{{Cite journal |last1=Sibelle |first1=Maksoud |last2=Dany |first2=Azar |date=2023 |title=Lebanese Amber: A Fantastic Journey into the Time of Dinosaurs |url=http://jogg.cug.edu.cn/en/article/doi/10.15964/j.cnki.027jgg.2023.04.012.pdf |journal=Journal of Gems & Gemmology |language=en |volume=25 |issue=4 |pages=136–145 |doi=10.15964/j.cnki.027jgg.2023.04.012 |issn=2096-9120}}

|File:Tragichrysa ovoruptora Lebanese amber fig1 a.jpg (Tragichrysa ovoruptora)]]

Grès du Liban Alloformation

|~125 Ma (early Barremian)

|Lebanon

|A deposit of dysodiles (sedimentary rocks with high organic matter content) representing a well-preserved freshwater ecosystem, including plants, pollen grains, invertebrates (including insects), turtles, and a diverse fish fauna.{{Cite journal |last1=El Hajj |first1=Layla |last2=Baudin |first2=François |last3=Gèze |first3=Raymond |last4=Cavin |first4=Lionel |last5=Dejax |first5=Jean |last6=Garcia |first6=Géraldine |last7=Horne |first7=David J. |last8=Maksoud |first8=Sibelle |last9=Otero |first9=Olga |last10=Azar |first10=Dany |date=2021-04-01 |title=Dysodiles from the lower Barremian of Lebanon: Insights on the fossil assemblages and the depositional environment reconstruction |url=https://www.sciencedirect.com/science/article/abs/pii/S0195667120304195 |journal=Cretaceous Research |volume=120 |pages=104732 |doi=10.1016/j.cretres.2020.104732 |issn=0195-6671}}{{Cite journal |last=Schädel |first=Mario |last2=Azar |first2=Dany |last3=El Hajj |first3=Layla |last4=Maksoud |first4=Sibelle |last5=Robin |first5=Ninon |date=2025-04-02 |title=A 125 million-year-old freshwater isopod shines new light on the origin of subterranean freshwater species |url=https://royalsocietypublishing.org/doi/full/10.1098/rsos.241512 |journal=Royal Society Open Science |volume=12 |issue=4 |pages=241512 |doi=10.1098/rsos.241512 |pmc=11962533 |pmid=40177107}}

|

Las Hoyasabout 125 Ma (Barremian)Cuenca, Spain

|The site is mostly known for its exquisitely preserved dinosaurs, especially enantiornithines. The lithology of the formation mostly consists of lacustarine limestone deposited in a freshwater wetland environment.

|File:Concornis lacustris 34.jpg, an early enantiornithean ]]

Yixian Formationabout 125–121 Ma (Barremian-Aptian)Liaoning, China

|The Yixian Formation is well known for its great diversity of well-preserved specimens and its feathered dinosaurs, such as the large tyrannosauroid Yutyrannus, the therizinosaur Beipiaosaurus, and various small birds, along with a selection of other dinosaurs, such as the iguanodontian Bolong, the sauropod Dongbeititan and the ceratopsian Psittacosaurus. Other biota included the troodontid Mei, the dromaeosaurid Tianyuraptor, and the compsognathid Sinosauropteryx.

|File:Sinosauropteryxfossil.jpg, the first non-avain dinosaur with evidence of feathers to have been recognized]]

Jiufotang Formationabout 122-119 Ma (Aptian)Liaoning, China

|This formation overlies the slightly older Yixian Formation and preserved very similar species, including a wide variety of dinosaurs such as the ceratopsian Psittacosaurus and the early bird Confuciusornis, both of which are also found in the Yixian Formation. Also notable are the very abundant specimens of the dromaeosaurid Microraptor, which is known from up to 300 specimens and is among the most common animals found here.

|File:Microraptor_gui_holotype.png independently of true birds.]]

Khasurty Fossil Site

|? (Aptian)

|Buryatia, Russia

|One of the largest fossil insect sites in northern Asia, with over 6000 fossilized insect specimens preserved in mudstones, representing over 16 orders and 130 families. Taxa have both Jurassic & Cretaceous affinities. Fossils of other invertebrates such as arachnids & crustaceans are also known, in addition to small plants and fragmentary vertebrate remains such as fish scales and bird feathers.{{Cite journal |last1=Kopylov |first1=D. S. |last2=Rasnitsyn |first2=A. P. |last3=Aristov |first3=D. S. |last4=Bashkuev |first4=A. S. |last5=Bazhenova |first5=N. V. |last6=Dmitriev |first6=V. Yu. |last7=Gorochov |first7=A. V. |last8=Ignatov |first8=M. S. |last9=Ivanov |first9=V. D. |last10=Khramov |first10=A. V. |last11=Legalov |first11=A. A. |last12=Lukashevich |first12=E. D. |last13=Mamontov |first13=Yu. S. |last14=Melnitsky |first14=S. I. |last15=Ogłaza |first15=B. |date=2020-12-01 |title=The Khasurty Fossil Insect Lagerstätte |url=https://doi.org/10.1134/S0031030120110027 |journal=Paleontological Journal |language=en |volume=54 |issue=11 |pages=1221–1394 |doi=10.1134/S0031030120110027 |issn=1555-6174|url-access=subscription }}

|

Shengjinkou Formationabout 120 MaXinjiang, China

|Part of the finds from this site consisted of dense concentrations of pterosaur bones, associated with soft tissues and eggs. The site represented a nesting colony that storm floods had covered with mud. Dozens of individuals could be secured from a total that in 2014 was estimated to run into the many hundreds.

|File:Hamititan skeletal.jpg, a large titanosaurian sauropod]]

Xiagou Formationabout 120–115? MaGansu, China

|This site is known outside the specialized world of Chinese geology as the site of a lagerstätte in which the fossils were preserved of Gansus yumenensis, the earliest true modern bird.

|File:Gansus zheni - early cretaceous Liaoning IMG 5188 Beijing Museum of Natural History.jpg is among the earliest known modern birds ]]

Paja Formation130-113 MaColombia

|This site is famous for its vertebrate fossils and is the richest Mesozoic fossiliferous formation of Colombia. Several marine reptile fossils of plesiosaurs, pliosaurs, ichthyosauras and turtles have been described from the formation and it hosts the only dinosaur fossils described in the country to date; the titanosauriform sauropod Padillasaurus.

|File:Desmatochelys padillai.jpg, a protostegid and one of the oldest known turtle fossils]]

Koonwarra Fossil Bed{{Cite journal |last1=Poropat |first1=Stephen F. |last2=Martin |first2=Sarah K. |last3=Tosolini |first3=Anne-Marie P. |last4=Wagstaff |first4=Barbara E. |last5=Bean |first5=Lynne B. |last6=Kear |first6=Benjamin P. |last7=Vickers-Rich |first7=Patricia |last8=Rich |first8=Thomas H. |date=2018-04-03 |title=Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date |url=https://www.tandfonline.com/doi/full/10.1080/03115518.2018.1453085 |journal=Alcheringa: An Australasian Journal of Palaeontology |language=en |volume=42 |issue=2 |pages=157–229 |doi=10.1080/03115518.2018.1453085 |bibcode=2018Alch...42..157P |s2cid=133845914 |issn=0311-5518}}around 118-115 MaVictoria, Australia

|This site is composed of mudstone sediment thought to have been laid down in a freshwater lake. Arthropods, fish and plant fossils are known from this site.

|File:Fossil-flea-292224-large.jpg, a large sized stem-flea]]

Crato Formation113 Manortheast Brazil

|The Crato Formation earns the designation of lagerstätte due to an exceedingly well preserved and diverse fossil faunal assemblage. Some 25 species of fossil fishes are often found with stomach contents preserved, enabling paleontologists to study predator-prey relationships in this ecosystem. There are also fine examples of pterosaurs, reptiles and amphibians, invertebrates (particularly insects), and plants. Also known from this site is Ubirajara, the first non-avian dinosaur from the southern hemisphere with evidence of feathers. Additionally, the formation abounds with evidence of plant-insect interaction.{{Cite journal |last1=Filho |first1=Edilson Bezerra dos Santos |last2=Adami-Rodrigues |first2=Karen |last3=Lima |first3=Flaviana Jorge de |last4=Bantim |first4=Renan Alfredo Machado |last5=Wappler |first5=Torsten |last6=Saraiva |first6=Antônio Álamo Feitosa |date=9 August 2019 |title=Evidence of plant–insect interaction in the Early Cretaceous Flora from the Crato Formation, Araripe Basin, Northeast Brazil |url=https://www.tandfonline.com/doi/full/10.1080/08912963.2017.1408611 |journal=Historical Biology |language=en |volume=31 |issue=7 |pages=926–937 |doi=10.1080/08912963.2017.1408611 |issn=0891-2963 |access-date=21 May 2024 |via=Taylor and Francis Online|url-access=subscription }}

|File:Tupandactylus navigans skeleton.PNG, a fossil pterosaur that was preserved with feathers and other soft tissues intact ]]

Amargosa Bed (Marizal Formation)? (Aptian-Albian)northeast Brazil

|Fluvial site which preserved fish, crustacean and plant fossils.{{Cite journal |last1=Voltani |first1=Cibele Gasparelo |last2=Osés |first2=Gabriel Ladeira |last3=Freitas |first3=Bernardo Tavares |last4=Prado |first4=Gustavo Marcondes Evangelista Martins |last5=Rohn |first5=Rosemarie |last6=Pacheco |first6=Mírian Liza Alves Forancelli |last7=Anelli |first7=Luiz Eduardo |last8=de Almeida |first8=Renato Paes |last9=Simões |first9=Marcello Guimarães |last10=Caldeira do Prado |first10=Ludmila Alves |last11=Araripe |first11=Rilda Verônica Cardoso de |last12=Galante |first12=Douglas |last13=Rangel |first13=Elidiane Cipriano |date=2023 |title=Taphonomy of fish, invertebrates and plant remains in the first Tethyan-South Atlantic marine ingression along Cretaceous rift systems in NE-Brazil |url=http://dx.doi.org/10.1016/j.cretres.2023.105508 |journal=Cretaceous Research |volume=147 |pages=105508 |doi=10.1016/j.cretres.2023.105508 |issn=0195-6671|url-access=subscription }}

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Pietraroja Plattenkalk

|113-110 Ma

|Campania, Italy

|A konservat-lagerstätte famous for its diverse and well-preserved fish and plant fossils. Also known from this formation is Scipionyx, one of Europe's most well-preserved dinosaurs.{{Cite web |title=Vertebrates from the uppermost stratigraphic sequence of the Pietraroja plattenkalk (Early Cretaceous, Southern Italy) |url=https://iris.unito.it/handle/2318/151109 |access-date=2024-07-12 |website=iris.unito.it}}{{Cite journal |last=Bartiromo |first=Antonello |date=2013-11-01 |title=Plant remains from the Lower Cretaceous Fossil-Lagerstätte of Pietraroja, Benevento, southern Italy |url=https://www.sciencedirect.com/science/article/pii/S0195667113001316 |journal=Cretaceous Research |volume=46 |pages=65–79 |doi=10.1016/j.cretres.2013.08.013 |issn=0195-6671|url-access=subscription }}

|File:Scipionyx samniticus.JPG, a small dinosaur with preserved internal organs.]]

Jinju Formation112.4–106.5 MaSouth Korea

|The Jinju Formation is notable for the post-Jehol Group insect assemblage,{{Cite journal |last1=Lee |first1=Soo Bin |last2=Li |first2=Yan-Da |last3=Cai |first3=Chenyang |last4=Engel |first4=Michael S. |last5=Nam |first5=Gi Soo |last6=Park |first6=Jong Kyun |last7=Nel |first7=André |last8=Jenkins Shaw |first8=Josh |last9=Jouault |first9=Corentin |last10=Legalov |first10=Andrei |last11=Kundrata |first11=Robin |date=June 2024 |title=Cretaceous beetles of the Jinju Formation (Coleoptera): An overview of the Jinju Formation, its coleopteran diversity, and past and future research |url=https://linkinghub.elsevier.com/retrieve/pii/S1226861524000414 |journal=Journal of Asia-Pacific Entomology |language=en |volume=27 |issue=2 |pages=102236 |doi=10.1016/j.aspen.2024.102236 |access-date=8 June 2024 |via=Elsevier Science Direct|url-access=subscription }} as well as other fauna such as isopods and fish.{{Cite journal |last1=Kim |first1=Haang-Mook |last2=Chang |first2=Mee-Mann |last3=Wu |first3=Feixiang |last4=Kim |first4=Yang-Hee |date=2014-01-01 |title=A new ichthyodectiform (Pisces, Teleostei) from the Lower Cretaceous of South Korea and its paleobiogeographic implication |url=https://www.sciencedirect.com/science/article/pii/S0195667113001778 |journal=Cretaceous Research |volume=47 |pages=117–130 |doi=10.1016/j.cretres.2013.11.007 |issn=0195-6671|url-access=subscription }}{{Cite journal |last1=Park |first1=Tae-Yoon |last2=Wilson |first2=George D. F. |last3=Lee |first3=Dong-Chan |last4=Choi |first4=Duck K. |date=2012 |title=Occurrence of the isopod Archaeoniscus coreaensis new species from the Lower Cretaceous Jinju Formation, Korea |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/occurrence-of-the-isopod-archaeoniscus-coreaensis-new-species-from-the-lower-cretaceous-jinju-formation-korea/50CB97E8DA651B48F396B3F88FB8A79A |journal=Journal of Paleontology |language=en |volume=86 |issue=4 |pages=626–640 |doi=10.1666/11-131R.1 |issn=0022-3360|url-access=subscription }} The site is also notable for its abundance and diversity of tetrapod trackways.{{Cite journal |last1=Kim |first1=Kyung Soo |last2=Lockley |first2=Martin G. |last3=Lim |first3=Jong Deock |last4=Kim |first4=Dong Hee |date=April 2019 |title=The oldest known anuran (frog) trackways from the Jinju Formation, Lower Cretaceous, Korea |url=https://linkinghub.elsevier.com/retrieve/pii/S0195667118304166 |journal=Cretaceous Research |language=en |volume=96 |pages=142–148 |doi=10.1016/j.cretres.2018.12.008 |access-date=8 June 2024 |via=Elsevier Science Direct|url-access=subscription }}

|File:Larva of Coptoclava.jpg beetle larva preserved at this locality]]

Tlayúa Formation110 MaPuebla, Mexico

|A marine lagerstätte preserving Albian actinopterygians and lepidosaurs.{{cite journal |last1=Than-Marchese |first1=Bruno Andrés |last2=Alvarado-Ortega |first2=Jesús |date=October 2022 |title=Armigatus felixi sp. nov. An Albian double armored herring (Clupeomorpha, Ellimmichthyiformes) from the Tlayúa lagerstätte, Mexico |url=https://www.sciencedirect.com/science/article/abs/pii/S0895981122001948 |journal=Journal of South American Earth Sciences |volume=118 |page=103905 |doi=10.1016/j.jsames.2022.103905 |bibcode=2022JSAES.11803905T |access-date=29 June 2023|url-access=subscription }}

|File:Cipactlichthys_scutatus_holotype.png, example of fish from this site]]

Romualdo Formation108–92 Manortheast Brazil

|The Romualdo Formation is a part of the Santana Group and has provided a rich assemblage of fossils; flora, fish, arthropods insects, turtles, snakes, dinosaurs, such as Irritator, and pterosaurs, including the genus Thalassodromeus. The stratigraphic units of the group contained several feathers of birds, among those the first record of Mesozoic birds in Brazil.

|File:Cast of Tapejara wellnhoferi - Pterosaurs Flight in the Age of Dinosaurs.jpg, a common pterosaur in Brazil during the Early Cretaceous ]]

Muhi Quarry (El Doctor Formation)? (Albian to Cenomanian, probably Late Albian){{Cite journal |last1=López-Palomino |first1=Isabel |last2=González-Rodríguez |first2=Katia Adriana |last3=Schultze |first3=Hans-Peter |last4=Palma-Ramírez |first4=Arturo |last5=Contreras-Cruz |first5=Diana |date=2021-11-01 |title=Ammonites from the La Negra Facies (El Doctor Formation, late Albian) of the Muhi Quarry, Hidalgo, central Mexico |url=https://www.sciencedirect.com/science/article/pii/S0895981121002479 |journal=Journal of South American Earth Sciences |volume=111 |pages=103400 |doi=10.1016/j.jsames.2021.103400 |issn=0895-9811|url-access=subscription }}Hidalgo, Mexico

|While this site produced limestones for construction, rocks in that locality contain a diverse Cretaceous marine biota such as fish, ammonites and crustaceans.

|File:Motlayoichthys_sergioi.jpg, a member of Pachyrhizodontidae]]

Puy-Puy Lagerstätte100.5 MaFrance

|A paralic site preserving a variety of ichnofossils,{{cite journal |last1=Néraudeau |first1=Didier |last2=Vullo |first2=Romain |last3=Bénéfice |first3=Pierre |last4=Breton |first4=Gérard |last5=Dépré |first5=Éric |last6=Gaspard |first6=Danièle |last7=Girard |first7=Vincent |last8=Le Couls |first8=Matthieu |last9=Moreau |first9=Jean-David |last10=Nel |first10=André |last11=Perrichot |first11=Vincent |last12=Solórzano-Kraemer |first12=Mónica M. |last13=Wappler |first13=Torsten |date=July 2020 |title=The paralic Albian–Cenomanian Puy-Puy Lagerstätte (Aquitaine Basin, France): An overview and new data |url=https://www.sciencedirect.com/science/article/abs/pii/S0195667118304038 |journal=Cretaceous Research |volume=111 |page=104124 |doi=10.1016/j.cretres.2019.03.022 |bibcode=2020CrRes.11104124N |s2cid=133649140 |access-date=18 June 2023}} along with some vertebrate remains.{{cite journal |last1=Vullo |first1=Romain |last2=Néraudeau |first2=Didier |last3=Dépré |first3=Eric |date=October 2013 |title=Vertebrate remains from the Cenomanian (Late Cretaceous) plant-bearing Lagerstätte of Puy-Puy (Charente-Maritime, France) |url=https://www.sciencedirect.com/science/article/abs/pii/S0195667113001043 |journal=Cretaceous Research |volume=45 |pages=314–320 |doi=10.1016/j.cretres.2013.06.002 |bibcode=2013CrRes..45..314V |access-date=18 June 2023|url-access=subscription }} The site preserves evidence of plant-insect interaction.{{cite journal |last1=Santos |first1=Artai A. |last2=Xiao |first2=Lifang |last3=Labandeira |first3=Conrad C. |last4=Néraudeau |first4=Didier |last5=Dépré |first5=Eric |last6=Moreau |first6=Jean-David |last7=Perrichot |first7=Vincent |last8=Wappler |first8=Torsten |date=1 July 2022 |title=Plant–insect interactions from the mid-Cretaceous at Puy-Puy (Aquitaine Basin, western France) indicates preferential herbivory for angiosperms amid a forest of ferns, gymnosperms, and angiosperms |url=https://www.tandfonline.com/doi/abs/10.1080/23818107.2022.2092772 |journal=Botany Letters |volume=169 |issue=4 |pages=568–587 |doi=10.1080/23818107.2022.2092772 |s2cid=250241945 |access-date=18 June 2023}}

Burmese amber101-99 Ma (latest Albian/earliest Cenomanian)Myanmar

|More than 1,000 species of taxa have been described from ambers from Hukawng Valley. While it is important for understanding the evolution of biota, mainly insects, during the Cretaceous period, it is also extremely controversial by facing ethical issues due to its association with conflicts and labor conditions.

|File:Oculudentavis_naga_specimen.jpg, small-sized lizard]]

El Chango Lagerstätte

|Earliest Cenomanian

|Chiapas

|A site from the beginning of the Angiosperm Terrestrial Revolution containing early evidence of insect predation on angiosperms.{{Cite journal |last1=Santos |first1=Artai A. |last2=McLoughlin |first2=Stephen |last3=Rubalcava-Knoth |first3=Marco A. |last4=Hernández-Damián |first4=Ana L. |last5=Villanueva-Amadoz |first5=Uxue |last6=Cevallos-Ferriz |first6=Sergio R. S. |date=10 July 2024 |title=Plant-insect interactions in the mid-Cretaceous paleotropical El Chango Lagerstätte (Cintalapa Fm., Mexico)—patterns of herbivory during the Angiosperm Terrestrial Revolution |journal=Frontiers in Ecology and Evolution |volume=12 |doi=10.3389/fevo.2024.1381539 |doi-access=free |issn=2296-701X }}

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English Chalk

|100-90 Ma (Cenomanian to Turonian)

|England

|Two subsections of England's famous chalk formation, the Grey Chalk Subgroup and the lower sections of the White Chalk Subgroup, yield three-dimensionally preserved fossils of marine fishes. This exquisite level of preservation is unlike fish fossils from other deposits from around the same time, which are only preserved as two-dimensional compression fossils.{{Cite journal |last1=Friedman |first1=Matt |last2=Beckett |first2=Hermione T. |last3=Close |first3=Roger A. |last4=Johanson |first4=Zerina |date=2016 |title=The English Chalk and London Clay: two remarkable British bony fish Lagerstätten |url=https://www.lyellcollection.org/doi/10.1144/SP430.18 |journal=Geological Society, London, Special Publications |language=en |volume=430 |issue=1 |pages=165–200 |doi=10.1144/SP430.18 |issn=0305-8719|url-access=subscription }}

|File:Aulolepis (36908826841).jpg]]

Haqel/Hjoula/al-Nammoura95-94 MaLebanon

|Famous Lebanese konservat-lagerstätten of the Late Cretaceous (middle to late Cenomanian) age, which contain a well-preserved variety of different fossils. Small animals like shrimp, octopus, stingrays, and bony fishes are common finds at these sites. Some of the rarest fossils from this locality include those of octopuses.{{Cite journal |last1=George |first1=Hady |last2=Bazzi |first2=Mohamad |last3=Hossny |first3=Tamara El |last4=Ashraf |first4=Nida |last5=Saad |first5=Pierre Abi |last6=Clements |first6=Thomas |date=2024-04-29 |title=The famous fish beds of Lebanon: the Upper Cretaceous Lagerstätten of Haqel, Hjoula, Nammoura, and Sahel Aalma |journal=Journal of the Geological Society |volume=181 |issue=5 |language=en |doi=10.1144/jgs2023-210 |issn=0016-7649|doi-access=free }}

|File:Ichthyoceros_fossil.jpg, a pycnodont actinopterygian]]

Ein Yabrud (Amminadav Formation or Bet-Meir Formation)CenomanianWest Bank, Palestine

|The marine fossil site with well-preserved marine vertebrate fossils, especially fossils of early legged snakes like Haasiophis and Pachyrhachis has quality of preservation rivals that of other famous Lagerstatten.{{Cite journal |last1=Rieppel |first1=Olivier |last2=Zaher |first2=Hussam |last3=Tchernov |first3=Eitan |last4=Polcyn |first4=Michael J. |date=2003 |title=The anatomy and relationships of Haasiophis terrasanctus, a fossil snake by well-developed hind limbs from the Mid-Cretaceous of the Middle East |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/anatomy-and-relationships-of-haasiophis-terrasanctus-a-fossil-snake-by-welldeveloped-hind-limbs-from-the-midcretaceous-of-the-middle-east/7241E4CCA42E98D8C3747A74146F1C1B |journal=Journal of Paleontology |language=en |volume=77 |issue=3 |pages=536–558 |doi=10.1666/0022-3360(2003)077<0536:TAAROH>2.0.CO;2 |issn=0022-3360|url-access=subscription }}

|File:Image_of_SMNK-PAL._8613.jpg, a pycnodont actinopterygian]]

Komen Limestone

|95-94 Ma

|Komen, Slovenia

|A Late Cenomanian locality in the Karst of Slovenia with a high diversity of articulated fossil fish, in addition to small reptiles and invertebrates.{{Cite journal |last1=Palci |first1=Alessandro |last2=Jurkovšek |first2=Bogdan |last3=Kolar-Jurkovšek |first3=Tea |last4=Caldwell |first4=Michael W. |date=2008 |title=New palaeoenvironmental model for the Komen (Slovenia) Cenomanian (Upper Cretaceous) fossil lagerstätte |url=https://doi.org/10.1016/j.cretres.2007.05.003 |journal=Cretaceous Research |volume=29 |issue=2 |pages=316–328 |doi=10.1016/j.cretres.2007.05.003 |issn=0195-6671|url-access=subscription }}

|File:Abhandlungen der Geologischen Bundesanstalt (1892) (16142481744).jpg]]

Hesseltal Formation

|94–93 Ma

|Saxony & North Rhine-Westphalia, Germany

|Deposited during the anoxic conditions of the Cenomanian-Turonian boundary event, this formation has a high number of well-preserved, articulated fish skeletons, in addition to exceptionally preserved ammonites with soft parts.{{Cite journal |last1=Cawley |first1=John |last2=Lehmann |first2=Jens |last3=Wiese |first3=Frank |last4=Kriwet |first4=Jürgen |date=2020 |title=Njoerdichthys dyckerhoffi gen. et sp. nov. (Pycnodontiformes, lower Turonian) northward migration caused by the Cretaceous Thermal Maximum |url=https://doi.org/10.1016/j.cretres.2020.104590 |journal=Cretaceous Research |volume=116 |pages=104590 |doi=10.1016/j.cretres.2020.104590 |issn=0195-6671 |pmc=7611863 |pmid=34690488}}{{Cite journal |last1=Klug |first1=Christian |last2=Riegraf |first2=Wolfgang |last3=Lehmann |first3=Jens |date=2012 |title=Soft–part preservation in heteromorph ammonites from the Cenomanian–Turonian Boundary Event (OAE 2) in north–west Germany |url=https://onlinelibrary.wiley.com/doi/10.1111/j.1475-4983.2012.01196.x |journal=Palaeontology |language=en |volume=55 |issue=6 |pages=1307–1331 |doi=10.1111/j.1475-4983.2012.01196.x |issn=0031-0239}}

|File:Rhamphoichthys flipped.jpg-like Rhamphoichthys]]

Vallecillo (Agua Nueva Formation)94–92 MaNuevo León, Mexico

|The site is noted for its qualities as a konservat-lagerstätte, with notable finds including the plesiosaur Mauriciosaurus and the possible shark Aquilolamna.

|File:Mauriciosaurus fernandezi.jpg, a short-necked plesiosaur ]]

Conulus Bed

|Turonian

|Poland

|A crinoid konzentrat-lagerstätte.{{Cite journal |last1=Salamon |first1=Mariusz A. |last2=Gorzelak |first2=Przemyslaw |last3=Borszcz |first3=Tomasz |last4=Gajerski |first4=Artur |last5=Kaźmierczak |first5=Jolanta |date=May–June 2009 |title=A crinoid concentration Lagerstätte in the Turonian (Late Cretaceous) Conulus Bed (Miechów-Wolbrom area, Poland) |url=https://www.sciencedirect.com/science/article/pii/S0016699509000229 |journal=Geobios |language=en |volume=42 |issue=3 |pages=351–357 |doi=10.1016/j.geobios.2008.10.008 |access-date=9 March 2025 |via=Elsevier Science Direct|url-access=subscription }}

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Akrabou Formation (Gara Sbaa/Agoult & Goulmima)? (Turonian)Asfla, Morocco

|Marine site known for exceptionally preserved, three-dimensional fish fossils.{{Cite journal |last1=Cooper |first1=Samuel L. A. |last2=Martill |first2=David M. |date=2020-12-01 |title=Pycnodont fishes (Actinopterygii, Pycnodontiformes) from the Upper Cretaceous (lower Turonian) Akrabou Formation of Asfla, Morocco |journal=Cretaceous Research |volume=116 |pages=104607 |doi=10.1016/j.cretres.2020.104607 |issn=0195-6671 |pmc=7442934 |pmid=32863512 |doi-access=free}}{{Cite journal |last1=Martill |first1=David M. |last2=Ibrahim |first2=Nizar |last3=Brito |first3=Paulo M. |last4=Baider |first4=Lahssen |last5=Zhouri |first5=Samir |last6=Loveridge |first6=Robert |last7=Naish |first7=Darren |last8=Hing |first8=Richard |date=2011 |title=A new Plattenkalk Konservat Lagerstätte in the Upper Cretaceous of Gara Sbaa, south-eastern Morocco |url=https://doi.org/10.1016/j.cretres.2011.01.005 |journal=Cretaceous Research |volume=32 |issue=4 |pages=433–446 |doi=10.1016/j.cretres.2011.01.005 |issn=0195-6671|url-access=subscription }}

|File:Grandemarinus.jpg, an early gar]]

Orapa diamond mine

|Turonian

|Botswana

|An insect lagerstätte known for being one of the few entomofaunas from southern Africa, containing a variety of insects,{{Cite journal |last1=Brothers |first1=D. J. |last2=Rasnitsyn |first2=A. P. |date=1 September 2003 |title=Diversity of Hymenoptera and other insects in the Late Cretaceous (Turonian) deposits at Orapa, Botswana : a preliminary review |url=https://journals.co.za/content/ento/11/2/EJC32559 |journal=African Entomology |language=en |volume=11 |issue=2 |pages=221–226 |issn=1021-3589 |access-date=8 June 2024 |via=Sabinet}} particularly beetles.{{Cite journal |last=Woolley |first=Christopher |date=6 June 2016 |title=The first scarabaeid beetle (Coleoptera, Scarabaeidae, Melolonthinae) described from the Mesozoic (Late-Cretaceous) of Africa |url=http://africaninvertebrates.pensoft.net/articles.php?id=8416 |journal=African Invertebrates |volume=57 |issue=1 |pages=53–66 |doi=10.3897/AfrInvertebr.57.8416 |doi-access=free |issn=2305-2562 |access-date=8 June 2024}}{{Cite journal |last1=Mnguni |first1=Sandiso |last2=McKay |first2=Ian James |last3=Badenhorst |first3=Shaw |date=29 April 2024 |title=A Fossil Paederinae from a Lacustrine Deposit at Orapa Diamond Mine in Botswana1 |url=https://meridian.allenpress.com/jes/article/doi/10.18474/JES23-99/500466/A-Fossil-Paederinae-from-a-Lacustrine-Deposit-at |journal=Journal of Entomological Science |volume=59 |issue=4 |language=en |doi=10.18474/JES23-99 |issn=0749-8004 |access-date=8 June 2024}}

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New Jersey amber

|91-89 Ma

|New Jersey, US

|Turonian-aged amber from the Raritan & Magothy Formations of New Jersey, with a high diversity of well-preserved insects, plants and fungi.{{Cite journal |last1=Grimaldi |first1=David |last2=Agosti |first2=Donat |date=2000-12-05 |title=A formicine in New Jersey Cretaceous amber (Hymenoptera: Formicidae) and early evolution of the ants |journal=Proceedings of the National Academy of Sciences |language=en |volume=97 |issue=25 |pages=13678–13683 |doi=10.1073/pnas.240452097 |doi-access=free |issn=0027-8424 |pmc=17635 |pmid=11078527}}

|File:Sphecomyrma freyi AMNH-NJ943A.jpg, an early ant]]

Lower Idzików beds

|87-86 Ma (Coniacian)

|Lower Silesian Voivodeship, Poland

|An exposure of these beds near Stary Waliszów contains a konzentrat-lagerstätte of numerous Cretaceous marine invertebrates in concretions, including decapods, molluscs and echinoderms, as well as well-preserved plant fossils that indicate a nearshore environment. Very well-preserved phosphatized decapod remains are known.{{Cite journal |last1=Trzęsiok |first1=Dawid |last2=Krzykawski |first2=Tomasz |last3=Niedźwiedzki |first3=Robert |last4=Brom |first4=Krzysztof |last5=Gorzelak |first5=Przemysław |last6=Salamon |first6=Mariusz A. |date=2014-05-01 |title=Palaeoenvironment of the Upper Cretaceous (Coniacian) concretion-bearing Lagerstätten from Poland |url=https://www.sciencedirect.com/science/article/pii/S0031018214001047 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=401 |pages=154–165 |doi=10.1016/j.palaeo.2014.02.030 |issn=0031-0182|url-access=subscription }}

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Smoky Hill Chalk87–82 MaKansas and Nebraska, US

|A Cretaceous konservat-lagerstätte known primarily for its exceptionally well-preserved marine reptiles. Also known from this site are fossils of large bony fish such as Xiphactinus, mosasaurs, flying reptiles or pterosaurs (namely Pteranodon), flightless marine birds such as Hesperornis, and turtles.

|File:Xiphactinus in Denver Museum.jpg is famous for being found with another fish (Gillicus) preserved in its stomach. ]]

Ingersoll Shale85 MaAlabama, US

|A Late Cretaceous (Santonian) informal geological unit in eastern Alabama. Fourteen theropod feathers assigned to birds and possibly dromaeosaurids have been recovered from the unit.

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Sahel Alma

|~84 Ma

|Lebanon

|A Late Cretaceous (Santonian) konservat-lagerstätte with similar excellent preservation of marine organisms as the nearby, older Sannine Lagerstätte, but in a deep-water environment. Includes a high number of well-preserved shark body fossils, in addition to cephalopods and deepwater arthropods.

|File:Pristiophorus tumidens Sahel Alma Santonian.jpg, an early sawshark]]

Calcare di Aurisina

|~80-70.6 Ma

|Italy & Slovenia

|A late cretaceous shallow marine series of carbonate platforms dominated by rudists, with fossils of invertebrates and vertebrates, specially fishes.{{Cite journal |last1=Dalla Vecchia |first1=F.M |last2=Tentor |first2=M. |date=2004 |title=Il Carso 85 milioni di anni fa: Gli straordinari fossili di Polazzo |url=https://www.academia.edu/download/51467252/Il_Carso_85_milioni_di_anni_fa_gli_strao20170122-22471-vnp25m.pdf |journal=Gruppo Speleologico Monfalconese A.d.F |pages=1–75}} Its best known outcrop is the Villaggio del Pescatore site, that yielded the holotype of Tethyshadros, as well other exceptionally preserved taxa like Acynodon adriaticus.{{Cite journal |last1=Chiarenza |first1=Alfio Alessandro |last2=Fabbri |first2=Matteo |last3=Consorti |first3=Lorenzo |last4=Muscioni |first4=Marco |last5=Evans |first5=David C. |last6=Cantalapiedra |first6=Juan L. |last7=Fanti |first7=Federico |date=2021-12-02 |title=An Italian dinosaur Lagerstätte reveals the tempo and mode of hadrosauriform body size evolution |journal=Scientific Reports |language=en |volume=11 |issue=1 |pages=23295 |bibcode=2021NatSR..1123295C |doi=10.1038/s41598-021-02490-x |issn=2045-2322 |pmc=8640049 |pmid=34857789}}{{Cite journal |last1=Marco |first1=Muscioni |last2=Chiarenza |first2=Alfio Alessandro |last3=Delfino |first3=Massimo |last4=Fabbri |first4=Matteo |last5=Milocco |first5=Kevin |last6=Fanti |first6=Federico |date=2023 |title=Acynodon adriaticus from Villaggio del Pescatore (Campanian of Italy): Anatomical and chronostratigraphic integration improves phylogenetic resolution in Hylaeochampsidae (Eusuchia) |url=http://dx.doi.org/10.1016/j.cretres.2023.105631 |journal=Cretaceous Research |volume=151 |pages=105631 |bibcode=2023CrRes.15105631M |doi=10.1016/j.cretres.2023.105631 |issn=0195-6671 |hdl-access=free |hdl=11093/5418}}

|File:Tethyshadros_insularis.JPG]]

Auca Mahuevo80 MaPatagonia, Argentina

|A Cretaceous lagerstätte in the eroded badlands of the Patagonian province of Neuquén, Argentina. The sedimentary layers of the Anacleto Formation at Auca Mahuevo were deposited between 83.5 and 79.5 million years before the present and offers a view of a fossilized titanosaurid nesting site.

|File:Sauropod egg Auca Mahuevo Museum of Anchient Life.jpg

Ellisdale Fossil Site

|79-76 Ma

|New Jersey, US

|A middle Campanian konzentrat-lagerstätte from the Marshalltown Formation with one of the most diverse Mesozoic vertebrate faunas of eastern North America, likely originating from a flood event. A high number of disarticulated bones of dinosaurs, fish, reptiles, amphibians, and small mammals is known, most of which are microfossils.{{Cite journal |last=Brownstein |first=Chase Doran |date=2019-04-01 |title=First Record of a Small Juvenile Giant Crocodyliform and its Ontogenetic and Biogeographic Implications |url=https://bioone.org/journals/bulletin-of-the-peabody-museum-of-natural-history/volume-60/issue-1/014.060.0104/First-Record-of-a-Small-Juvenile-Giant-Crocodyliform-and-its/10.3374/014.060.0104.full |journal=Bulletin of the Peabody Museum of Natural History |volume=60 |issue=1 |pages=81 |doi=10.3374/014.060.0104 |issn=0079-032X|url-access=subscription }}

|

Coon Creek Formation

|76.8-76.0 Ma{{Cite journal |last1=Self-Trail |first1=Jean M. |last2=Gardner |first2=Kristina F. |last3=O'Keefe |first3=Jennifer |last4=Mason |first4=Patricia H. |last5=Puckett |first5=Mark |last6=Gibson |first6=Michael A. |last7=McCarty |first7=M. Maeve |date=2024 |title=Microfossils and biostratigraphy of the Upper Cretaceous Coon Creek Formation Lagerstätte, Mississippi Embayment, USA |url=https://www.lyellcollection.org/doi/10.1144/SP545-2023-137 |journal=Geological Society, London, Special Publications |language=en |volume=545 |issue=1 |pages=717–738 |doi=10.1144/SP545-2023-137 |issn=0305-8719|url-access=subscription }}

|Tennessee and Mississippi, US

|This late Campanian formation has some of the world's best-preserved remains of Cretaceous marine invertebrates (primarily mollusks and decapod crustaceans), with many retaining their original aragonitic shells and exoskeletons.{{Cite journal |last1=Vrazo |first1=M. B. |last2=Diefendorf |first2=A. F. |last3=Crowley |first3=B. E. |last4=Czaja |first4=A. D. |date=2018 |title=Late Cretaceous marine arthropods relied on terrestrial organic matter as a food source: Geochemical evidence from the Coon Creek Lagerstätte in the Mississippi Embayment |url=https://onlinelibrary.wiley.com/doi/10.1111/gbi.12270 |journal=Geobiology |language=en |volume=16 |issue=2 |pages=160–178 |doi=10.1111/gbi.12270 |issn=1472-4677 |pmid=29350453}}

|File:Avitelmessus grapsoides.JPG Avitelmessus grapsoides, which occurs in great numbers at Coon Creek]]

Baumberge Formation

|~75-72 Ma

|North Rhine-Westphalia, Germany

|A Late Campanian formation in the Baumberge of Germany with a high number of articulated fossil fish remains, in addition to shark body fossils.{{Cite web |title=PBDB |url=https://paleobiodb.org/classic/displaySearchStrataResults?group_formation_member=Baumberge |access-date=2024-07-12 |website=paleobiodb.org}}

|File:Sphenocephalus fissicaudus 349.jpg fish Sphenocephalus]]

Nardò (Calcari di Melissano){{Cite journal |last1=Paparella |first1=Ilaria |last2=Palci |first2=Alessandro |last3=Nicosia |first3=Umberto |last4=Caldwell |first4=Michael W. |date=2018 |title=A new fossil marine lizard with soft tissues from the Late Cretaceous of southern Italy |journal=Royal Society Open Science |volume=5 |issue=6 |pages=172411 |doi=10.1098/rsos.172411 |issn=2054-5703|doi-access=free |pmid=30110414 |pmc=6030324 }}~72-70 Ma (upper Campanian-lower Maastrichtian)Apulia, Italy

|This site is especially famous for its limestones containing abundant fossil fish remains.

|File:Nardovelifer_altipinnis.jpg, the earliest lampriform]]

Harrana (Muwaqqar Chalk Marl Formation)66.5-66.1 Ma (Late Maastrichtian){{Cite journal |last1=Jagt |first1=John W.M. |last2=Jagt-Yazykova |first2=Elena A. |last3=Kaddumi |first3=Hani F. |last4=Lindgren |first4=Johan |date=2018-10-02 |title=Ammonite dating of latest Cretaceous mosasaurid reptiles (Squamata, Mosasauroidea) from Jordan—preliminary observations |url=https://www.tandfonline.com/doi/full/10.1080/03115518.2017.1308011 |journal=Alcheringa: An Australasian Journal of Palaeontology |language=en |volume=42 |issue=4 |pages=587–596 |doi=10.1080/03115518.2017.1308011 |issn=0311-5518|url-access=subscription }}Jordan

| Phosphatic deposits formed in this site are known to preserve vertebrate fossils with soft tissue, such as mosasaurs, plesiosaurs, sharks, bony fish, turtles and crocodylians.{{Cite journal |last1=Lindgren |first1=Johan |last2=Kaddumi |first2=Hani F. |last3=Polcyn |first3=Michael J. |date=2013-09-10 |title=Soft tissue preservation in a fossil marine lizard with a bilobed tail fin |journal=Nature Communications |language=en |volume=4 |issue=1 |pages=2423 |doi=10.1038/ncomms3423 |issn=2041-1723|doi-access=free |pmid=24022259 }}

|File:Postredectes_harranaensis.jpg, an example of fish fossil from this site]]

Tanis{{Cite journal |title=A seismically induced onshore surge deposit at the KPg boundary, North Dakota |last=DePalma |first=Robert |display-authors=etal |date=2 April 2019 |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=116 |issue=17 |pages=8190–8199 |doi=10.1073/pnas.1817407116 |pmid=30936306 |pmc=6486721 |bibcode=2019PNAS..116.8190D |doi-access=free}}

|66.0 Ma

|North Dakota, US

|Tanis is part of the heavily studied Hell Creek Formation, a group of rocks spanning four states in North America renowned for many significant fossil discoveries from the Upper Cretaceous and lower Paleocene. Tanis is a significant site because it appears to record the events from the first minutes until a few hours after the impact of the giant Chicxulub asteroid in extreme detail. This impact, which struck the Gulf of Mexico 66.043 million years ago, wiped out all non-avian dinosaurs and many other species (the so-called "K-Pg" or "K-T" extinction).

|File:Goddard Scientific Colloquium, Tanis fish fossils.jpg from the Tanis site]]

=Paleogene=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Baunekule Facies

|64-63 Ma (Danian)

|eastern Denmark

|These facies of the Faxe Formation document an extremely well-preserved cold-water coral mound ecosystem dominated by Dendrophyllia corals, and also includes gastropods, tubeworms, bivalves, bryozoans and gastropods.{{Cite journal |last1=Lauridsen |first1=B. W. |last2=Bjerager |first2=M. |date=2021-12-15 |title=The fauna of a Danian fossil Conservation Lagerstätten from the cold-water coral mound complex at Faxe, Denmark |url=https://www.sciencedirect.com/science/article/pii/S0031018221004855 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=584 |pages=110700 |doi=10.1016/j.palaeo.2021.110700 |issn=0031-0182|url-access=subscription }}

|

Tenejapa-Lacandón Formation

|~63 Ma

|Chiapas, Mexico

|A formation with a high number of well-preserved fish fossils indicative of mass mortality events.{{Cite journal |last1=Cantalice |first1=Kleyton Magno |last2=Martínez-Melo |first2=Alejandra |last3=Romero-Mayén |first3=Violeta Amparo |date=2019-08-30 |title=The paleoichthyofauna housed in the Colección Nacional de Paleontología of Universidad Nacional Autónoma de México |url=https://zse.pensoft.net/article/35435/ |journal=Zoosystematics and Evolution |language=en |volume=95 |issue=2 |pages=429–452 |doi=10.3897/zse.95.35435 |doi-access=free |issn=1860-0743}} One of the most important formations for documenting the recovery of ocean ecosystems in the wake of the K-Pg extinction, due to being deposited just a few million years after and being located only {{Convert|500|km|mi}} away from the Chixculub impact site.{{Cite journal |last1=Alvarado-Ortega |first1=J |last2=Cuevas-García |first2=M |last3=Melgarejo-Damián |first3=M |last4=Cantalice |first4=KS. |last5=Alaniz-Galvan |first5=A |last6=Solano-Templos |first6=G |last7=Than-Marchese |first7=B |date=2015 |title=Paleocene fishes from Palenque, Chiapas, southeastern Mexico |url=http://dx.doi.org/10.26879/536 |journal=Palaeontologia Electronica |doi=10.26879/536 |issn=1094-8074}}

|File:Paleoichthyofauna (10.3897-zse.95.35435) Figure 4.jpg, Kelmejtubus, and Paleoserranus, three unique fish]]

Menat Formation

|

60 Ma

|

Auvergne, France

|

A Palaeocene maar lake containing three-dimensional plant remains.{{cite journal |last1=Wedmann |first1=Sonja |last2=Uhl |first2=Dieter |last3=Lehmann |first3=Thomas |last4=Garrouste |first4=Romain |last5=Nel |first5=Andre |last6=Gomez |first6=Bernard |last7=Smith |first7=Krister T. |last8=Schaal |first8=S. F. K. |date=June 2018 |title=The Konservat-Lagerstätte Menat (Paleocene; France) – an overview and new insights |url=https://www.researchgate.net/publication/326081432 |journal=Geologica Acta |volume=16 |issue=2 |pages=189–213 |doi=10.1344/GeologicaActa2018.16.2.5 |access-date=29 April 2023}} It is particularly notable for preserving one of the oldest known bees.{{Cite journal |last1=Michez |first1=Denis |last2=De Meulemeester |first2=Thibaut |last3=Rasmont |first3=Pierre |last4=Nel |first4=André |last5=Patiny |first5=Sébastien |date=18 February 2009 |title=New fossil evidence of the early diversification of bees: Paleohabropoda oudardi from the French Paleocene (Hymenoptera, Apidae, Anthophorini) |url=https://onlinelibrary.wiley.com/doi/10.1111/j.1463-6409.2008.00362.x |journal=Zoologica Scripta |language=en |volume=38 |issue=2 |pages=171–181 |doi=10.1111/j.1463-6409.2008.00362.x |issn=0300-3256 |access-date=7 July 2024 |via=Wiley Online Library|url-access=subscription }}

|File:Paleohabropoda oudardi MNHN.F.R63891 direct lighting.jpg.]]

Danata Formation

|56-53 Ma

|western Turkmenistan

|Outcropping in the Kopet Dag range, this formation preserves numerous fossil fish from a northeastern arm of the Tethys Ocean during the Paleocene-Eocene thermal maximum. 38 taxa from 13 orders are known, the vast majority of which are acanthomorphs.{{Cite journal |last1=Bannikov |first1=A. F. |last2=Erebakan |first2=I. G. |date=2023 |title=On the Evolution of Some Groups of Marine Bony Fishes in the Cenozoic of the Tethys and Paratethys |url=https://link.springer.com/10.1134/S0031030123050015 |journal=Paleontological Journal |language=en |volume=57 |issue=5 |pages=475–490 |doi=10.1134/S0031030123050015 |issn=0031-0301|url-access=subscription }}

|File:Eospinus daniltshenkoi.jpg, a bizarre tetraodontiform fish.]]

Fur Formation

|

55–53 Ma

|

Fur & Mors, Denmark

|

Preserves abundant fossil fish, insects, reptiles, birds and plants. The Fur Formation was deposited about 55 Ma, just after the Palaeocene-Eocene boundary, and its tropical or sub-tropical flora indicate that the climate after the Paleocene-Eocene Thermal Maximum was moderately warm (approximately 4-8 degrees warmer than today).

|

File:Tasbacka danica.JPG, immature sea turtle]]

London Clay

|

54–48 Ma

|

England, UK

|

Collected for close to 300 years, Plant fossils, especially seeds and fruits, are found in abundance.
Some 350 named species of plant have been found, making the London Clay flora one of the world's most diverse for fossil seeds and fruits. The flora includes tropical taxa found in modern Asia, reflecting the much warmer climate of the early Eocene.

|File:Brychaetus muelleri skull and anterior body.jpg, fish skull]]

Eocene Okanagan Highlands

|

52 - 48 Ma

|

British Columbia, Canada & Washington, USA

|

Includes McAbee Fossil Beds, Princeton chert & Klondike Mountain Formation; Recognized as temperate/subtropical uplands right after the Paleocene–Eocene Thermal Maximum and spanning the Early Eocene Climatic Optimum, preserves highly detailed uplands lacustrine fauna and flora.

|

File:Florissantia quilchenensis 01 SRIC.jpg flower, an extinct malvaceous genus from western North America]]

Monte Solane

|51-49 MA

|Verona, Italy

|Slightly older than the nearby, more well-known Monte Bolca site, the Monte Solane site also preserves numerous marine fish and plants, but documents an entirely different ecosystem that appears to be of a bathypelagic habitat, forming one of the few known lagerstätte to preserve a deep-sea ecosystem.{{Cite journal |last1=Giusberti |first1=Luca |last2=Bannikov |first2=Alexander |last3=Boscolo Galazzo |first3=Flavia |last4=Fornaciari |first4=Eliana |last5=Frieling |first5=Joost |last6=Luciani |first6=Valeria |last7=Papazzoni |first7=Cesare Andrea |last8=Roghi |first8=Guido |last9=Schouten |first9=Stefan |last10=Sluijs |first10=Appy |last11=Bosellini |first11=Francesca R. |last12=Zorzin |first12=Roberto |date=2014-06-01 |title=A new Fossil-Lagerstätte from the Lower Eocene of Lessini Mountains (northern Italy): A multidisciplinary approach |url=https://www.sciencedirect.com/science/article/pii/S0031018214001199 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=403 |pages=1–15 |doi=10.1016/j.palaeo.2014.03.012 |hdl=1874/309076 |issn=0031-0182|hdl-access=free }}

|

Green River Formation

|

50 Ma

|

Colorado/Utah/Wyoming, US

|

An Eocene aged site that is noted for the fish fauna preserved. Other fossils include the crocodilians, birds, and mammals.

|

File:Diplomystus and Knightia Green River Fm WY.jpg and Knightia]]

Monte Bolca

|

50-49 Ma

|

Verona, Italy

|

A fossil site with specimens of fish and other organisms that are so highly preserved that their organs are often completely intact in fossil form, and even the skin color can sometimes be determined. It is assumed that mud at the site was low in oxygen, preventing both decay and the mixing action of scavengers from harming the fossils.{{Cite journal |last1=Friedman |first1=Matt |last2=Carnevale |first2=Giorgio |date=2018 |title=The Bolca Lagerstätten: shallow marine life in the Eocene |url=https://www.lyellcollection.org/doi/10.1144/jgs2017-164 |journal=Journal of the Geological Society |language=en |volume=175 |issue=4 |pages=569–579 |doi=10.1144/jgs2017-164 |issn=0016-7649|url-access=subscription }}

|

File:Naturkundemuseum Berlin - Archaeophis proavus Massalongo - Monte Bolca.jpg]]

Messel Formation

|

47 Ma

|

Hessen, Germany

|

This site has significant geological and scientific importance. Over 1000 species of plants and animals have been found at the site. After almost becoming a landfill, strong local resistance eventually stopped these plans and the Messel Pit was declared a UNESCO World Heritage Site on 9 December 1995. Significant scientific discoveries about the early evolution of mammals and birds are still being made at the Messel Pit, and the site has increasingly become a tourist site as well.

|

File:Masillamys Senckenberg 2007-01.JPG sp., an ischyromyid rodent]]

Baltic amber

|47-35 Ma (Lutetian to Priabonian)

|Pomeranian Voivodeship, Poland & Kaliningrad Oblast, Russia

|The largest amber deposit on Earth, this amber is part of the Prussian Formation, and preserves a high diversity of exceptionally well-preserved fossil invertebrates, plants, and small vertebrates that inhabited eastern Europe during the warmer, subtropical conditions of the middle Eocene. It is the largest world's single largest repository of fossil insects.{{cite journal |last1=Perkovsky |first1=E. E. |last2=Rasnitsyn |first2=A. P. |last3=Vlaskin |first3=A. P. |last4=Taraschuk |first4=M. V. |year=2007 |title=A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples |journal=African Invertebrates |volume=48 |issue=1 |pages=229–245}}{{Citation |last1=Rikkinen |first1=Jouko |title=Morphological Convergence in Forest Microfungi Provides a Proxy for Paleogene Forest Structure |date=2018 |work=Transformative Paleobotany |pages=527–549 |url=https://doi.org/10.1016/B978-0-12-813012-4.00022-X |access-date=2024-07-12 |publisher=Elsevier |doi=10.1016/b978-0-12-813012-4.00022-x |isbn=978-0-12-813012-4 |last2=Schmidt |first2=Alexander R.|url-access=subscription }}{{Citation |last1=Heinrichs |first1=Jochen |title=A Comprehensive Assessment of the Fossil Record of Liverworts in Amber |date=2018 |work=Transformative Paleobotany |pages=213–252 |url=https://doi.org/10.1016/B978-0-12-813012-4.00012-7 |access-date=2024-07-12 |publisher=Elsevier |doi=10.1016/b978-0-12-813012-4.00012-7 |isbn=978-0-12-813012-4 |last2=Feldberg |first2=Kathrin |last3=Bechteler |first3=Julia |last4=Regalado |first4=Ledis |last5=Renner |first5=Matthew A.M. |last6=Schäfer-Verwimp |first6=Alfons |last7=Gröhn |first7=Carsten |last8=Müller |first8=Patrick |last9=Schneider |first9=Harald|url-access=subscription }}{{Cite journal |last1=Wolfe |first1=Alexander P. |last2=McKellar |first2=Ryan C. |last3=Tappert |first3=Ralf |last4=Sodhi |first4=Rana N.S. |last5=Muehlenbachs |first5=Karlis |date=2016 |title=Bitterfeld amber is not Baltic amber: Three geochemical tests and further constraints on the botanical affinities of succinite |url=https://doi.org/10.1016/j.revpalbo.2015.11.002 |journal=Review of Palaeobotany and Palynology |volume=225 |pages=21–32 |doi=10.1016/j.revpalbo.2015.11.002 |issn=0034-6667|url-access=subscription }}

|File:Deutsches Bernsteinmuseum 04 (cropped).jpg, a gecko]]

Kishenehn Formation

|46.2 Ma

|Montana, US

|A Middle Eocene site preserving exquisitely detailed insect specimens in oil shale.{{cite journal |last1=Greenwalt |first1=Dale E. |last2=Rose |first2=Tim R. |last3=Siljestrom |first3=Sandra M. |last4=Goreva |first4=Julia S. |last5=Constenius |first5=Kurt N. |last6=Wingerath |first6=Jonathan G. |date=24 June 2014 |title=Taphonomy of the Fossil Insects of the Middle Eocene Kishenehn Formation |journal=Acta Palaeontologica Polonica |volume=60 |issue=4 |pages=931–947 |doi=10.4202/app.00071.2014 |s2cid=53364401 |doi-access=free }}

|

Mahenge Formation

|46 Ma

|Tanzania

|A terrestrial Middle Eocene lagerstätte preserving fish, plant and arthropod fossils.{{cite journal |last1=Herendeen |first1=Patrick S. |last2=Jacobs |first2=Bonnie F. |date=September 2000 |title=Fossil Legumes from the Middle Eocene (46.0 Ma) Mahenge Flora of Singida, Tanzania |url=https://www.jstor.org/stable/2656727 |journal=American Journal of Botany |volume=87 |issue=9 |pages=1358–1366 |doi=10.2307/2656727 |jstor=2656727 |pmid=10991905 |access-date=29 June 2023|url-access=subscription }}{{cite journal |last1=Schlüter |first1=Thomas |date=2018 |title=Eocene insects from a Maar Lagerstätte at Mahenge, northern Tanzania |url=https://web.s.ebscohost.com/abstract?direct=true&profile=ehost&scope=site&authtype=crawler&jrnl=01718177&AN=131864569&h=vgyyeqtF3nqPl6DDjZPzEZDaiMzmmlGhBHslRi4WBMg0%2bZZfueAEmB4PBBKiKWkKtPlPxqxwLYpWzjwMRwYdDw%3d%3d&crl=c&resultNs=AdminWebAuth&resultLocal=ErrCrlNotAuth&crlhashurl=login.aspx%3fdirect%3dtrue%26profile%3dehost%26scope%3dsite%26authtype%3dcrawler%26jrnl%3d01718177%26AN%3d131864569 |journal=Entomologia Generalis |volume=37 |issue=3–4 |pages=375–392 |doi=10.1127/entomologia/2017/0653 |s2cid=91844938 |access-date=29 June 2023|url-access=subscription }}

|File:Singedella NT.jpg]]

Quercy Phosphorites Formation{{cite journal| last1=Lalloy |first1=F. | last2=Rage|first2=J. C. | last3=Evans | first3=S.E. | last4=Boistel | first4=R. | last5=Lenoir | first5=N. | last6=Laurin | first6=M. |year=2013 |title=A re-interpretation of the Eocene anuran Thaumastosaurus based on microCT examination of a 'mummified' specimen |journal=PLOS ONE |volume=8 |issue=9 |pages=1–11 | doi = 10.1371/journal.pone.0074874 | pmid=24086389 | pmc=3783478|bibcode=2013PLoSO...874874L |doi-access=free }}

|

45-25 Ma

|

Occitania, France

|

This site qualifies as a lagerstätte because beside a large variety of mammals, birds, turtles, crocodiles, flora and insects, it also preserves the soft tissues of amphibians and squamates, in addition to their articulated skeleton in what has been called natural mummies.

|

File:Xenomorphia resurrecta fossil tomography.png parasitic wasps preserved in fly pupae]]

Na Duong

|Priabonian

|Northern Vietnam

|A Late Eocene site notable for its highly detailed coprolite preservation.{{Cite journal |last1=Halaçlar |first1=Kazım |last2=Rummy |first2=Paul |last3=Liu |first3=Jia |last4=Hunt |first4=Adrian P. |last5=Van Do |first5=Truong |last6=Minh |first6=Nguyen Trung |last7=Deng |first7=Tao |date=15 September 2023 |title=Exceptionally well-preserved crocodilian coprolites from the Late Eocene of Northern Vietnam: Ichnology and paleoecological significance |journal=iScience |language=en |volume=26 |issue=9 |pages=107607 |doi=10.1016/j.isci.2023.107607 |pmc=10470398 |pmid=37664591 }}

|

Bitterfeld amber

|38-34 Ma

|Saxony, Germany

|One of three major Paleogene deposits of European amber, this deposit of the Cottbus Formation shares a similar biota to the Baltic Amber, indicating a concurrent formation, but appears to have a geologically distinct origin from Baltic amber.

|File:Bitterfeld amber pseudoscorpions (10.3897-evolsyst.2.22581) Figure 4.jpgs from the Bitterfeld amber]]

Rovno amber

|38-34 Ma

|Rivne Oblast, Ukraine & Gomel, Belarus

|One of three major Paleogene deposits of European amber, this deposit of the Obukhov Formation preserves a high diversity of invertebrates, many of which are shared with the Baltic amber but others of which are unique. A drier habitat compared to the Baltic amber is suggested based on some of the insect taxa preserved.

|File:Eocenomyrma ukrainica CASENT0917550 holotype profile.jpg, an ant]]

Florissant Formation

|

34 Ma

|

Colorado, US

|

A late Eocene (Priabonian) aged site that is noted for the finely preserved plant and insect paleobiota. Fossils are preserved in diatom blooms of a lahar dammed lake system and the formation is noted for the petrified stumps of Sequoia affinis

|

File:Palaeovespa florissantia.jpg ]]

Babaheidar (Pabdeh Formation)

|late Eocene/early Oligocene

|Chaharmahal and Bakhtiari province, Iran

|A well-preserved marine fossil site in the Zagros Mountains with thousands of known fish fossils, as well as plants, crustaceans, insects, and birds.{{Cite journal |last1=Mirzaie Ataabadi |first1=Majid |last2=Bahrami |first2=Ali |last3=Yazdi |first3=Mehdi |last4=Nel |first4=André |date=2019-05-28 |title=A locust witness of a trans-oceanic Oligocene migration between Arabia and Iran (Orthoptera: Acrididae) |url=https://www.tandfonline.com/doi/full/10.1080/08912963.2017.1378651 |journal=Historical Biology |language=en |volume=31 |issue=5 |pages=574–580 |doi=10.1080/08912963.2017.1378651 |issn=0891-2963}}

|File:Iraniplectus bakhtiari.jpg, an unusual tetraodontiform]]

Tremembé Formation

|30-25 Ma

|São Paulo, Brazil

|A Late Oligocene-aged freshwater lake deposit, containing well-preserved fossil fishes.{{Cite journal |last=Gasparelo Voltani |first=Cibele |last2=Menegazzo |first2=Mirian Costa |last3=Galeazzi |first3=Cristiano Padalino |last4=Montefeltro |first4=Felipe Chinaglia |last5=Almeida |first5=Renato Paes de |last6=Anelli |first6=Luiz Eduardo |date=2023-09-01 |title=Fish biostratinomy applied to lacustrine paleoenvironmental reconstructions: Oligocene deposits of Taubaté Basin, Brazil |url=https://linkinghub.elsevier.com/retrieve/pii/S0895981123003413 |journal=Journal of South American Earth Sciences |volume=129 |pages=104530 |doi=10.1016/j.jsames.2023.104530 |issn=0895-9811|url-access=subscription }}

|File:Lignobrycon ligniticus (P9012).tif Lignobrycon ligniticus]]

Menilite Formation

|Early Oligocene

|Poland

|A deep-sea flysch deposit along the Carpathian Mountains, originally deposited in the Paratethys Sea with exceptionally preserved deepwater fish fossils and microbial mats.{{Cite journal |last=Leonowicz |first=Paulina |last2=Bienkowska-Wasiluk |first2=Małgorzata |last3=Ochmanski |first3=Tomasz |date=2021-05-15 |title=Benthic microbial mats from deep-marine flysch deposits (Oligocene Menilite Formation from S Poland): Palaeoenvironmental controls on the MISS types |url=https://www.sciencedirect.com/science/article/abs/pii/S0037073821000336 |journal=Sedimentary Geology |volume=417 |pages=105881 |doi=10.1016/j.sedgeo.2021.105881 |issn=0037-0738|url-access=subscription }}

|File:Serranus-budensis 2.jpg{{Cite journal |last=Bienkowska-Wasiluk |first=M. |last2=Paldyna |first2=M. |date=2018-03-26 |title=Taxonomic revision of the Oligocene percoid fish Oligoserranoides budensis (Heckel, 1856), from the Paratethys and paleobiogeographic comments |url=https://revistes.ub.edu/index.php/GEOACTA/article/view/GeologicaActa2018.16.1.5 |journal=Geologica Acta |language=en |volume=16 |issue=1 |pages=75–92 |doi=10.1344/GeologicaActa2018.16.1.5 |issn=1696-5728}}]]

Canyon Ferry Reservoir

|32.0 ± 0.1 Ma

|Montana, US

|A diverse Early Oligocene plant and insect fossil site.{{Cite journal |last=CoBabe |first=E. A. |date=1 June 2002 |title=A new insect and plant Lagerstatte from a Tertiary lake deposit along the Canyon Ferry Reservoir, southwestern Montana |url=http://rmg.geoscienceworld.org/cgi/doi/10.2113/gsrocky.37.1.13 |journal=Rocky Mountain Geology |language=en |volume=37 |issue=1 |pages=13–30 |doi=10.2113/gsrocky.37.1.13 |issn=1555-7332 |access-date=8 June 2024 |via=GeoScienceWorld|url-access=subscription }}

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Luberon (Campagne-Calavon Formation)

|~30 Ma

|Cereste, France

|A group of early Oligocene localities deposited along a large freshwater lake, preserving the fossils of plants, insects, fish, and terrestrial vertebrates, often with articulated skeletons, skin outlines, feathers, and original pigmentation patterns.{{Cite journal |last1=Coster |first1=Pauline |last2=Legal |first2=Stephane |date=2021-11-08 |title=An Early Oligocene Fossil Lagerstätten from the Lacustrine Deposits of the Luberon UNESCO Global Geopark |url=https://oiccpress.com/gcr/article/view/2800/986 |journal=Geoconservation Research |language=en |volume=4 |issue=2 |doi=10.30486/gcr.2021.1915524.1068 |issn=2588-7343}}

|File:Dapalis macrurus de l'oligocène de Céreste.JPG, one of the most common fossils of the site]]

Rauenberg/Frauenweiler Clay Pit

|30 Ma

|Baden-Württemberg, Germany

|A marine fossil site with an Arctic-like invertebrate fauna and a Paratethyan vertebrate fauna displaying evidence of intermittent anoxia.{{cite journal |last1=Maxwell |first1=Erin E. |last2=Alexander |first2=Stefanie |last3=Bechly |first3=Günter |last4=Eck |first4=Kristina |last5=Frey |first5=Eberhard |last6=Grimm |first6=Kirsten |last7=Kovar-Eder |first7=Johanna |last8=Mayr |first8=Gerald |last9=Micklich |first9=Norbert |last10=Rasser |first10=Michael |last11=Roth-Nebelsick |first11=Anita |last12=Salvador |first12=Rodrigo B. |last13=Schoch |first13=Rainier R. |last14=Schweigert |first14=Günter |last15=Stinnescbeck |first15=Wolfgang |last16=Wolf-Schwenniger |first16=Karin |last17=Ziegler |first17=Reinhard |date=1 December 2016 |title=The Rauenberg fossil Lagerstätte (Baden-Württemberg, Germany): A window into early Oligocene marine and coastal ecosystems of Central Europe |url=https://www.sciencedirect.com/science/article/abs/pii/S0031018216305521 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=463 |pages=238–260 |doi=10.1016/j.palaeo.2016.10.002 |bibcode=2016PPP...463..238M |access-date=29 April 2023|url-access=subscription }}

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Sangtang Lagerstätte

|~28 – 23 Ma

|Guangxi, China

|A section of the Late Oligocene Yongning Formation with one of the very few known Cenozoic assemblages of mummified plant fossils,{{Cite journal |last1=Quan |first1=Cheng |last2=Fu |first2=QiongYao |last3=Shi |first3=GongLe |last4=Liu |first4=YuSheng |last5=Li |first5=Long |last6=Liu |first6=XiaoYan |last7=Jin |first7=JianHua |date=2016-01-09 |title=First Oligocene mummified plant Lagerstätte at the low latitudes of East Asia |url=http://dx.doi.org/10.1007/s11430-015-5250-z |journal=Science China Earth Sciences |volume=59 |issue=3 |pages=445–448 |doi=10.1007/s11430-015-5250-z |issn=1674-7313|url-access=subscription }} including mummified fruits.{{Cite journal |last1=Shi |first1=Xiang-Gang |last2=Fu |first2=Qiong-Yao |last3=Jin |first3=Jian-Hua |last4=Quan |first4=Cheng |date=21 June 2017 |title=Mummified Oligocene fruits of Schima (Theaceae) and their systematic and biogeographic implications |journal=Scientific Reports |language=en |volume=7 |issue=1 |page=4009 |doi=10.1038/s41598-017-04349-6 |issn=2045-2322 |pmc=5479851 |pmid=28638066 }}{{Cite journal |last1=Fu |first1=Qiong-Yao |last2=Li |first2=Long |last3=Jin |first3=Jian-Hua |last4=Liu |first4=Xiao-Yan |last5=Quan |first5=Cheng |date=17 May 2017 |title=Mummified fruits of Choerospondias nanningensis sp. nov. (Anacardiaceae) from the upper Oligocene of a low latitude site in East Asia |url=https://onlinelibrary.wiley.com/doi/10.1111/jse.12255 |journal=Journal of Systematics and Evolution |language=en |volume=55 |issue=5 |pages=477–483 |doi=10.1111/jse.12255 |issn=1674-4918 |access-date=12 May 2024 |via=Wiley Online Library|url-access=subscription }}{{Cite journal |last1=Han |first1=Meng |last2=Manchester |first2=Steven R. |last3=Wu |first3=Yan |last4=Jin |first4=Jianhua |last5=Quan |first5=Cheng |date=2 August 2017 |title=Fossil fruits of Canarium (Burseraceae) from Eastern Asia and their implications for phytogeographical history |url=https://www.tandfonline.com/doi/full/10.1080/14772019.2017.1349624 |journal=Journal of Systematic Palaeontology |language=en |volume=16 |issue=10 |pages=841–852 |doi=10.1080/14772019.2017.1349624 |issn=1477-2019 |access-date=12 May 2024 |via=Taylor and Francis Online|url-access=subscription }}

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Enspel Lagerstätte

|

24.79-24.56 Ma

|

Rhineland-Palatinate, Germany

|

A Chattian maar deposit famous for its insect fossils.{{cite journal |last1=Wedmann |first1=Sonja |last2=Poschmann |first2=Markus |last3=Hörnschemeyer |first3=Thomas |date=19 November 2009 |title=Fossil insects from the Late Oligocene Enspel Lagerstätte and their palaeobiogeographic and palaeoclimatic significance |journal=Palaeobiodiversity and Palaeoenvironments |volume=90 |pages=49–58 |doi=10.1007/s12549-009-0013-5 |s2cid=84729592 |doi-access=free }}

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Aix-en-Provence Formation

|~24 Ma

|Provence, France

|A terminal Oligocene brackish palaeoenvironment.{{cite journal |last1=Gaudant |first1=Jean |last2=Nel |first2=André |last3=Nury |first3=Denise |last4=Véran |first4=Monette |last5=Carnevale |first5=Giorgio |date=August–September 2018 |title=The uppermost Oligocene of Aix-en-Provence (Bouches-du-Rhône, Southern France): A Cenozoic brackish subtropical Konservat-Lagerstätte, with fishes, insects and plants |url=https://www.sciencedirect.com/science/article/pii/S1631068317300799 |journal=Comptes Rendus Palevol |volume=17 |issue=7 |pages=460–478 |doi=10.1016/j.crpv.2017.08.002 |bibcode=2018CRPal..17..460G |s2cid=134885555 |access-date=29 June 2023|url-access=subscription }}

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=Neogene=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

Dominican amber30–10 MaDominican Republic

|Dominican amber differentiates itself from Baltic amber by being nearly always{{citation needed|date=January 2023}} transparent, and it has a higher number of fossil inclusions. This has enabled the detailed reconstruction of the ecosystem of a long-vanished tropical forest.George Poinar, Jr. and Roberta Poinar, 1999. The Amber Forest: A Reconstruction of a Vanished World, (Princeton University Press) {{ISBN|0-691-02888-5}}

|File:Larval Amblyomma hard tick in Dominican amber-12.jpg tick preserved in amber.]]

Riversleigh25–15 MaQueensland, Australia

|This locality is recognised for the series of well preserved fossils deposited from the Late Oligocene to the Miocene. The fossiliferous limestone system is located near the Gregory River in the north-west of Queensland, an environment that was once a very wet rainforest that became more arid as the Gondwanan land masses separated and the Australian continent moved north.

|File:Reconstruction of Nimbadon lavarackorum mother and juvenile - journal.pone.0048213.g002.png marsupial Nimbadon lavarackorum.]]

Foulden Maar23 MaOtago, New Zealand

|These layers of diatomite have preserved exceptional fossils of fish from the crater lake, and plants, spiders, and insects from the sub-tropical forest that developed around the crater,{{cite journal |last1=Lee |first1=Daphne E. |last2=Kaulfuss |first2=Uwe |last3=Conran |first3=John G. |last4=Bannister |first4=Jennifer M. |last5=Lindqvist |first5=Jon K. |date=11 August 2016 |title=Biodiversity and palaeoecology of Foulden Maar: an early Miocene Konservat-Lagerstätte deposit in southern New Zealand |url=https://www.tandfonline.com/doi/abs/10.1080/03115518.2016.1206321 |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=40 |issue=4 |pages=525–541 |doi=10.1080/03115518.2016.1206321 |bibcode=2016Alch...40..525L |s2cid=132004293 |access-date=29 April 2023|url-access=subscription }} along with in situ pollen.

|File:Galaxias effusus whole (cropped).jpg fish species Galaxias effusus.]]

Ebelsberg Formation

|23-22 Ma

|Upper Austria, Austria

|This Aquitanian-aged konservat-lagerstätte records an exceptional fossil assemblage of an enormous number of plants, fish, marine mammals, and marine invertebrates from a section of the central Paratethys Sea.{{Cite journal |last1=Grunert |first1=Patrick |last2=Harzhauser |first2=Mathias |last3=Rögl |first3=Fred |last4=Sachsenhofer |first4=Reinhard |last5=Gratzer |first5=Reinhard |last6=Soliman |first6=Ali |last7=Piller |first7=Werner E. |date=2010 |title=Oceanographic conditions as a trigger for the formation of an Early Miocene (Aquitanian) Konservat-Lagerstätte in the Central Paratethys Sea |url=https://doi.org/10.1016/j.palaeo.2010.04.001 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=292 |issue=3–4 |pages=425–442 |doi=10.1016/j.palaeo.2010.04.001 |issn=0031-0182|url-access=subscription }}

|File:Austromola angerhoferi.jpg, a sunfish]]

Chiapas amber23-15 MaChiapas, Mexico{{Cite journal |title=Two Flat-Backed Polydesmidan Millipedes from the Miocene Chiapas-Amber Lagerstätte, Mexico | first1 = Francisco | last1 = Riquelme | first2 = Miguel | last2 = Hernández-Patricio | first3 = Arnulfo | last3 = Martínez-Dávalos | display-authors = etal | date = 2014 | doi = 10.1371/journal.pone.0105877 | journal = PLOS ONE |volume=9 |issue=8 |pages=e105877 |pmid=25162220 |pmc=4146559 | bibcode = 2014PLoSO...9j5877R | doi-access = free }}

|As with other ambers, a wide variety of taxa have been found as inclusions including insects and other arthropods, as well as plant fragments and epiphyllous fungi.

|File:Plos One e105877 Figure 4-A Anbarrhacus adamantis.png.]]

Clarkia fossil beds20-17 MaIdaho, US

|The Clarkia fossil beds site is best known for its fossil leaves. Their preservation is exquisite; fresh leaves are unfossilized, and sometimes retain their fall colors before rapidly oxidizing in air. It has been reported that scientists have managed to isolate small amounts of ancient DNA from fossil leaves from this site. However, other scientists are skeptical of the validity of this reported occurrence of Miocene DNA.

|File:Clarkia leaf.jpg

Barstow Formation19–13.4 MaCalifornia, US

|The sediments are fluvial and lacustrine in origin except for nine layers of rhyolitic tuff. It is well known for its abundant vertebrate fossils including bones, teeth and footprints. The formation is also renowned for the fossiliferous concretions in its upper member, which contain three-dimensionally preserved arthropods.

|File:CamelFootprintBarstowMiocene.jpg.]]

Shanwang Formation18-17 MaShandong Province, China

|Fossils have been found at this site in dozens of categories, representing over 600 separate species. Animal fossils include insects, fish, spiders, amphibians, reptiles, birds and mammals. Insect fossils have clear, intact veins. Some have retained beautiful colours.

|File:Lusorex taishanensis.jpg]]

Morozaki Group{{Cite journal |last1=Mah |first1=Christopher L. |last2=Blake |first2=Daniel B. |date=2012-04-27 |title=Global Diversity and Phylogeny of the Asteroidea (Echinodermata) |journal=PLOS ONE |language=en |volume=7 |issue=4 |pages=e35644 |doi=10.1371/journal.pone.0035644 |issn=1932-6203 |pmc=3338738 |pmid=22563389 |bibcode=2012PLoSO...735644M |doi-access=free }}18-17 MaAichi Prefecture, Japan

|Known from well-preserved deep sea fauna including fish, starfish and arthropods like crabs, shrimps and giant amphiopods.

|

Sandelzhausen16 MaBavaria, Germany

|A Middle Miocene vertebrate locality.{{cite journal |last1=Moser |first1=Markus |last2=Rössner |first2=Gertrud E. |last3=Göhlich |first3=Ursula B. |last4=Böhme |first4=Madelaine |last5=Fahlbusch |first5=Volker |date=7 February 2009 |title=The fossil lagerstätte Sandelzhausen (Miocene; southern Germany): history of investigation, geology, fauna, and age |url=https://link.springer.com/article/10.1007/s12542-009-0012-x |journal=PalZ |volume=83 |pages=7–23 |doi=10.1007/s12542-009-0012-x |s2cid=55172519 |access-date=29 April 2023}}

McGraths Flat~16-11 MaNSW, Australia

|Deposited in unusual conditions that record microscopic details of soft tissues and delicate structures. Fossil evidence of animals with soft bodies, unlike the bones of mammals and reptiles, is rare in Australia, and discoveries at McGraths' Flat have revealed unknown species of invertebrates such as insects and spiders.{{Cite journal |title=A Lagerstätte from Australia provides insight into the nature of Miocene mesic ecosystems | first1 = Matthew | last1 = McCurry | first2 = David | last2 = Cantrill | first3 = Patrick | last3 = Smith | display-authors = etal | date = 2022 | doi = 10.1126/sciadv.abm1406 | journal = Science Advances|volume=8 |issue=1 |pages=eabm1406 | pmid = 34995110 | pmc = 8741189 | bibcode = 2022SciA....8.1406M }}

|File:Megamonodontium mccluskyi (AM F.145559).jpg ]]

Dolnja Stara

|~15 Ma

|Slovenia

|A barnacle fossil site preserving barnacles shortly after settlement attached to mangrove leaves.{{Cite journal |last1=Kočí |first1=Tomáš |last2=GašParič |first2=Rok |last3=Buckeridge |first3=John |last4=KočOvá Veselská |first4=Martina |last5=šOster |first5=Aleš |date=29 May 2023 |title=The first record of a Konservat-Lagerstätten in which early post-settlement stages of fossil archaeobalanids (Cirripedia: Balanomorpha) are preserved |url=https://onlinelibrary.wiley.com/doi/10.1111/1749-4877.12728 |journal=Integrative Zoology |language=en |volume=19 |issue=2 |pages=200–223 |doi=10.1111/1749-4877.12728 |pmid=37248329 |issn=1749-4877 |access-date=12 May 2024 |via=Wiley Online Library|url-access=subscription }}

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Pisco Formation15-2 MaArequipa & Ica, Peru

|Several specialists consider the Pisco Formation one of the most important lagerstätten, based on the large amount of exceptionally preserved marine fossils, including sharks (most notably megalodon), penguins, whales, dolphins, birds, marine crocodiles and aquatic giant sloths.

|File:Acrophyseter deinodon restoration.jpg stem-physeteroid whale Acrophyseter.]]

Hindon Maar14.6 MaNew Zealand

|A maar preserving a Southern Hemisphere lake-forest ecosystem, including body fossils of plants, insects, fish, and birds, along with in situ pollen{{cite journal |last1=Kaulfuss |first1=Uwe |last2=Bannister |first2=Jennifer M. |last3=Conran |first3=John G. |last4=Kennedy |first4=Elizabeth M. |last5=Mildenhall |first5=Dallas C. |last6=Lee |first6=Daphne E. |date=April 2023 |title=Review of flowers and inflorescences with in situ pollen from the Miocene Foulden and Hindon Konservat-Lagerstätten, southern New Zealand |url=https://www.sciencedirect.com/science/article/abs/pii/S0034666722002287 |journal=Review of Palaeobotany and Palynology |volume=311 |page=104830 |doi=10.1016/j.revpalbo.2022.104830 |bibcode=2023RPaPa.31104830K |s2cid=255325092 |access-date=29 April 2023|url-access=subscription }} and coprolites of both fish and birds.{{cite journal |last1=Kaulfuss |first1=Uwe |last2=Lee |first2=Daphne E. |last3=Wartho |first3=Jo-Anne |last4=Bowie |first4=Elliot |last5=Lindqvist |first5=Jon K. |last6=Conran |first6=John G. |last7=Bannister |first7=Jennifer M. |last8=Mildenhall |first8=Dallas C. |last9=Kennedy |first9=Elizabeth M. |last10=Gorman |first10=Andrew R. |date=1 July 2018 |title=Geology and palaeontology of the Hindon Maar Complex: A Miocene terrestrial fossil Lagerstätte in southern New Zealand |url=https://www.sciencedirect.com/science/article/abs/pii/S003101821830052X |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=500 |pages=52–68 |doi=10.1016/j.palaeo.2018.03.022 |bibcode=2018PPP...500...52K |s2cid=134962038 |access-date=29 April 2023}}

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Ngorora Formation

|13.3-9 Ma

|Tanzania

|The alkaline palaeolake deposits of the Ngorora Formation contains articulated fish fossils that died en masse from asphyxiation during episodic ash falls or from rapid acidification.{{cite journal |last1=Rasmussen |first1=Cornelia |last2=Reichenbacher |first2=Bettina |last3=Lenz |first3=Olaf |last4=Altner |first4=Melanie |last5=Penk |first5=Stefanie B. R. |last6=Prieto |first6=Jerome |last7=Brüsch |first7=Dennis |date=28 December 2015 |title=Middle–late Miocene palaeoenvironments, palynological data and a fossil fish Lagerstätte from the Central Kenya Rift (East Africa) |url=https://pubs.geoscienceworld.org/geolmag/article-abstract/154/1/24/350474/Middle-late-Miocene-palaeoenvironments |journal=Geological Magazine |volume=154 |issue=1 |pages=24–56 |doi=10.1017/S0016756815000849 |s2cid=130521301 |access-date=29 June 2023|url-access=subscription }}

|File:Megistotherium.jpg was a large sized hyaenodont mammal ]]

Öhningen Maar

|13 Ma

|Baden-Württemberg, Germany

|This unit of the Upper Freshwater-Molasse contains a former crater lake that has produced long-renowned fossils for centuries, including several foundational to the science of paleontology. About 1,500 species have been described from these deposits.{{Cite journal |last=Uhl |first=Dieter |last2=Wuttke |first2=Michael |last3=Aiglstorfer |first3=Manuela |last4=Gee |first4=Carole T. |last5=Grandi |first5=Federica |last6=Höltke |first6=Olaf |last7=Kaiser |first7=Thomas M. |last8=Kaulfuss |first8=Uwe |last9=Lee |first9=Daphne |last10=Lehmann |first10=Thomas |last11=Oms |first11=Oriol |last12=Poschmann |first12=Markus J. |last13=Rasser |first13=Michael W. |last14=Schindler |first14=Thomas |last15=Smith |first15=Krister T. |date=2024-12-01 |title=Deep-time maar lakes and other volcanogenic lakes as Fossil-Lagerstätten – An overview |url=https://link.springer.com/article/10.1007/s12549-024-00635-0 |journal=Palaeobiodiversity and Palaeoenvironments |language=en |volume=104 |issue=4 |pages=763–848 |doi=10.1007/s12549-024-00635-0 |issn=1867-1608|doi-access=free }}

|File:Andrias scheuchzeri 3478.jpg Andrias scheuchzeri]]

Pi Gros13 MaCatalonia, Spain

|An ichnofossil lagerstätte containing annelid, mollusc, and sponge trace fossils. The fossil site no longer exists due to having been quarried for the construction of an industrial park.{{cite journal |last1=Belaústegui|first1=Zain |last2=Domènech |first2=Rosa |last3=Martinell |first3=Jordi |date=16 January 2018 |title=An Ichnofossil-Lagerstätte From the Miocene Vilanova Basin (NE Spain): Taphonomic and Paleoecologic Insights Related to Bioerosion Structures |url=https://pubs.geoscienceworld.org/sepm/palaios/article-abstract/33/1/16/526007/AN-ICHNOFOSSIL-LAGERSTATTE-FROM-THE-MIOCENE |journal=PALAIOS |volume=33 |issue=1 |pages=16–28 |doi=10.2110/palo.2017.082 |bibcode=2018Palai..33...16B |s2cid=135386238 |access-date=1 May 2023|url-access=subscription }}

Bullock Creek12 MaNorthern Territory, Australia

|Among the fossils at the Bullock Creek site have been found complete marsupial crania with delicate structures intact. New significant taxa identified from the Bullock Creek mid Miocene include a new genus of crocodile, Baru (Baru darrowi), a primitive true kangaroo, Nambaroo, with high-crowned lophodont teeth; and a new species of giant horned tortoise, Meiolania. New marsupial lion, thylacine, and dasyurid material has also been recovered.

|

Tunjice? (Middle Miocene)Slovenia

|This site is known worldwide for the earliest fossil records of seahorses.{{Cite journal |last1=Žalohar |first1=Jure |last2=Hitij |first2=Tomaž |date=2017-04-01 |title=The first known fossil record of pipehorses (Teleostei: Syngnathidae: Haliichthyinae) from the Miocene Coprolitic Horizon from the Tunjice Hills, Slovenia |url=https://www.sciencedirect.com/science/article/pii/S0753396917300253 |journal=Annales de Paléontologie |volume=103 |issue=2 |pages=113–125 |doi=10.1016/j.annpal.2017.04.001 |issn=0753-3969|url-access=subscription }}

|File:Tunjice_Hills_Hippocampus.jpg and H. slovenicus from this site.]]

Ashfall Fossil Beds11.83 MaNebraska, US

|The Ashfall Fossil Beds of Antelope County in northeastern Nebraska are rare fossil sites of the type called lagerstätten that, due to extraordinary local conditions, capture an ecological "snapshot" in time of a range of well-preserved fossilized organisms. Ash from a Yellowstone hotspot eruption 10-12 million years ago created these fossilized bone beds.

|File:Ashfall Fossil Beds - Teleoceras and Cormohipparion.jpg and the three-toed horse Cormohipparion.]]

Alcoota Fossil Beds8 MaNorthern Territory, Australia

|It is notable for the occurrence of well-preserved, rare, Miocene vertebrate fossils, which provide evidence of the evolution of the Northern Territory's fauna and climate. The Alcoota Fossil Beds are also significant as a research and teaching site for palaeontology students.

|File:Alcoota miscellaneous fossils.jpg marsupials ]]

Saint-Bauzile7.6-7.2 MaArdèche, France

|A Late Miocene site preserving articulated mammal skeletons with skin and fur impressions.{{cite journal |last1=Métais |first1=Grégoire |last2=Sen |first2=Sevket |date=August–September 2018 |title=The late Miocene mammals from the Konservat-Lagerstätte of Saint-Bauzile (Ardèche, France) Conservation exceptionnelle de mammifères du Miocène supérieur à Saint-Bauzile (Ardèche, France) |journal=Comptes Rendus Palevol |volume=17 |issue=7 |pages=479–493 |doi=10.1016/j.crpv.2018.05.001 |s2cid=134016889 |doi-access=free }}

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Capo San Marco Formation~7 MaSardinia, Italy

|A microbialite containing exceptionally preserved Girvanella-type filaments.{{Cite journal |last=Saint Martin |first=Jean-Paul |date=April–June 2010 |title=The Girvanella-like remains from Messinian marine deposits (Sardinia, Italy): Lagerstätten paradigm for microbial biota? |url=https://linkinghub.elsevier.com/retrieve/pii/S0753396910000200 |journal=Annales de Paléontologie |language=en |volume=96 |issue=2 |pages=33–50 |doi=10.1016/j.annpal.2010.10.002 |access-date=12 May 2024 |via=Elsevier Science Direct|url-access=subscription }}

Tresjuncos6 MaCuenca, Spain

|A Late Miocene lacustrine konservat-lagerstätte containing fossils of diatoms, plants, crustaceans, insects, and amphibians.{{cite journal |last1=Bustillo |first1=María Ángeles |last2=Díaz-Molina |first2=Margarita |last3=López-García |first3=María José |last4=Delclòs |first4=Xavier |last5=Peláez-Campomanes |first5=Pablo |last6=Peñalver |first6=Enrique |last7=Rodríguez-Talavera |first7=Rosario |last8=Sanchiz |first8=Borja |date=3 August 2017 |title=Geology and paleontology of Tresjuncos (Cuenca, Spain), a new diatomaceous deposit with Konservat-Lagerstätte characteristics from the European late Miocene |url=https://link.springer.com/article/10.1007/s41513-017-0032-4 |journal=Journal of Iberian Geology |volume=43 |issue=3 |pages=395–411 |doi=10.1007/s41513-017-0032-4 |hdl=2445/163339 |s2cid=133950518 |access-date=29 April 2023|hdl-access=free }}

Libros

|Late Miocene

|Teruel, Spain

|A Late Miocene lacustrine lagerstätte located inside a sulfur mine, including well-preserved remains of frogs, as well as beavers, birds, snake, insects, arachnids, and plant remains.{{Cite journal |last=Mcnamara |first=M. E. |last2=Orr |first2=P. J. |last3=Alcala |first3=L. |last4=Anadon |first4=P. |last5=Penalver |first5=E. |date=2012-03-20 |title=WHAT CONTROLS THE TAPHONOMY OF EXCEPTIONALLY PRESERVED TAXA--ENVIRONMENT OR BIOLOGY? A CASE STUDY USING FROGS FROM THE MIOCENE LIBROS KONSERVAT-LAGERSTATTE (TERUEL, SPAIN) |url=https://pubs.geoscienceworld.org/palaios/article/27/2/63-77/100147 |journal=PALAIOS |language=en |volume=27 |issue=2 |pages=63–77 |doi=10.2110/palo.2010.p10-126r |issn=0883-1351|url-access=subscription }}

|File:Pelophylax pueyoi 20170301.jpg]]

Gray Fossil Site4.9-4.5 MaTennessee, US

|As the first site of its age known from the Appalachian region, the Gray Fossil Site is a unique window into the past. Research at the site has yielded many surprising discoveries, including new species of red panda, rhinoceros, pond turtle, hickory tree, and more. The site also hosts the world's largest known assemblage of fossil tapirs.

|File:Skull of Pristinailurus bristoli.jpg, a North American relative of the modern Red Panda]]

Willershausen

|Late Pliocene

|Lower Saxony, Germany

|A lacustrine fossil site containing well preserved beetles.{{Cite journal |last1=Kolibáč |first1=Jiří |last2=Adroit |first2=Benjamin |last3=Gröning |first3=Elke |last4=Brauckmann |first4=Carsten |last5=Wappler |first5=Torsten |date=8 August 2016 |title=First record of the family Trogossitidae (Insecta, Coleoptera) in the Late Pliocene deposits of Willershausen (Germany) |url=http://link.springer.com/10.1007/s12542-016-0316-6 |journal=PalZ |language=en |volume=90 |issue=4 |pages=681–689 |doi=10.1007/s12542-016-0316-6 |issn=0031-0220 |access-date=8 June 2024 |via=Springer Link}}

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=Quaternary=

class="wikitable" align="center" width="100%"
Site(s)

!Age

!Location

!Significance

!Notable fossils/organisms

The Mammoth Site26 KaSouth Dakota, US

|The facility encloses a prehistoric sinkhole that formed and was slowly filled with sediments during the Pleistocene era. As of 2016, the remains of 61 mammoths, including 58 North American Columbian and 3 woolly mammoths had been recovered. Mammoth bones were found at the site in 1974, and a museum and building enclosing the site were established.

|File:Arctodus_Simus,_Hot_Springs,_South_Dakota.jpg, a large species of "short-faced" bear that was one of North America's largest predators during the Pleistocene.]]

Rancho La Brea Tar Pits40–12 KaCalifornia, US

|A group of tar pits where natural asphalt (also called asphaltum, bitumen, or pitch; brea in Spanish) has seeped up from the ground for tens of thousands of years. Over many centuries, the bones of trapped animals have been preserved. Among the prehistoric species associated with the La Brea Tar Pits are Pleistocene mammoths, dire wolves, short-faced bears, American lions, ground sloths, and, the state fossil of California, the saber-toothed cat (Smilodon fatalis).

|File:Mammuthus columbi Page.jpg excavated from the tar pits]]

Waco Mammoth National Monument65–51 KaTexas, US

|A paleontological site and museum in Waco, Texas, United States where fossils of 24 Columbian mammoths (Mammuthus columbi) and other mammals from the Pleistocene Epoch have been uncovered. The site is the largest known concentration of mammoths dying from a (possibly) reoccurring event, which is believed to have been a flash flood.

|File:Waco mammoth site QRT.jpg

El Breal de Orocual2.5–1 MaMonagas, Venezuela

|The largest asphalt well on the planet. Like the La Brea Tar Pits, this site preserves a number of megafauna like toxodonts, glyptodonts, camelids, and the felid Homotherium venezuelensis.

|File:El Breal de Orocual.png, Coragyps, Dasypus and Myrmecophaga]]

El Mene de Inciarte28–25.5 KaZulia, Venezuela

|Another series of tar pits. These also preserve a similar assemblage of megafauna.

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Naracoorte Caves500-1 KaSouth Australia, Australia

|A series of caves that preserve numerous pleistocene megafauna, like Thylacoleo, and is recognized as a World heritage site alongside the older, but geographically similar Riversleigh site.

|File:Thylacoleo skeleton in Naracoorte Caves.jpg

Mare aux Songes

|4 Ka

|Mauritius

|A marsh that preserves a diversity of subfossil animals and plants, many of which were driven to extinction without proper documentation following human arrival, most notably the famous dodo. The mortality assemblages may have formed from a freshwater lake that was occasionally impacted by catastrophic droughts.{{Cite journal |last1=Rijsdijk |first1=Kenneth F. |last2=Hume |first2=Julian P. |last3=Bunnik |first3=Frans |last4=Florens |first4=F. B. Vincent |last5=Baider |first5=Claudia |last6=Shapiro |first6=Beth |last7=van der Plicht |first7=Johannes |last8=Janoo |first8=Anwar |last9=Griffiths |first9=Owen |last10=van den Hoek Ostende |first10=Lars W. |last11=Cremer |first11=Holger |last12=Vernimmen |first12=Tamara |last13=De Louw |first13=Perry G. B. |last14=Bholah |first14=Assenjee |last15=Saumtally |first15=Salem |date=2009-01-01 |title=Mid-Holocene vertebrate bone Concentration-Lagerstätte on oceanic island Mauritius provides a window into the ecosystem of the dodo (Raphus cucullatus) |url=https://www.sciencedirect.com/science/article/pii/S0277379108002473 |journal=Quaternary Science Reviews |volume=28 |issue=1 |pages=14–24 |doi=10.1016/j.quascirev.2008.09.018 |issn=0277-3791}}

|File:Dodo-Skeleton Natural History Museum London England.jpg skeleton from Mare aux Songes]]

Crawford Lake

|800 Ya-present

|Ontario, Canada

|This lake is notable for its detailed preservation of rotifer and dinoflagellate fossils even after centuries, documenting the ecological shifts that occurred in and around the lake following the establishment of a Iroquoian village from 1268–1486 CE, and later following European colonization of the region in the early 19th century.{{Cite journal |last1=Krueger |first1=Andrea |last2=McCarthy |first2=Francine |date=2016 |title=Great Canadian Lagerstätten 5. Crawford Lake – A Canadian Holocene Lacustrine Konservat-Lagerstätte with Two-Century-Old Viable Dinoflagellate Cysts |url=https://www.erudit.org/en/journals/geocan/2016-v43-n2-geocan02561/1036779ar/ |journal=Geoscience Canada |language=en |volume=43 |issue=2 |pages=123–132 |doi=10.12789/geocanj.2016.43.086 |issn=0315-0941}}

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See also

  • List of fossil sites (with link directory)
  • Hoard, a concentration of human artifacts useful for similar reasons in archaeology

References

{{Reflist}}

Further reading

  • Penney, D. (ed.) 2010. Biodiversity of Fossils in Amber from the Major World Deposits. Siri Scienfic Press, Manchester, 304 pp.
  • {{cite web

| year=2003

| url=http://palaeo.gly.bris.ac.uk/Palaeofiles/Lagerstatten/

| title=Fossil Lagerstätten

| publisher=Department of Earth Sciences, University of Bristol

| access-date=2005-11-21

| archive-url=https://web.archive.org/web/20070612022023/http://palaeo.gly.bris.ac.uk/Palaeofiles/Lagerstatten/

| archive-date=12 June 2007

| url-status=dead

}} – A catalogue of sites of exceptional fossil preservation produced by MSc palaeobiology students at University of Bristol's Department of Earth Sciences.

  • {{cite journal |doi= 10.1144/jgs.157.1.173 |last= Orr|first= Patrick J.|author2=David J. Siveter|date=1 January 2000|title= Three-dimensional preservation of a non-biomineralized arthropod in concretions in Silurian volcaniclastic rocks from Herefordshire, England|journal= Journal of the Geological Society|volume= 157|issue = 1|pages= 173–86|bibcode= 2000JGSoc.157..173O|s2cid= 129635160|url= http://jgs.geoscienceworld.org/cgi/content/abstract/157/1/173|access-date= 2006-10-26 |author-link2= David J. Siveter}}
  • {{cite journal |doi= 10.1016/j.tree.2024.04.004 |last= Kimmig|first= Julien|author2=James D. Schiffbauer|date=25 April 2024|title= A modern definition of Fossil-Lagerstätten|journal= Trends in Ecology and Evolution|volume= 39|issue = 6|pages= 621–624|pmid= 38670863|url= https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(24)00086-7|access-date= 2024-05-05|url-access= subscription}}

{{Burgess shale type preservation}}

{{Authority control}}

{{DEFAULTSORT:Lagerstatte}}